Potential Global Standard Stratotype-section and Point (GSSP) for a Cambrian stage boundary defined by the first appearance of the trilobite Ptychagnostus atavus, Drum Mountains, Utah, USA

The base of the Ptychagnostus (or Acidusus) atavus Zone is one of the most clearly recognizable horizons on an intercontinental scale in the Cambrian System, and would serve as an excellent position for the base of a new stage-level chronostratigraphic subdivision. Among well-exposed, readily accessible sections in Laurentia, the “Stratotype Ridge” section, Drum Mountains, western Utah, USA, is perhaps the most suitable for a Global Standard Stratotype-section and Point (GSSP) defined by the first appearance datum (FAD) of the cosmopolitan agnostoid trilobite P. atavus. In the “Stratotype Ridge” section, the FAD of P. atavus occurs near the base of a calcisiltite bed 62 m above the base of the Wheeler Formation. A position corresponding closely to this horizon can be recognized with precision in Gondwana, Siberia, Kazakhstan, and Baltica using a combination of stratigraphic tools, the most useful of which are trilobite biostratigraphy, conodont biostratigraphy, and sequence stratigraphy. Brachiopod biostratigraphy and chemostratigraphy provide general constraints on the position of the horizon intercontinentally.     

Advances in Cambrian stratigraphy and paleontology: Integrating correlation techniques, paleobiology, taphonomy and paleoenvironmental reconstruction

Papers resulting from the Fourth International Symposium on the Cambrian System, held in Nanjing, China, in 2005 cover three major aspects of geology and paleontology: (1) the developing global standard for Cambrian chronostratigraphy or regional correlation schemes; (2) regional lithostratigraphy, sedimentology and paleoenvironments; (3) organismal paleobiology, phylogenetic affinities and taphonomy.A generalized curve of carbon isotopes (δ¹³C) through the Cambrian suggests a relationship between major biotic events, sea level history and the development of deposits of exceptional preservation (Lagerstätten). Recognition of this relationship increases the importance of the δ¹³C profile as a tool for intercontinental and intracontinental correlation. Significant δ¹³C excursions in the Cambrian are: BACE (negative excursion at the base of the Cambrian System); ZHUCE (positive excursion in the lower part of Stage 2); SHICE (negative excursion in the middle part of Stage 2); CARE (positive excursion near the base of Stage 3); MICE (positive excursion in the lower part of Stage 4); AECE (negative excursion in the middle part of Stage 4); ROECE (negative excursion near the base of Stage 5); DICE (negative excursion beginning near the base of the Drumian Stage); SPICE (positive excursion beginning at the base of the Paibian Stage); TOCE (negative excursion near the top of Stage 10). All acronyms other than SPICE are newly proposed.     

Taxonomy of Ptychagnostus atavus: A key trilobite in defining a global Cambrian stage boundary

The taxonomy of Ptychagnostus atavus (Tullberg, 1880) and Ptychagnostus intermedius (Tullberg, 1880) has been confused, in part at least, because of inadequate or erroneous illustrations of specimens from syntype series. From further examination and new illustrations we conclude that those series are conspecific and that P. atavus is the senior synonym of P. intermedius. Laurentian agnostoids previously assigned to P. intermedius are reassigned to Ptychagnostus sinicus Lu, 1957. As revised, P. atavus is a common, globally distributed species in open-marine lithofacies. Clarification of its taxonomy is important because its first appearance datum (FAD) will likely define the base of a new global Cambrian stage.     

The Early Cambrian acritarch Skiagia and its significance for global correlation

Global records of Early Cambrian acritarchs from the Skiagia plexus reveal that they are stratigraphically significant and can be used as biostratigraphic markers. Skiagia ornata and Skiagia ciliosa have a cosmopolitan distribution and the levels of their first appearance datums are proposed to serve as biostratigraphic horizons for interregional correlation. These levels are close to the lower boundary of the undefined Cambrian Stage 3, and within the Stage 3, respectively. The acritarch assemblage diagnostic of the Heliosphaeridium dissimilare-S. ciliosa Zone of Baltica is recorded worldwide and some species are traced on four of the Cambrian continents that include South Australia, South China and Laurentia. This supports the intercontinental correlation of certain Lower Cambrian strata and establishes the time equivalence between the regional trilobite zones to which these strata are attributed. The regional zonation is briefly reviewed for the purpose of comparing and evaluating the acritarch ranges. The biostratigraphic correlation of the Lower Cambrian strata, belonging to different faunal provinces, is based for the first time on the same fossil species, which have a cosmopolitan distribution and well-defined first appearance datums, and on the entire diagnostic acritarch assemblages.     

