Genetic variation in competition traits at different ages and time periods and correlations with traits at field tests of 4-year-old Swedish Warmblood horses

For many years, the breeding value estimation for Swedish riding horses has been based on results from Riding Horse Quality Tests (RHQTs) of 4-year-olds only. Traits tested are conformation, gaits and jumping ability. An integrated index including competition results is under development to both get as reliable proofs as possible and increases the credibility of the indexes among breeders, trainers and riders. The objectives of this study were to investigate the suitability of competition data for use in genetic evaluations of horses and to examine how well young horse performance agrees with performance later in life. Competition results in dressage and show jumping for almost 40 000 horses from the beginning of the 1960s until 2006 were available. For RHQT data of 14 000 horses judged between 1988 and 2007 were used. Genetic parameters were estimated for accumulated competition results defined for different age groups (4 to 6 years of age, 4 to 9 years of age and lifetime), and for different birth year groups. Genetic correlations were estimated between results at RHQT and competitions with a multi-trait animal model. Heritabilities were higher for show jumping than dressage and increased with increasing age of the horse and amount of information. For dressage, heritabilities increased from 0.11 for the youngest group to 0.16 for lifetime results. For show jumping corresponding values increased from 0.24 to 0.28. Genetic correlations between competition results for the different age groups were highly positive (0.84 to 1.00), as were those between jumping traits at RHQT and competition results in show jumping (0.87 to 0.89). For dressage-related traits as 4-year-old and dressage competition results the estimated genetic correlations were between 0.47 and 0.77. We suggest that lifetime results from competitions should be integrated into the genetic evaluation system. However, genetic parameters showed that traits had changed during the over 35-year period covered due to the development of the sport, which needs to be considered in future genetic evaluations.     

Effects of long-time series of data on genetic evaluations for performance of Swedish Warmblood riding horses

For Swedish Warmblood sport horses, breeding values (BVs) are predicted using a multiple-trait animal model with results from competitions and young horse performance tests. Data go back to the beginning of the 1970s, and earlier studies have indicated that some of the recorded traits have changed through the years. The objective of this study was to investigate the effects of including all performance data or excluding the older ones compared to a bivariate model (BM) considering performance traits in early and late periods as separate traits. The bivariate approach was assumed to give the most correct BVs for the actual breeding population. Competition results in dressage and show jumping for almost 40 000 horses until 2006 were available. For riding horse quality test (RHQT), data of 14 000 horses judged between 1973 and 2007 were used. Genetic correlations of 0.69 to 1.00 were estimated between traits recorded at different time periods (RHQT data) or different birth year groups (competition data). A cross-validation study and comparison of BVs using different sets of data showed that most accurate and similar results were obtained when BVs were predicted from either the BM or the univariate model including all data from the beginning of the recording. We recommend using all data and applying the univariate model to minimise the computational efforts for genetic evaluations and for provision of reliable BVs for as many horses as possible.     

Constraining free riding in public goods games: designated solitary punishers can sustain human cooperation

Much of human cooperation remains an evolutionary riddle. Unlike other animals, people frequently cooperate with non-relatives in large groups. Evolutionary models of large-scale cooperation require not just incentives for cooperation, but also a credible disincentive for free riding. Various theoretical solutions have been proposed and experimentally explored, including reputation monitoring and diffuse punishment. Here, we empirically examine an alternative theoretical proposal: responsibility for punishment can be borne by one specific individual. This experiment shows that allowing a single individual to punish increases cooperation to the same level as allowing each group member to punish and results in greater group profits. These results suggest a potential key function of leadership in human groups and provides further evidence supporting that humans will readily and knowingly behave altruistically.     

Blood Sport. A Social History of Spanish Bullfighting (With an Essay and Bibliography by Rosario Cambria).

Abstract

The objective of this research was to determine the effect of a five-day therapeutic riding day camp on children's anger, quality of life and perceived self-competence. Our study sample involved 16 able-bodied individuals (11±4.4 years) with no known physical or psychological disability, and no known history of psychotropic medications. The Children's Anger Inventory, Peds Quality of Life, and Self Perception Profile for Children were administered prior to riding on day one and after riding on day five. Results show that after five days of therapeutic riding camp, the total score of the anger inventory and scores for all sub-scales except frustration decreased significantly. No other differences were noted. Data analysis suggests that five days of therapeutic riding day camp can significantly impact on anger. These changes may be related to the child's relationship with the horse, the social environment of camp, the horse and riding, increased contact with nature, or a combination of these factors.     

