菰属系统与演化研究－－叶片表皮微形态

菰属叶片表皮微形态以其具有稻型的硅质体,指状的双胞微毛,三角形的副卫细胞,应位在禾本科(Gramineae)稻族(Oryzeae)内,但因其硅质体不整齐排列于脉上,而且有时呈现十字形或浅节结形,双胞微毛的顶细胞较粗壮等独特性状,结合染色体2n=2x=30,34,而区别于稻族中其他各属(x=12),且其所具竹亚科(Bambusoideae)的胚型(F+PP)在稻族中也鲜有,故应在稻族中独立成菰亚族(subtribeZizaniinae)。其属下种间的叶表皮微形态特征,大都吻合孢粉、外稃表皮微形态特征的研究结果,揭示菰属在全世界有4种,2亚种。

STUDIES ON THE SYSTEMATIC AND EVOLUTIONRY OF ZIZANIA L.(GRAMINEAE)--LEAF EPIDERMIS

This paper deals with the characters of leaf epidermis of Zizania and its relative genera. We find that Zizania L. is similar to Gen. Oryza, Leersia etc. by the Oryza-type silica-bodies, triangular subsidiary-cells and filiform micro-hairs, but it is clearly distinguished from them by its silicabodies usually irregularely arranged on the veins (usually regularly arranged over the veins in Oryza, Leersia ctc.)and the stomata are sually overarched by papillae. Hence, this genus may be considered as an only genus of subtribe Zizaniinae in tribe Oryzeae by connected with its chromosome number 2n=2x=30, 34 different from every one with basic chromosome number x=12. There are 4 species 2 subspecies(Z. aquatica subsp, aquatica, Z. aquatica subsp. brevis(Fass.)S. L. Chen, Z. palustris subsp. palustris, Z. palustris subsp. interior(Fass.) S. L. Chert, Z. tenana Hitchc. and Z. latifolia(Griseb.)Turcz. ex Stapf) in the world.