Pteridryaceae: A new fern family of Polypodiineae (Polypodiales) including taxonomic treatments

We undertook phylogenetic analyses to resolve the relationships of Pteridrys and related taxa based on six plastidmarkers (atpA, atpB, matK& rps16-matK, rbcL, rps4& rps4-trnS, and trnL & trnL-F) and nuclear pgiC.We included 195 accessions representing approximately 147 species in 38 genera, and seven of the nine families in Polypodiineae (eupolypods I). Tectariaceae s.l. (i.e., Arthropteris, Draconopteris, Hypoderris, Malaifilix, Pteridrys, Tectaria, and Triplophyllum in addition to Polydictyum) is recovered asmonophyletic (97% maximum likelihood bootstrap value), but with low (<50%) maximum parsimony jackknife value. The family Tectariaceae s.l. is therefore the only family in ferns without a corresponding non-parametric-based strong support in spite of our data totaling 9616 aligned base pairs. Tectariaceae s.l. can not be unambiguously recognized by any of the 13morphological characters analyzed. However, if the clade composed of Draconopteris, Malaifilix, Polydictyum, and Pteridrys (DMPP) is recognized as a distinct family, at least four morphological characters enable the distinction of the DMPP clade from its sister clade. Considering the uncertainty in the monophyly, the diagnosability, and the deep divergence, we propose to establish a new family, Pteridryaceae, to accommodate the DMPP clade. Species of Pteridryaceae share mostly the following characteristics: erect to suberect rhizomes, reduced basal pinnae, anastomosing or free venation, absence of catenate hairs at the leaf surface, and perine ornamentation lacking spines or spinules. Identification keys are provided for the four genera and 31 species of the DMPP clade (or Pteridryaceae). Reflecting the presented results, the recognition of Arthropteridaceae is the preferred taxonomic status of the Arthropteris clade.     

广西石灰岩地区蕨类新植物

广西石灰岩地区蕨类新植物周厚高,黎桦,黄玉源（广西农业大学,南宁５３０００５）关键词桂南爬树蕨;广西耳蕨;节毛耳蕨;石灰岩地区;广西ＮＥＷＳＰＥＣＩＥＳＯＦＰＴＥＲＩＤＯＰＨＹＴＥＦＲＯＭＬＩＭＥＳＴＯＮＥＡＲＥＡＯＦＧＵＡＮＧＸＩ￥ＺｈｏｕＨｏｕｇ...     

桂南爬树蕨的组织结构研究

本文对桂南爬树蕨的主要器官进行了解剖结构的研究,结果表明：茎的中柱为仅有两个新月型的维管束,中柱内及其周围的机械组织形成“８”字形。与爬树蕨相同,为新的中柱类型,中柱周围的几层皮层组织特化为机械组织的起始位置为对正维管束长轴的中央。而髓部的机械组织分化的起点是在中轴的中间位置,呈线形,同时向两侧分化。木质部分化为沿圆周线方向两端式,为新的分化方式。叶为等面叶,与爬树蕨相比,叶的厚度小,叶肉细胞层数少且细胞排列紧密,机械组织明显少。叶缘近圆形。根为二原型,直径比后者略小些。孢子类型与爬树蕨相同,孢子囊群无囊群盖。