Temporal and Spatial Meiofauna Fluctuations in an Inlet of the South West Baltic (Darss-Zingst Bodden Chain) with Special Reference to the Harpacticoida (Copepoda,Crustacea)

Diurnal, seasonal and annual fluctuations of the meiofauna population density were recorded during long term investigations carried out in an inlet of the southwestern coast of the Baltic from 1970 to 1980. Field experiments have shown a diurnal vertical migration of harpacticoids, which leave the sediment at night to colonize the pelagic zone 14 times more densely than during day. Besides the seasonal fluctuations considerable differences were observed between the meiofauna densities in two successive years from samples taken twice in a grid consisting of 98 stations: the abundances of all meiofauna taxa decreased by approximately 50 %. These fluctuations are not only caused by the reproductive activities of the meiofauna under different environmental conditions, but they are to a great extent a result of the active and passive movements of the meiofauna.     

Horse riding in urban conservation areas: Reviewing scientific evidence to guide management

Summary We undertook a literature review of the impacts of horse riding in conservation areas, and used it to guide management of horse riding in Canberra Nature Park (CNP), a large, fragmented semi‐natural park in and around the city of Canberra in the Australian Capital Territory. The literature review established that, because of their relatively large weight and small area in contact with the ground, horses have a relatively high potential for doing environmental damage. Impacts tend to be generally lowest for hikers, followed by motorcycles, horses and four‐wheeled vehicles. One study showed horse traffic caused more damage on established trails than motorcycles, off‐road bicycles or hikers. Most published studies of horse‐riding impacts in Australia have been conducted in alpine and subalpine environments, and in temperate woodlands and forests on sandstone near Sydney. They have shown that impacts are generally highest in previously untracked areas. Impacts on established trails are generally most marked on sections of trail that are wet, boggy or steep, and on unplanned and unmaintained trails. Impacts are lowest on constructed and maintained trails. Trail proliferation, associated with avoidance of untrafficable sections and short‐cutting, can be a major problem. Horses also have potential to spread weeds, because pastures and dried stock feeds contain large numbers of weed seeds that retain high levels of viability in horse manure. The risk of weed establishment is highest when manure is deposited in disturbed, damp sites, particularly when riding off‐track. Much less weed establishment is apparent when horse riders remain on‐track. Horse riding is a popular activity, but one that is relatively expensive to provide for, and one that may reduce opportunities for lower‐impact recreational park users. For all these reasons it appears socially equitable that provision is made for lower numbers of horse riders compared to numbers of park users involved in lower impact, more passive, recreational activities. We conclude by describing how this information was used to develop principles to guide management of horse riding and assess risk at individual sites in Canberra Nature Park.     

Genetic correlations between conformation traits and radiographic findings in the limbs of German Warmblood riding horses

Studbook inspection (SBI) data of 20 768 German Warmblood mares and radiography results (RR) data of 5102 Hanoverian Warmblood horses were used for genetic correlation analyses. The scores on a scale from 0 to 10 were given for conformation and basic quality of gaits, resulting in 14 SBI traits which were used for the correlation analyses. The radiographic findings considered included osseous fragments in fetlock (OFF) and hock joints (OFH), deforming arthropathy in hock joints (DAH) and distinct radiographic findings in the navicular bones (DNB) which were analyzed as binary traits, and radiographic appearance of the navicular bones (RNB) which was analyzed as a quasi-linear trait. Genetic parameters were estimated multivariately in linear animal models with REML using information on 24 448 horses with SBI and/or RR records. The ranges of heritability estimates were h² = 0.14–0.34 for the RR traits and h² = 0.09–0.50 for the SBI traits. Negative additive genetic correlations of r_g = -0.19 to -0.56 were estimated between OFF and conformation of front and hind limbs and walk at hand, and between DNB and hind limb conformation. There were indications of negative additive genetic correlations between DAH and all SBI traits, but because of low prevalence and low heritability of DAH, these results require further scrutiny. Positive additive genetic correlations of r_g = 0.37–0.52 were estimated between OFF and withers height and between OFH and withers height, indicating that selection for taller horses will increase disposition to develop OFF and OFH. Selection of broodmares with regards to functional conformation will assist, but cannot replace possible selection against radiographic findings in the limbs of young Warmblood riding horses, particularly with regards to OFF.     

