印尼爪哇动物群与周口店地区及华南动物群对比

Recently the vertebrate faunal succession established by von Koenigswald in 1930s was replaced by a new one which was proposed by Sondaar (1984), which arose some problems in regional correlations, because the relationship between the two schemes are not very clear,and some of the guide fossils mentioned by von Koenigswald were not included in the new scheme. In China, we used to compare the Jetis Fauna with Gongwangling fauna, Trinil Fauna with Zhoukoudian and Yanjinggou Faunas, but at the present time, both of the faunal names were given up. The new dating works changed the old scheme of faunas, the oldest mammalian fossil record in Java is only 1.5 Ma, which had been thought to be Middle Pliocene in the past half century. The new results from the dating works are not well correlated yet with the turnover of faunal assemblages. It's sure that the Java Faunas have close relationship with that of China, but how we can correlate them precisely is still to be done in the future.
     

The late Cenozoic Thiaridae (Mollusca, Gastropoda, Cerithioidea) of the Albertine Rift Valley (Uganda-Congo) and their bearing on the origin and evolution of the Tanganyikan thalassoid malacofauna

The recent cerithioid malacofauna of meromictic Lake Tanganyika is unique in its degree of thalassoid convergence with marine molluscs. This is generally considered the result of a long-lasting intensive escalation and cladogenesis caused by a coevolutionary prey–predator interaction in a freshwater ecosystem with sea-like characteristics, i.e. exceptional longevity (ca. 7–12 Ma) and vast dimensions (present surface area: 32900 km², present maximum depth: 1470 m). In the Albertine Basin, ca 300 km north of the Tanganyika Trough, Palaeolake Obweruka existed during Mio-Pliocene times. In many aspects, it can be considered as a 'sister lake' of Lake Tanganyika, being also long-lived (from ca 7.5 to 2.5 Ma), extensive (surface: 27000 km²) and meromictic. Like Lake Tanganyika it belonged to the Congo catchment. Although thalassoid molluscs have been known from the Albertine deposits since the beginning of the 20th Century, previous researchers recognised only four polymorphic thiarid species, two of which were thalassoid. Detailed in situ collecting shows that the perceived low species diversity and high variability of the Obwerukan thalassoids is a phenomenon of the museum drawer. In the field the thiarid species and other molluscs are not extremely polymorphic, as formerly thought, but tend to occur in discrete morphometric packages according to stratigraphic level and geographic area. The species richness of the Palaeolake Obweruka thiarids has thus been severely underestimated, as is also the case for the Recent Tanganyikan thiarids. The present authors discern 35 species and 7 genera in the Albertine deposits (2 of which are new to science), and consider this to be a conservative estimate. In their degree of diversification and morphological escalation, the Obwerukan thiarids are thus comparable to the Tanganyika thalassoids. Arguments are presented that the thalassoids of these lakes are polyphyletic, that they are derived from the same genera (mainly Potadoma, Potadomoides and Pseudocleopatra) and that the tempo and mode of their intralacustrine evolution must have been largely similar. As for the tempo of evolution, the Albertine fossil record shows that escalation in shell morphology and ornamentation in the Thiaridae was extremely rapid, as in the Ampullariidae and Viviparidae (Van Damme & Pickford, 1995, 1999). In the Thiaridae heavily ornate shells evolved from an unornamented ancestor in two consecutive bursts, each lasting less than 0.1 million years over a total period of ca. 1 million years. The punctuated escalation in the thiarids was initiated millions of years after the lake was formed. This implies that the degree of escalation is not related to age of a lake and that the Tanganyikan thalassoids or part of them may not be particularly old, even though Lake Tanganyika itself is. As to the mode of evolution, it appears that escalation is a coevolutionary reaction to predation but that this selective pressure only triggers a reaction under specific conditions and in synergy with other environmental factors. The Obwerukan thiarid fossil record also indicates that iterative evolution of molluscan shells may be of frequent occurrence. What is considered to be a single polymorphic chronospecies, static over a great length of time, may actually be a sequence of several species that branched at different moments from the same ancestral phenotype. Finally, there is the inference that bottle-necking may be, geologically speaking, so common that it forms an essential force in macroevolution.     

Reconsidering locomotor habits and life style of the Balearic insular giant rodent Hypnomys Bate, 1918 from the allometry of the limb long bones

