青海曲麻莱-治多地区巴颜喀拉山群(上三叠统)遗迹化石

首次对青海南部曲麻莱-治多地区巴颜喀拉山群上部(上三叠统)遗迹化石进行了系统采集,共采获遗迹化石33属54种。据遗迹化石的形态、习性、水深等环境成因特征,可划分为两类组合:一类以Cosmorhaphe,Helminthopsis,Helminthoida,Megagrapton,Paleodictyon等为主,产生于浊流事件前;另一类以Chondrites,Neo-nereites,Palaeophycus,Phycosiphon,Planolites,Zoophycos等为主,产生于浊流事件后,两者均为半深海-深海Nereites遗迹相的典型代表,反映晚三叠世研究区应为浊流沉积发育的复理石相沉积。     

Ichnology of deep‐sea fan overbank deposits of the Ganei slates (Eocene,Switzerland)— a classical flysch trace fossil locality studied first by Oswald Heer

Ganei (Switzerland) is a classical locality for trace fossils. At this site, Heer (1877) described a large number of trace fossils, several of which were new taxa. The trace fossils occur in thin‐bedded turbidites in which the basal divisions of the Bouma sequence are typically absent; the turbidites are assigned to the Ganei Slates and are Eocene in age. They are interpreted to have been deposited in an overbank environment within an upper to middle fan area distal to a channel. Two trace‐fossil associations occur: the first (I) is characterized by bulldozing organisms producing biodeformational structures, Scolica, and Nereites irregularis; the second (II) association shows a distinct tiering pattern with near‐surface graphoglyptids and a mixed layer with simple tubes such as cf. Palaeophycus and Planolites, plus patterned tubes such as Nereites cirrinalis, and Chondrites. Deeper turbidite layers were colonized by Chondrites and Gyro‐phyllites. All trace fossils show a normal size spectrum compared to previously studied trace‐fossil associations, so the degree of oxygenation probably did not influence the composition of either trace‐fossil association. Seafloor sediment was probably soft and did not affect the trace‐fossil associations. Sedimentation rate and event frequency did not change and are estimated to have been in a range of 5–10 cm/1000 years and 2–5 events per 1000 years, respectively. The composition of trace‐fossil associations I and II is therefore interpreted to have been controlled by the benthic food content being higher for trace‐fossil association I than for II.     

Rhizocorallium hamatum (Fischer-Ooster 1858), a Zoophycos-like trace fossil from deep-sea Cretaceous-Neogene sediments

Rhizocorallium hamatum (Fischer-Ooster 1858) is a trace fossil of the Zoophycos group, which is distinguished by its mostly horizontal, branched spreite lobes. It has so far, been ascribed mainly to Zoophycos, but the latter should be limited to forms having helical whorls, which are absent in R. hamatum. It has also been ascribed to Phycosiphon, which, however, shows J-shaped spreite lobes, while the lobes of R. hamatum are U-shaped. R. hamatum is very similar to R. commune var. irregulare, but the latter displays a distinctly wider marginal tunnel with respect to lobe width. R. hamatum occurs from the Turonian to Eocene, possibly from the Hauterivian to Oligocene, but mostly from Maastrichtian to Palaeocene, deep sea, mainly turbiditic sediments rich in marl. The tracemaker, probably a ‘worm’-like invertebrate, ingested an organic-rich mud of the background sediment and relocated it into the middle to deep tiers within the underlying turbiditic marl, mostly in form of faecal pellets packed within the spreite lobes, for further use as a food resource. This way of feeding was a response to food deficiency on the deep-sea floor.     

Ordovician Bathyal Trace Fossils From Metasiliciclastics in Central Norway and Their Sedimentological and Paleogeographical Implications

Numerous trace fossils are recognized in metasiliciclastic rocks of the Lower-Middle Ordovician Lower Hovin Group, Upper Ordovician Ekne Group and on roofing slates from an unknown locality (probably Ordovician) from the Trondheim region, central Norway. The trace fossil assemblages are dominated by meandering pascichnial forms, i.e., Helminthoidichnites in the Lower Hovin Group and Helminthoidichnites and Nereites in the slates. Protovirgularia, Dictyodora, ?Planolites and ?Palaeophycus are less common. Alcyonidiopsis, ?Trichophycus, Chondrites, cf. Chondrites, Gordia, ?Phycodes, ?Helminthopsis, cf. Naviculichnium, Treptichnus, Saerichnites, Megagrapton and ?Paleodictyon (Squamodictyon) are rare. The trace fossils were collected or observed at 17 different localities and represent assemblages belonging to the Nereites ichnofacies (deep-sea flysch deposits with thinly bedded turbidites). They reflect a distal Nereites subfacies (fan-fringe or basin-plain deposits) transitional to a Nereites/Paleodictyon subfacies (proximal fan facies). The trace fossil assemblages indicate an opportunistic style of colonization related to fluctuations in food supply introduced by turbidity currents.     

Giant Paleodictyon in Eocene flysch

A giant Paleodictyon gomezi with a maximum mesh diameter of 13 cm and covering more than 0.5 m² occurs in the lower Eocene flysch near Zumaya. The giant Paleodictyon occurs with flute casts and, therefore, must have been produced deeply in the sediment. The lengthening of mesh cells parallel and perpendicular to the flute casts implies changing current directions during trace production. Rows of elongate and offset mesh cells reflect the row-by-row production of the trace; however, the base line followed by the trace fossil producer (meandering or spiral) cannot be reconstructed because of incomplete preservation. The extraordinary size of this giant Paleodictyon contradicts the evolutionary trend of deep-sea trace fossils toward optimization and miniaturization, since similar large Paleodictyon specimens occur in Silurian flysch.     

Intrastratal trace fossil zonation,Cretaceous flysch of northern California

Dissection of fine‐grained turbidites within the Franciscan complex at Trinidad Harbor reveals intrastratal zonation of biogenic structures. The pre‐depositional association comprises only three ichnotaxa (preserved in convex hyporelief) and is less diverse than the tiered post‐depositional association (comprised by at least seven ichnotaxa preserved in full relief). The latter association is dominated by Melatercichnus (n. ichnogen.). Post‐depositional trace‐makers exploited abundant plant detritus deposited with the turbidite sand. Temporal alternation of trace fossil associations resulted from the infrequent, but catastrophic, disturbance of pre‐turbidite communities and local replacement by assemblies of opportunistic organisms. Although probably short‐lived at specific sites, the post‐turbidite infauna was tiered, enabling a variety of deposit feeders to harvest phytodetritus of varied size or density simultaneously at different depths within newly deposited turbidite sand layers.     

Diverse tiering patterns in Paleogene flysch trace fossils,Magura nappe,Carpathian Mountains,Poland

Diversified tiering patterns of trace fossils were observed in three Paleogene lithostratigraphic units of the Magura nappe: the Szczawnica Formation (Paleocene‐early Eocene); the Belove?a beds (middle Eocene); and the Piwniczna Sandstone Member of the Magura Formation (middle‐upper Eocene). Sediments of the first unit were deposited under oxygen‐poor conditions, and those of the second and third units under well‐aerated conditions. Thick‐bedded turbidites and fluxoturbidites of the third unit accumulated quickly, in contrast to ‘normal flysch’ of the first and second units. Analyses of tiering patterns show a thickening of tiers following the improvement of sediment oxygenation. Slowly accumulated deposits of ‘normal’ flysch exhibit a more mature, well‐developed tiering pattern than those which accumulated quickly.