Horses do not exhibit motor bias when their balance is challenged

In many equestrian pursuits such as dressage and show-jumping, it is important that the horse exhibits the same level of balance when ridden to the left as when ridden to the right in canter - that is, to show no motor bias. It is a long-held belief within such disciplines that to reduce bias that exists in horses and thus to enhance symmetry of performance to the left and right, the horse needs to be worked equally in both directions, although there is a lack of scientific evidence of this influencing bias. There also is little compelling evidence for either the existence or absence of motor bias in unridden (and therefore younger) or ridden (and therefore older) horses. In this study, we tested whether there was a difference in motor bias between unridden (n = 15) and ridden (n = 15) horses when their balance was challenged by cantering them in circles both to the left and to the right on the lunge. As indicators of a difference in balance between the left and right and thus as indicators of motor bias, we conducted three lunging tests - time spent in canter, whether the horse cantered on the correct lead and whether it became disunited. A grazing stance test, where the extended foreleg during grazing was recorded as the preferred forelimb, was also used to compare responses in a test where balance was not actively challenged, to the three lunging tests where balance was actively challenged. No bias was found in either the unridden or ridden groups when their balance was challenged, but ridden horses exhibited a motor bias in grazing stance - when their balance was not challenged. There was also a correlation between the responses in all three lunging tests, but none between the grazing stance test and any of the three lunging tests. We therefore conclude that neither ridden nor unridden horses are biased when their balance is challenged; thus it cannot be concluded that ambidextrous training affects an inherent bias, and that estimation of motor bias in horses is affected by the test conditions. Finally, if ridden horses are truly unbiased, strong human motor bias might be responsible for the common perception amongst riders that horses are biased.     

Case studies of adults receiving horse-riding therapy

Abstract

The purpose of this study was to examine the psychotherapeutic effect of riding therapy (RT) on a group of adult users of a social services mental health team in South Devon. The benefits of animal-assisted therapy (AAT) and pet ownership on human health and well-being are well documented. However, whilst research has been conducted on the benefits of hippotherapy (the employment of horse riding as a physiotherapeutic aid) little has been undertaken on the psychotherapeutic benefits of riding therapy. Through case studies, the current study sought to explore whether the participants benefited in terms of confidence, increased self-esteem and social/interaction skills. A further objective was to find out whether these benefits, if identified, were transferable to other areas of the riders' lives. The study was conducted utilizing a case study, participant observational methodology following the progress of six women with various mental health problems receiving RT on a weekly basis. The sessions comprised of learning to look after the horses and carrying out stable management tasks, in addition to the riding; the chief instructor was aware that an important factor of the RT was building up a relationship and trust with the horses on the ground. Methods employed to record the sessions, apart from participant observation, included interviews and questionnaires. The prime objective of the study was for the riders' experience of the therapy to be expressed in their own words. It was found that the participants benefited in areas ranging from increased confidence and self-concept, and that the therapy aided social stimulation and led to transferable skills being acquired.     

Genetic parameters for traits evaluated at field tests of 3- and 4-year-old Swedish Warmblood horses

There are two types of 1-day field tests available for young Swedish Warmblood sport horses; one test for 3-year olds and one more advanced test for 4-year olds. Conformation, gaits and jumping ability are evaluated at both tests. Studies on various genetic parameters were based on about 20 000 tested horses. The data for 4-year olds consisted of 30 years of testing. The aims of the study were to estimate genetic parameters for results from different time periods, and to estimate heritabilities for, and genetic correlations between, traits scored in the two tests. The judgement of traits was shown to have been changed during the 30 years of testing, resulting in changes in higher heritabilities in, and stronger genetic correlations between, later time periods. In the first time period, records showed higher residual and lower genetic variances than in the subsequent time periods. Genetic correlations between traits recorded in the first and last time period deviated considerably from unity. Further studies are needed to investigate how to treat data from the early period in genetic evaluations. Heritabilities were moderate to high for conformation traits (0.24 to 0.58) at both types of tests, except for correctness of legs (0.08). The heritabilities for gait traits were also moderate to high (0.37 to 0.53). For jumping traits, the heritabilities ranged between 0.17 and 0.33. The highly positive genetic correlations (0.82 to 0.99) between corresponding traits tested at the simpler test for 3-year olds and at the ridden test of 4-year olds implied that it would be desirable to include the test results of 3-year olds into the genetic evaluation as breeding values for Swedish Warmbloods for many years has only been based on results from 4-year olds.