The genus Hypnomys Bate, 1918 includes some endemic Neogene chronospecies from Mallorca and Menorca, evolved in insularity conditions from the Lower Pliocene to the human arrival at the archipelago. The study of the allometric proportions (functional length and sagittal diameter) of the limbs’ long bones of Hypnomys eliomyoides Agusti, 1980 (Lower Pleistocene of Menorca), Hypnomys sp. (Pleistocene of Menorca), Hypnomys onicensis (Reumer, 1994) (Upper Pliocene-Lower Pleistocene of Mallorca) and Hypnomys morpheus Bate, 1918 (Upper Pleistocene of Mallorca) only indicates small differences with the garden dormouse (Eliomys quercinus Linnaeus, 1776) and other mainland rodents and insectivores. The study about the locomotion type by means of Similarity Matrix of Bou indicates that 1) Hypnomys spp. presents the greater similarities with the arboreal, jumping and gliding species; 2) the jumping and digging life style of Hypnomys spp. are more developed (25%) than in E. quercinus; 3) the walking locomotion of Hypnomys spp. is only a 7% greater than in E. quercinus; and 4) the degree of similarity with the gliding locomotion of Hypnomys sp. is greater (17%) than in E. quercinus. Some of this data indicates that Hypnomys spp. was better adapted to the arboreal life than E. quercinus, due to the relationship between the jumping and gliding capacities with the arboreal locomotion. The greater similarity of Hypnomys spp. with the walking locomotion does not necessarily involve more terrestrial habits. Our interpretation contrasts with earlier conclusions of Hypnomys life style (Hypnomys cf. onicensis and Hypnomys morpheus) where the terrestrial locomotion was overestimated in front of the arboreal locomotion. The locomotion type of Hypnomys would be related with the presence of important forest masses on the Balearic Islands during the Neogene, a different degree of environmental stress from that of the mainland ecosystems and a probable expansion of the ecological niche occupied by these species.     

The skull of Cteniogenys, a choristodere (Reptilia: Archosauromorpha) from the Middle Jurassic of Oxfordshire

The genus Ctemogenys was originally described as a lizard on the basis of isolated jaw fragments from the Upper Jurassic deposits of Como Bluff, Wyoming. The discovery of new material from a Middle Jurassic locality at Kirtlington, Oxfordshire, showed that Cteniogenys was not a lizard, but an early choristodere. The skull is represented by a collection of isolated bones, associated on the basis of fit and dermal sculpture pattern. The bones are here described and compared with those of the Late Cretaceous and Palaeocene choristoderes Champsosaurus and Simoedosaurus. Allowing for the much smaller size of the Middle Jurassic form, there is a close correspondence between the isolated bones of the three genera. Comparison with known choristoderes, based on an analysis of 53 derived character states, suggests that Ctemogenys is the most primitive of known genera. In general, the character states shared by all choristoderes support recent analyses which conclude that choristoderes arc derived from archosauromorph diapsids, not lepidosauromorphs as once thought. At Kirtlington, Cteniogenys forms part of a diverse microvertebrate assemblage including amiids, sharks, frogs, salamanders, lizards, mammals, crocodiles, pterosaurs, turtles and small dinosaurs.     

The colonisation of Madagascar by land-bound vertebrates

Despite discussions extending back almost 160 years, the means by which Madagascar's iconic land vertebrates arrived on the island remains the focus of active debate. Three options have been considered: vicariance, range expansion across land bridges, and dispersal over water. The first assumes that a group (clade/lineage) occupied the island when it was connected with the other Gondwana landmasses in the Mesozoic. Causeways to Africa do not exist today, but have been proposed by some researchers for various times in the Cenozoic. Over-water dispersal could be from rafting on floating vegetation (flotsam) or by swimming/drifting. A recent appraisal of the geological data supported the idea of vicariance, but found nothing to justify the notion of past causeways. Here we review the biological evidence for the mechanisms that explain the origins of 28 of Madagascar's land vertebrate clades [two other lineages (the geckos Geckolepis and Paragehyra) could not be included in the analysis due to phylogenetic uncertainties]. The podocnemid turtles and typhlopoid snakes are conspicuous for they appear to have arisen through a deep-time vicariance event. The two options for the remaining 26 (16 reptile, five land-bound-mammal, and five amphibian), which arrived between the latest Cretaceous and the present, are dispersal across land bridges or over water. As these would produce very different temporal influx patterns, we assembled and analysed published arrival times for each of the groups. For all, a ‘colonisation interval’ was generated that was bracketed by its ‘stem-old’ and ‘crown-young’ tree-node ages; in two instances, the ranges were refined using palaeontological data. The synthesis of these intervals for all clades, which we term a colonisation profile, has a distinctive shape that can be compared, statistically, to various models, including those that assume the arrivals were focused in time. The analysis leads us to reject the various land bridge models (which would show temporal concentrations) and instead supports the idea of dispersal over water (temporally random). Therefore, the biological evidence is now in agreement with the geological evidence, as well as the filtered taxonomic composition of the fauna, in supporting over-water dispersal as the mechanism that explains all but two of Madagascar's land-vertebrate groups.     

On Permian and Triassic insect faunas in relation to biogeography and the Permian-Triassic crisis

The taxonomic diversity dynamics of pterygote insects in the Permian and Triassic at the family/age level are considered. Different metrics of taxonomic diversity are compared. Biogeographic and taphonomic aspects of changes in the composition of insect faunas in the Permian and about the P-T transition are discussed. Some changes in the Permian insect faunas are of a biogeographic nature and do not indicate global changes in diversity. Insects with aquatic immatures were rather common in the Permian and Early Triassic, but these immatures are well represented in only few localities.