New fossil insects (Diaphanopterodea: Martynoviidae) from the Lower Permian of the Boskovice Basin, southern Moravia

Two new genera and new species assigned to the family Martynoviidae (Diaphanopterodea), Boskovicia celtai nov. gen., nov. sp. and Moravoptera hladilovae nov. gen., nov. sp. are described and illustrated from the Lower Permian (Artinskian/Sakmarian) locality of Obora, southern Moravia (Czech Republic). Discovery of the species Martynovia insignis Tillyard, originally described from the Lower Permian of Kansas (USA), supports close relationship of both faunas across Euramerica.     

New Middle Permian insects from Salagou Formation of the Lodève Basin in southern France (Insecta: Pterygota)

Summary

New insects are described from the Middle Permian (Guadalupian) of Salagou Formation in the Lodève Basin (Languedoc region, France). Palaeopterans are represented by new odonatopterans viz. Huangiopterum lodevense n. gen., n. sp. and Huangiopterum species B, attributed to clade Panodonata (Huangiopteridae n. fam.), already recorded through the family Saxonagrionidae in the same formation, and a diaphanopterodean Martynovia cf. halli Beckemeyer 2004 (Martynoviidae) known from the Lower Permian of Oklahoma. Archaeorthopterans are supplemented by Bethouxia ariegensis n. gen., n. sp., Paralongzhua elongata n. gen., n. sp., Lodevolongzhua incompleta n. gen., n. sp., the caloneurodean Jarmiloptera mouralensis n. gen., n. sp., and a cnemidolestodean cf. Tococladus sp. (Tococladidae). Permophyllum rotundatum n. sp. (?Tettigoniidae) demonstrates the first presence of Orthoptera with wings probably mimicking Peltaspermales reproductive organs in Late Palaeozoic possibly analogous to the known crypsis of roachoid tegmina with pteridosperm pinnules. Frequent occurrence of hardly identifiable grylloblattid wing fragments support their significant role in term of abundance and diversity of the Permian faunas. New taxa based on wing venation pattern as Genera and species undetermined A, B (Liomopteridae), Permobaharellus salagousensis n. gen., n. sp. (family uncertain), Lodevophlebia reticulata n. gen., n. sp. (Sylvaphlebiidae), Oborella lodevensis n. sp. (Euryptilonidae), and Genus and species undetermined C, were described. Paoliida is represented by a paoliid gen. et sp. indet. and three anthracoptilids Strephocladus gandi n. sp., Strephocladus mouralensis n. sp., Lodevocladus subtilis n. gen., n. sp. The first record of miomopteran Palaeomantis sp. (Paleomanteidae) from Salagou Formation is presented. Acercarian insects are represented by the first record of Thripida and diverse new hemipterans, namely Euroscytina lutevanorum n. gen., n. sp. (Cicadomorpha: Scytinopteridae), Prosbole dio n. sp., Prosbole garrici n. sp., Pervestigia cf. veteris Becker-Migdisova 1961 (Prosbolidae), Archescytina species A (Paleorrhyncha: Archescytinidae) and Permopsyllidium lesclansis n. sp. (Sternorrhyncha: Psylloidea: Protopsyllidiidae). Entomofauna from the Salagou Formation demonstrate highly diverse insect communities with some taxa linked to early and late Permian ones known from assemblages in Euramerica and Russia, such as Obora (Czech Republic), Wellington Formation (Kansas and Oklahoma, USA), and Tshekarda (Ural, Russia).     

New Martynoviidae from the Permian of Southern France (Lodève basin) (Insecta: Palaeoptera: Diaphanopterodea)

A new genus and two new species, Salagouneura chimaira nov. gen. and nov. sp. and Phaneroneura minuta nov. sp., together with several other specimens of uncertain affinities, are herein assigned to the family Martynoviidae. This family was previously only known from the Lower Permian of Oklahoma and Kansas (USA). The new specimens have been discovered in the Salagou Formation (Permian; Hérault, France). Stratigraphical and palaeoecological implications of the occurrence of this family in the Salagou Formation are discussed. These discoveries seem correlated with similar palaeoenvironmental conditions between the Salagou Formation and the Midco Salt Lake Formation (Oklahoma).