An endemic brackish water faunule from the middle jurassic of southern qinghai province,China

An endemic brackish water faunule consisting of three taxa of bivalves,Hudsonella sinensis (Chen) n. gen.,Hudsonella sp., andProtocardia hepingxiangensis Chen & Lin occurs in the Bajocian of the southern Qinghai province, China.Hudsonella sinensis is a very variable species with a hinge structure characteristic of the subfamily Pachydontinae of the family Corbulidae. According to both geochemical data (stable carbon isotopes and the strontium/barium ratio) and ecological data the faunule lived in a brackish water embayment of mesohaline salinity (5#x2013;18 per mil) burrowing in soft substrate.     

Facies analysis of a large-scale Jurassic shelf-lagoon: the Kamar-e-Mehdi Formation of east-central Iran

The Callovian–Lower Kimmeridgian Kamar-e-Mehdi Formation of the Tabas Block (east-central Iran) is an up to 1,350-m-thick, fine-grained, marly-calcareous unit containing a basal Echellon Limestone Member (up to 180 m thick) and a terminal Nar Limestone Member (up to 100 m thick). The formation was deposited in a relatively deep shelf-lagoon that was part of the large-scale carbonate system of the Esfandiar Subgroup, extending N–S for about 500 km along the strike with a width of up to 100 km. The lagoonal Kamar-e-Mehdi Formation shows sedimentation rates of 150 m/myr, twice as high as those of the shelf-edge carbonate barrier (Esfandiar Platform). The repetitive lithologies and uniform depositional environment suggest equilibrium conditions between sedimentation and subsidence, related to constant slow rotation of the Tabas fault-block around a horizontal axis, the platform sitting on the crest, and the lagoon occupying the dip-slope. Lagoonal sedimentation was dominated by suspended carbonate mud and peloids from the eastern Esfandiar Platform whereas the subordinate siliciclastic material was derived from the west (Yazd Block). The diverse macrobenthos (mainly bivalves) suggests fully marine conditions for the major part of the Kamar-e-Mehdi Formation. However, towards the upper part, biotic impoverishment and the deposition of skeletal-poor, evaporitic sediments indicate increasing restriction. The overlying Magu Gypsum Formation marks the end of an arid basin-fill cycle and possibly forms an effective seal for hydrocarbon reservoirs in that area. The Esfandiar Subgroup was a Neotethys-facing carbonate margin, forming part of a belt of carbonate systems tracking the margins of the Iran Plate during Callovian to Late Jurassic times.     

Comparative ecological analysis of Toarcian (Lower Jurassic) benthic faunas from southern France and east-central Spain

Toarcian (Lower Jurassic) argillaceous-marly sediments of southern France (Causses) and east-central Spain (Barranco de la Puta section) were deposited in mid to lower shelf settings. They contain a low to moderately diverse, autochthonous fauna of benthic macroinvertebrates dominated by bivalves and gastropods, in the case of the Causses, and by bivalves and brachiopods in the Barranco de la Puta section. Five benthic associations are recognized at the latter locality and four at the former. In the Causses, low species diversity, evidence of seasonal mass mortality, presence of the opportunistic bivalve Parvamussium pumilum, local dominance of the soft bottom coral Thecocyathus mactrus and the dark-grey, fine-grained sediment point to high rates of sedimentation, high turbidity, intermittently lowered oxygen values and soft to soupy substrate conditions. These factors, together with eutrophic water masses, were the main environmental parameters governing faunal distribution. Soupy substrate conditions are thought to be largely biologically produced by the activity of infaunal depositfeeders, chiefly nuculoid bivalves. In the Barranco de la Puta section, abundant brachiopods, higher carbonate content of the sediment, and evidence of intermittent in-situ reworking and winnowing point to an influence of storm-induced currents, a shallower depositional depth, lower turbidity, a lower rate of sedimentation, and a somewhat firmer substrate. As a consequence, the fauna is largely dominated by epifaunal suspension-feeders. Compared with maximum and average sizes from elsewhere in the Jurassic, most bivalve species are distinctly smaller, a feature interpreted as stunting. Brachiopods, in contrast, reach normal size. The lack of nuculids, scarcity of deposit-feeders in general, and stunting are interpreted as evidence of an oligotrophic environment. Lowered oxygen conditions appear to have played only a very limited role in shaping the distribution pattern. Many of the differences between both areas can be explained by the different trophic regimes that reflect considerably higher run-off and consequently higher input of terrigenous sediment and dissolved nutrients to the basin in the case of the Causses. This in turn is thought to possibly reflect differences in regional climate; humid in southern France and more arid in eastcentral Spain.     

Platform-basin transect of a middle to late Jurassic large-scale carbonate platform system (Shotori mountains,Tabas area,east-central Iran)

