The history and chronology of siwalik cercopithecids

Cercopithecid monkeys make a comparatively late appearance in the Miocene Siwalik formations of the Indian Subcontinent. The oldest well-dated specimen is a 6.3 MY old colobine, ?Presbytis sivalensis, and it is doubtful that any cercopithecids were present in the Siwalik faunas prior to 7.0 MY, a date that is considerably later than estimates for their appearance elsewhere in Eurasia. Cercopithecines appear later than colobines. Although their temporal ranges are uncertain, cercopithecines evidently were not present before 3.2 MY and possibly not until 2.5 MY. They are represented by three species,?Macaca palaeindica, Procynocephalus subhimalayanus, andTheropithecus delsoni. In the Siwaliks the appearances of colobines and cercopithecines coincide with or follow shortly after major faunal changes that also bring other northern Eurasian and African taxa into the region.     

Boselaphines (Artiodactyla, Ruminantia, Bovidae) from the Middle Siwaliks of Hasnot, Pakistan

In this paper, boselaphine material from several localities in the area of the Hasnot Pakistan, is described, identified, and discussed. Four species that belong to three different genera of the tribe Boselaphini have been found: Selenoportax vexillarius, S. lydekkeri, Pachyportax latidens and Eotragus sp. Eotragus sp. is reported for the first time from the Hasnot and consequently from other Upper Middle Siwalik sediments of Pakistan and equivalent strata of the world, extending the range of the genus from the Lower to the Middle Siwaliks. Reviewing the Siwaliks’ Selenoportax species, S. dhokpathanensis Akhtar and S. tatrotensis Akhtar are synonymized with S. lydekkeri and S. vexillarius, respectively.     

Les principales étapes de l’occupation humaine en bordure de l’Himalaya occidentale

Lithic industries attest that Western Himalayas, Hindu Kush and Pamir as well as the sub-Himalayan belt that is the Siwalik range, were inhabited during the prehistoric times. However, in thin transition zone between plains and high mountains, distinction is to be kept between low-middle altitude regions and high altitude regions (above 2000 m). In the former regions, the first human occupation tenuously appears around 2 Ma. In the Middle Pleistocene, the Lower Palaeolithic industries belong to two technological facies, the Acheulian and the Early Soanian. In the North-Western Siwaliks, the Soanian occurs in plenty on the river terraces, which were all built after 0.4 Ma, and the Acheulian appears between 0.5 and 0.7 Ma. The Middle Palaeolithic is still rich in cobble tools and therefore named Late Soanian. The Upper Palaeolithic is missing, perhaps due to climatic factors and the Neolithic is characterised by polished axes/adzes, but actually many of the cobble tools usually considered as Early Soanian may well belong to the cobble tool technical tradition common in South-East Asia during the Late Pleistocene and Early Holocene. In the north of the mountain range, the entire Palaeolithic sequence yielded by the loess in Southern Tajikistan is characterised, right from the late Lower Pleistocene, by lithic industries mainly compose of flakes, sometimes with cobble tools. In the high altitude zones, prehistoric sites are unknown before the Holocene and therefore they are contemporaneous with the Neolithic but some technological facies display particular features. The question is to know whether the originality of the industries, observed at different periods of time, is linked to techno-cultural adaptations to particular environments or to the isolation of the mountain populations.     

Pre-Quaternary hominin settlements in Asia: Archaeology,bio-lithostratigraphy and magnetostratigraphy evidences at Masol,Siwaliks, Northwestern India

The National Museum of Natural History, Paris, the National Centre for Scientific Research (CNRS/UMR 7194 and 8148) and the Society for Archaeological and Anthropological Research, Chandigarh, conduct paleontological and archaeological surveys in a sub-Himalayan territory well known for its Upper Pliocene fossiliferous deposits and named the Quranwala Zone. Since 2007, the team is investigating this formation in the Masol anticline and has collected four fossils with marks of butchery as well as numerous lithic artifacts. In this paper we show the first chopper extracted in stratigraphy from the deepest fossiliferous layer at more than 2.6 Ma as shown by the undisputable magnetostratigraphy, we describe the lithostratigraphy of the excavation, the techno-typology of the chopper and compare it with the collection of its locality and two other major paleonto-archaeological sites of Masol. The characteristics of this Masol lithic assemblage do not match those of the early Lower Pleistocene African Oldowan (2.55 Ma), but they are similar to those of the Chinese lithic industry of Longgupo (2.5 Ma). These activities open new perspectives regarding the phylogenetic and geographical origins of these Asian hominins as well as on their descendants, and justify the pursuit of surveys and test pits in the Quranwala zone.     

