Three new hybrids of Ophioglossum (Ophioglossaceae) from Monte Pisano,Tuscany (Central Italy)

Based on general morphology, spore measurements and ornamentation (scanning electron microscope), genome size estimation, and molecular systematics (trnL-trnF IGS), we show the extreme systematic complexity within the European representatives of the genus Ophioglossum. In particular, three hybrids from Tuscany are described: the tetraploid O. × pierinii Peruzzi, Magrini, Marchetti & Viane, seen as the hybrid between diploid O. lusitanicum L. and hexaploid O. azoricum C.Presl; the tetraploid O. × giovanninii Peruzzi, Pierini, Magrini, Marchetti & Viane, seen as the homoploid hybrid between tetraploid O. vulgatum L. and tetraploid O. × pierinii Peruzzi, Magrini, Marchetti & Viane; the pentaploid O. × pseudoazoricum Peruzzi, Pierini, Magrini, Marchetti & Viane, seen as the hybrid between hexaploid O. azoricum C.Presl and tetraploid O. vulgatum L. All the three new taxa grow in different localities in the Monte Pisano mountain range.     

Etude chimiosystematique des lycopodiales,isoetales, selaginellales et psilotales

A chemosystematic survey has been made of the flavonoids in the three orders of Lycopsida (Lycopodiales, Isoetales, Selaginellales), and of the single order of Psilotopsida (Psilotales). On the basis of the flavonoid patterns observed, a phylogenetic scheme is proposed for these taxa and their origin from fossil Pteridophyta (Psilophytales, Protolepidodendrales, Lepidodendraceae, Sigillariaceae) is discussed.