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The morphospace of 54 species of Commelinaceae from nine genera was examined with simultaneous attention to constraints, adaptive hypotheses and relatedness. Eleven morphological traits, including leaf length and width, angle between the leaves and internode distances, were measured for each species and analysed by principal components analysis and nested analysis of variance. The results revealed a significant signal of relatedness in vegetative morphology; genus explained 20–50% of the variance in a single trait. The relationships between some traits are consistent with adaptive explanations. The findings are consistent with the prediction that evolution for optimal phyllotaxis should be relaxed as self‐shading decreases, and that light availability governs leaf size and branching patterns. Constraints potentially explain some trait correlations, and support was found for the hypothesis that structural constraints govern leaf size and internode size correlations. © 2008 The Linnean Society of London, Botanical Journal of the Linnean Society, 2008, 158 , 257–268.  相似文献   
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An Exploratory Model of Play   总被引:1,自引:0,他引:1  
Play is defined as a state of experience in which the actor's ability to act matches the requirements for action in his environment. It differs from anxiety, in which the requirements outnumber the ability, and from boredom, in which the requirements are too few for the ability level of the actor. Games are reviewed with illustrations from a cross-cultural context of traditional and modern societies. It is suggested that games of skill, strategy, and chance all share structural characteristics that allow the player to limit his experiences so as to maximize the play experience as defined. Further theoretical implications are drawn from the model in terms of the relationship of individuals and the social system.  相似文献   
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How much water does a river need?   总被引:58,自引:0,他引:58  
1. This paper introduces a new approach for setting streamflow-based river ecosystem management targets and this method is called the 'Range of Variability Approach' (RVA). The proposed approach derives from aquatic ecology theory concerning the critical role of hydrological variability, and associated characteristics of timing, frequency, duration, and rates of change, in sustaining aquatic ecosystems. The method is intended for application on rivers wherein the conservation of native aquatic biodiversity and protection of natural ecosystem functions are primary river management objectives.
2. The RVA uses as its starting point either measured or synthesized daily streamflow values from a period during which human perturbations to the hydrological regime were negligible. This streamflow record is then characterized using thirty-two different hydrological parameters, using methods defined in Richter et al . (1996). Using the RVA, a range of variation in each of the thirty-two parameters, e.g. the values at ± 1 standard deviation from the mean or the twenty-fifth to seventy-fifth percentile range, are selected as initial flow management targets.
3. The RVA targets are intended to guide the design of river management strategies (e.g. reservoir operations rules, catchment restoration) that will lead to attainment of these targets on an annual basis. The RVA will enable river managers to define and adopt readily interim management targets before conclusive, long-term ecosystem research results are available. The RVA targets and management strategies should be adaptively refined as suggested by research results and as needed to sustain native aquatic ecosystem biodiversity and integrity.  相似文献   
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