New specimen of Conohyus indicus (Lydekker, 1884) (Mammalia: Suidae) from the base of the Late Miocene, Jammu, India

Conohyus indicus is a poorly known tetraconodont suid from the summit of the Lower Siwaliks and the base of the Middle Siwaliks of the Indian Subcontinent. Even though it was first recorded well over a century ago, only 13 specimens have since been described in the literature, consisting of isolated teeth and incomplete mandible and maxilla fragments. We here describe another fragmentary mandible from the Ramnagar Member (uppermost Middle Miocene to basal Late Miocene) of the Siwalik Group, which contains the left canine lacking its tip, the alveolus of P/1 and the left P/2 – P/3 and right P/3 - M/1. The new specimen, albeit incomplete,provides interesting information about the anterior parts of the jaw and throws light on the systematic position of the species as well as the recently described species Conohyus thailandicus from Thailand.     

Siwalik synopsis: A long stratigraphic sequence for the Later Cenozoic of South Asia

The Tertiary sediments of northern Pakistan are an exceptional record of terrestrial sedimentation and represent most of Neogene time. Foremost is the Siwalik Group of the Potwar Plateau, for which multiple, superposed fossil levels span ∼18–6 Ma. Well-developed magnetostratigraphic control provides secure dating so that Siwalik fossil horizons may be interpolated into a time scale with resolution to 100,000 years. We describe the geographic setting of the Potwar, give an overview of the temporal distribution of faunas, and discuss changes in paleohabitat and paleoecology with coinciding faunal change, as seen from the Siwalik viewpoint. The long Siwalik biostratigraphy of many successive assemblages with its resolved time scale may be compared directly with other well-dated sequences. Immigrant arrival and timing of faunal change may be traced. The basins of the Iberian Peninsula show somewhat different timing of introduction of hipparionine horses, and faunal turnover in the Siwaliks clearly precedes the Vallesian crisis in Spain. In contrast to the increasingly seasonal precipitation of the late Miocene Potwar, the paleohabitat of coeval North China appears to have been moist and equable, with high diversity faunas. Continued development and comparison of resolved Neogene records allow increasing resolution of the patterns of faunal change on regional to global levels.     

New Bovid fossils from the Late Miocene Hasnot (Siwaliks,Pakistan)

The Siwalik Late Miocene bovids from the Hasnot deposits of Northern Pakistan are described here. The bovids are represented predominantly by boselaphines. The Hasnot outcrops range 7–5 Ma and correspond to the fauna of the Late Miocene–Early Pliocene of Eurasia and Africa. The associated fauna of Hasnot is suggestive a vast open land environment depicting sporadic dry and flood seasons, forcing a mosaic of ecotonal habitats with countless number of niches.     

New Alicornops (Rhinocerotidae) remains from Lower and Middle Siwaliks,Pakistan

Unpublished rhinocerotid remains from the Lower and Middle Siwaliks of Pakistan are described in this paper and recognized as two species of Alicornops. Alicornops complanatum (Heissig) is identified in the Dhok Pathan Formation of the Middle Siwaliks and Alicornops laogouense Deng in the Kamlial Formation of the Lower Siwaliks. The Dhok Pathan Formation levels with A. complanatum are roughly correlated with the late Miocene-Pliocene European mammal zones MN10-15. In turn, levels with A. laogouense of the Kamlial Formation would correlate with the middle–late Aragonian (middle Miocene) European MN5. The recognition of the Chinese species A. laogouense in the Potwar Plateau represents the first discovery of this taxon in Pakistan and increases the geographical and stratigraphic distributions of this species, and adds to the rhinocerotid association from the Siwaliks. In turn, the presence of A. complanatum in the Siwaliks of Potwar Plateau also enlarges its geographic distribution in Pakistan, as it was previously known from the Bugti Hills of Balouchistan. The absence of Alicornops from the Siwaliks in the Chinji and Nagri formations (between late MN5 and MN9 zones) might be due to an inadequate fossil record, as other rhinocerotid species are known from Kamlial to Dhok Pathan formations. However, the two recorded species of Alicornops could also reveal two independent migration waves as supported by the appearance of other taxa in different formations. A summary of fossil Cenozoic rhinocerotids from different areas of Pakistan is also presented.