Summary Following a phase of predominantly siliciclastic sedimentation in the Early and Middle Jurassic, a large-scale, low-latitude carbonate depositional system was established in the northern part of the Tabas Block, part of the central-east Iranian microplate, during the Callovian and persisted until the latest Oxfordian/Early Kimmeridgian. Running parallel to the present eastern block margin, a NNW/SSE-trending carbonate platform developed in an area characterized by reduced subsidence rates (Shotori Swell). The growth of this rimmed, flat-topped barrier platform strongly influenced the Upper Jurassic facies pattern and sedimentary history of the Tabas Block. The platform sediments, represented by the predominantly fine-grained carbonates of the Esfandiar Limestone Formation, pass eastward into slope to basin sediments of the Qal'eh Dokhtar Limestone Formation (platform-derived allochthonites, microbialites, and peri-platform muds). Towards the west, they interfinger with bedded limestones and marlstones (Kamar-e-Mehdi Formation), which were deposited in an extensive shelf lagoon. In a N−S direction, the Esfandiar Platform can be traced for about 170 km, in an E-W direction, the platform extended for at least 35–40 km. The width of the eastern slope of the platform is estimated at 10–15 km, the width of the western shelf lagoon varied considerably (>20–80 km). During the Late Callovian to Middle Oxfordian, the Esfandiar Platform flourished under arid climatic conditions and supplied the slope and basinal areas with large amounts of carbonates (suspended peri-platform muds and gravitational sediments). Export pulses of platform material, e.g. ooids and aggregate grains, into the slope and basinal system are interpreted as highstand shedding related to relative sealevel variations. The high-productivity phase was terminated in the Late Oxfordian when the eastern platform areas drowned and homogeneous deep water marls of the Upper Oxfordian to Kimmeridgian Korond Formation onlapped both the Qal'eh Dokhtar Limestone Formation and the drowned Esfandiar Limestone Formation. Tectonic instability, probably caused by faulting at the margins of the Tabas Block in connection with rotational movements of the east-central Iranian block assemblage, was responsible for the partial drowning of the eastern platform areas. In some areas, relicts of the platform persisted to produce shallow-water sediments into the Kimmeridgian.     

Preserved life positions of some Jurassic bivalves

The observed life positions of 23 species of Jurassic bivalves are recorded and compared to reconstructions derived from functional interpretations of their morphology and comparisons with Recent relatives. In the majority of cases, these inferences correspond with field observations. There are, however, cases such ashognomon where likely reconstructions differ. These could easily lead to an erroneous interpretation of the autecology of some species.     

A Late Cretaceous epeiric carbonate platform: the Haftoman Formation of Central Iran

An integrated study of the litho-, bio-, and microfacies of several sections has greatly improved the knowledge on the stratigraphy and depositional setting of the Coniacian to Campanian Haftoman Formation in the Khur area of the northern Yazd Block, Central Iran. Generally, the Haftoman Formation rests on a major tectonic unconformity and commences with a basal conglomerate followed by up to 900 m of shallow-water carbonates with local red sandstone intercalations. Five different depositional environments (from distal to proximal) characterize the facies associations (FA) of the Haftoman Formation: silty, spiculitic wackestone (proximal basin, FA I), bio-/intraclastic wacke-, pack-, and grainstone (marginal shoals, FA II), bioclastic rud-/float-/boundstone (outer platform, FA III), silty mud-/wackestone (lagoonal inner platform, FA IV), and sandstone/sandy limestone (areas close to the mouth of ephemeral streams, FA V). The litho-, micro-, and biofacies of the Haftoman Formation are typical for an epeiric carbonate platform characterized by an arid climate and lagoonal circulation, resulting in nutrient-poor waters, warm temperatures, and high salinities. The Haftoman Platform was attached to an emergent arid hinterland formed by the Anarak Metamorphic Complex to the west and southwest of the study area. Unconformity-bounded depositional units indicate sea-level changes that may correspond to 400-kyr high-frequency sequences but further studies are needed to fully exploit the potential of sequence stratigraphy for regional and inter-regional correlation of the Haftoman Formation.     

Sex determination of the Early Cretaceous clam shrimp Eosestheria middendorfii (Yixian Formation,China)

Clam shrimps have been recognized as a key group for the study of reproductive system evolution, owing to the diversity of sexual systems in extant members. However, there are comparatively little data on fossil taxa. In this study, we reveal the sexual system of the Early Cretaceous clam shrimp Eosestheria middendorfii (Yixian Formation, China). This is the first study that restricts the analysis to a single cohort, minimizing the otherwise considerable impact of ecophenotypic variation within this species. In addition, the presence of egg clutches, which identifies some individuals as either female or hermaphroditic, serves as an independent indicator for sex prior to the statistical treatment of the data. Obligate sexuality (‘dioecy’) is the inferred reproductive system for E. middendorfii and sexual dimorphism accounts for about 10% of the adult shape variation. Carapace shape variation resulting from malformation and deformation is more pronounced than the underlying sexual dimorphism. Subtle sexual dimorphism, smaller and slightly more elongate females, lateral egg clutches, and egg diameters of about 140 μm indicate that E. middendorfii is closely allied with the extant family Cyzicidae.     

Biometric methods for species recognition in <Emphasis Type="Italic">Trigonia</Emphasis> Bruguière (Bivalvia; Trigoniidae): a case study from the Upper Jurassic of Western Europe

From its first occurrence in the Middle Triassic to the Late Cretaceous, and thus for almost 200 Ma, Trigonia, type genus of the family Trigoniidae, has been a common constituent of global shallow-marine benthic faunas. The genus is highly over-split at the species level, which hampers sound biostratigraphic, palaeobiogeographic, and palaeoecologic applications. The present study focuses on two closely related species, i.e. Trigonia reticulata Agassiz and Trigonia pseudomeriani Choffat, and illustrates typical problems of species identification in Trigonia. T. reticulata is a well-known representative of the genus with a significant fossil record in Lower Oxfordian to Upper Kimmeridgian strata of western and central Europe, and likely is the ancestor of T. pseudomeriani, which is endemic to the Upper Oxfordian to Lower Kimmeridgian Alcobaça formation of the Lusitanian Basin (central Portugal). In order to effectively display intraspecific variability, digital analyses of a set of metric and non-metric parameters, including outline analysis, were applied to a representative number of specimens from different localities and stages. Originally, these methods were intended to show that both forms are conspecific. Instead, the analysis supports the distinctness of both species based on a very limited number of non-plastic characters. The methods applied can be transferred to other species in Trigonia and may lead to a modern species concept for the genus. Also, they may be successfully applied to other members of the Trigoniinae.