Significance of Middle Cambrian mixed carbonate-siliciclastic units for global correlation: southern Nevada, USA

Sequence-stratigraphic analysis of the Middle Cambrian Highland Peak, Bonanza King, Swasey, and Wheeler formations in the Great Basin refines platform-to-basin correlations and distinguishes local tectonic events from eustasy. This analysis provides a reliable sea-level history through the Ptychagnostus gibbus and Ptychagnostus atavus trilobite intervals and confirms that the Global Stratotype Section and Point (GSSP) at the first appearance datum (FAD) of P. atavus was deposited during an overall sea-level rise. Deposition during the Middle Cambrian Ehmaniella/Bolaspidella biozones in the western U.S. is represented by two lithologically distinct successions: (1) a poorly fossiliferous, shallow-water, mixed carbonate-siliciclastic succession that is widespread across southern Nevada and southeastern California and (2) a highly fossiliferous, deeper water, fine-grained, siliciclastic succession in central Nevada and western Utah. The deeper water succession was deposited within the fault-controlled House Range Embayment and contains the P. atavus GSSP. Correlation of these disparate successions had been hampered by a lack of high-resolution biostratigraphic data, and limited chemostratigraphy and sequence stratigraphy. In this study, sequence-stratigraphic analysis indicates that the Condor Member of the Highland Peak Formation and the “mixed unit” of the Bonanza King Formation are the shallow-water platform equivalents of the basal Wheeler Formation encompassing the P. gibbus and lower P. atavus zones. The deepening event that is recorded in the P. gibbus Zone represents a major flooding surface that may be used as an important event marker for regional and global correlation. The overlying P. atavus GSSP, however, is within the later stage of transgression and may represent a globally synchronous event that can be correlated from platform to basin.     

Lower Cambrian Small Shelly Fossils of northern Sichuan and southern Shaanxi (China), and their biostratigraphic importance

Extensive information has been acquired on the occurrence of Lower Cambrian Small Shelly Fossils (SSF) in the shallow water sediments of the western fringe of the Yangtze Platform (eastern Yunnan and central Sichuan Provinces) during the last two decades. However, less is known on the temporal and spatial distribution of SSFs on the northern margin of the Yangtze Platform. Here we report SSF associations from Kuanchuanpu (Ningqiang County, southern Shaanxi Province), Shatan, and Xinli (both Nanjiang County, northern Sichuan Province) representing the Southwest Shaanxi region and from Sanlangpu, and Xiaowan (Xixiang County, southern Shaanxi Province) of the Southeast Shaanxi - Northwest Hubei region. The thickness of the Lower Cambrian SSF-bearing strata strongly decreases in this region from West to East and the stratigraphic hiatus increases. An earliest Cambrian hiatus is widely distributed on the northern margin of the Yangtze Platform. At the same time, a syngenetic brecciation occurred in wide shallow shelf settings. A tectonic mobilization, possibly related to the assembly of Gondwana following the breakup of Rodinia, may be responsible for this. In sections near Kuanchuanpu (Ningqiang County), the Meishucunian (equivalent to the Nemakit-Daldynian to Tommotian of Siberia) succession comprises ca. 55 m of dark phosphatic carbonates, siltstones, and cherts. The SSF associations are similar to those of East Yunnan and, in ascending order, can be discriminated into the Anabarites trisulcatus - Protohertzina anabarica Zone, the Watsonella yunnanensis Zone (previously mentioned as Heraultipegma yunnanensis Zone), and the Sinosachites flabelliformis - Tannuolina zhangwentangi Zone. The Siphogonuchites triangularis - Paragloborilus subglobosus Zone cannot be unambiguously detected in the Southwest Shaanxi region. The occurrence of embryos and eggs within the Anabarites trisulcatus - Protohertzina anabarica Zone, which has been reported earlier, is remarkable. The previously reconstructed life cycle and taxonomic treatment of Olivooides / Punctatus are re-discussed. The earliest SSF associations (Anabarites trisulcatus - Protohertzina anabarica Zone to Watsonella yunnanesis Zone) are missing in the Southeast Shaanxi - Northwest Hubei region and the Ninella tarimensis - Cambroclavus fangxianensis Assemblage Zone and Rhombocorniculum cancellatum Taxon Range Zone are introduced herein.     

Proposed stratotype for the base of the highest Cambrian stage at the first appearance datum of Cordylodus andresi, Lawson Cove section, Utah, USA

We propose a candidate for the Global Standard Stratotype-section and Point (GSSP) for the base of the highest stage of the Furongian Series of the Cambrian System. The section is at Lawson Cove in the Ibex area of Millard County, Utah, USA. The marker horizon is the first appearance datum (FAD) of the conodont Cordylodus andresi Viira et Sergeyeva in Kaljo et al. [Kaljo, D., Borovko, N., Heinsalu, H., Khazanovich, K., Mens, K., Popov, L., Sergeyeva, S., Sobolevskaya, R., Viira, V., 1986. The Cambrian–Ordovician boundary in the Baltic–Ladoga clint area (North Estonia and Leningrad Region, USSR). Eesti NSV Teaduste Akadeemia Toimetised. Geologia 35, 97–108]. At this section and elsewhere this horizon also is the FAD of the trilobite Eurekia apopsis (Winston et Nicholls, 1967). This conodont characterizes the base of the Cordylodus proavus Zone, which has been recognized in many parts of the world. This trilobite characterizes the base of the Eurekia apopsis Zone, which has been recognized in many parts of North America. The proposed boundary is 46.7 m above the base of the Lava Dam Member of the Notch Peak Formation at the Lawson Cove section. Brachiopods, sequence stratigraphy, and carbon-isotope geochemistry are other tools that characterize this horizon and allow it to be recognized in other areas.     

Evolution of C isotopes in the Cambrian of China: implications for Cambrian subdivision and trilobite mass extinctions

Carbon and oxygen isotopes were studied in fossiliferous Cambrian carbonates in northwestern Hunan Province (South China) and in northern Anhui and southern Shandong provinces (North China). Two major C isotope excursions related to biological events occur in the Wangcun section (Yongshun County, northwestern Hunan), which consists of a slope carbonate sequence (510 m thick) containing abundant trilobites. The first C isotope excursion (δ¹³C value shifts from -2.3‰ to 2‰) occurs near the boundary between the Qingxudong and Aoxi formations, close to the traditional Lower-Middle Cambrian boundary. The second excursion (δ¹³C value shifts from 0‰ to 3‰) occurs in the interval between the Linguagnostus reconditus Zone and the Glyptagnostus reticulatus Zone. The base of the G. reticulatus Zone define the base of the Paibi Stage and Furongian Series. Similar C isotope excursions also occur in shallow - water carbonate sections in North China. In Jiagou section near Huainan (Anhui Province), recently considered an important interval for defining the lower-middle Cambrian boundary because of dramatic changes in the trilobite fauna (extinction of redlichiids and appearances of ptychopariids), a negative C isotope excursion (δ¹³C value shifts from +1.21‰ to -1.93‰) occurs at the top of the lower member of the Mantou Formation. In the Gushan section (Changqing County, Shandong Province), a C isotope excursion (δ¹³C value shifts from -0.04‰ to 2.23‰) occurs at the base of the Changshan Formation and is coincident with the base of the Chuangia Zone. This excursion can be correlated with the excursion in the lower part of Glyptagnostus reticulatus Zone in the Wangcun section. The above two distinct C isotope excursions, which occur both in slope carbonates in South China and in shallow - water carbonates in North China, have also been recognized in Cambrian sections on other continents, and they coincide with global mass extinctions of trilobites. The two excursions evidently reflect global changes of Cambrian sea level, and they have utility for Cambrian subdivisions and for both regional and global stratigraphic correlation. In addition, a negative carbon excursion below the base of the Ptychagnostus atavus Zone in the Wangcun section supports previous suggestions that the FAD of P. atavus can be considered as a global correlatable horizon within the middle Cambrian.     

Cambrian high-resolution biostratigraphy and carbon isotope chemostratigraphy in Scania, Sweden: first record of the SPICE and DICE excursions in Scandinavia

A core drilling (Andrarum‐3), from the classical locality at Andrarum, Scania, southernmost Sweden, penetrated a 28.90‐m‐thick Cambrian succession. The core comprises dark grey to black, finely laminated mudstones and shales with early concretionary carbonate lenses (stinkstones or orsten) and a few primary carbonate beds. The middle Cambrian (provisional Series 3) part of the core comprises 17.35 m, whereas the Furongian Series (upper Cambrian) part covers the remaining 11.55 m. Nineteen trilobite and two phosphatocopine genera are present in the middle Cambrian, whereas the less diverse Furongian interval yielded four trilobite and three phosphatocopine genera. Other, less frequent, faunal elements include conodonts (s. l.), brachiopods, sponge spicules, bradoriids, and coprolites. Trilobites and phosphatocopines were used to subdivide the core into seven biozones ranging from the Ptychagnostus atavus Zone to the Parabolina spinulosa Zone (P. spinulosa Subzone). Carbon isotopic analyses (δ¹³C_org) through the core show two important excursions, the negative DrumIan Carbon isotope Excursion (DICE) in the Pt. atavus Zone, and the Steptoean Positive Carbon Isotope Excursion (SPICE) beginning near the first appearance of Glyptagnostus reticulatus and extending upward into the Olenus and Agnostus (Homagnostus) obesus Zone. The DICE displays a peak value, in the samples at hand, of –30.45‰δ¹³C_org in the lower part of the P. atavus Zone. The δ¹³C_org values increase through the overlying L. laevigata and A. pisiformis zones and display peak values of c. –28.00‰δ¹³C_org in the lowermost Furongian Olenus wahlenbergi and O. attenuatus subzones. Thereafter the values decrease significantly through the O. scanicus Subzone. Both isotopic excursions have been documented from several palaeocontinents, but never before from Baltica. Moreover, for the first time these excursions are recorded from organic matter in an alum shale setting. The recorded shift of +1.50–2.00‰δ¹³C_org is approximately half the magnitude of the SPICE documented from other regions. This discrepancy may be related to temporal variations in the type, origin, or diagenesis of the organic fraction analysed.     

Evidence for seafloor microbial mats and associated metazoan lifestyles in Lower Cambrian phosphorites of Southwest China

The increase in the depth and intensity of bioturbation through the Proterozoic-­Phanerozoic transition changed the substrates on which marine benthos lived from being relatively firm with a sharp sediment-water interface to having a high water content and blurry sediment-water interface. Additionally, microbial mats, once dominant on normal marine Proterozoic seafloors, were relegated to stressed settings lacking intense metazoan activity. This change in substrates has been termed the 'agronomic revolution', and its impact on benthic metazoans has been termed the 'Cambrian substrate revolution'. The shallow marine phosphorites of the Lower Cambrian Meishucun Formation of southwest China contain evidence suggestive of the presence of seafloor microbial mats. This evidence includes abundant and distinctive red-colored bedding planes enriched in heavy iron minerals and mica, interpreted as resulting from mat-decay mineralization and mica trapping by microbial mats. The radular grazing trace fossil Radulichnus is also found in this formation, indicating a firm, microbial mat-bound substrate. These radular scratches are always preserved with circular impressions around 10 cm in diameter, possibly the fossils of soft-bodied organisms. The first relatively intense bioturbation in this region is found in this formation and is dominated by horizontal Thalassinoides burrows, which could represent undermat mining behavior. The evidence for the presence of microbial mats in the Lower Cambrian Meishucun Formation, and for metazoan lifestyles associated with such mat-bound seafloors, reveals that normal marine environments dominated by typical Proterozoic-style soft substrates still existed during the Cambrian substrate revolution.     

Lapworthellids and other skeletonised microfossils from the Cambrian Stage 3 of the northern Montagne Noire,southern France

The Montagne Noire (southern France) possesses one of the most complete Cambrian successions in the western peri-Gondwana margin and might provide a good stratigraphic reference for both regional charts and international correlations. However, to date, the lower Cambrian succession of the northern Montagne Noire has been supposed to be devoid of biostratigraphically significant fossils. The complex tectonostratigraphic framework of the area (a range divided into Axial Zone, northern and southern Montagne Noire) exacerbated problems related to regional correlations and palaeogeographic reconstructions. As a result, the chronostratigraphic context of the lower Cambrian of northern Montagne Noire is still uncertain and stratigraphic reports have broadly relied on putative lithostratigraphic correlations with the southern Montagne Noire. The purpose of this study is to characterise, for the first time, the fossil record of carbonate beds and lenses of the northern Montagne Noire occurring at the top of the siliciclastic-dominated Marcory Formation, in order to provide regional bio- and chronostratigraphic constraints on lower Cambrian strata. Moreover, this study is aimed at improving international chronostratigraphic correlation. Carbonate beds and lenses cropping out along the Orque Cliff, in the Mélagues thrust slice, were investigated. They yielded a faunal assemblage constituted of molluscs (Igorella cf. ungulata and Igorella moncereti n. sp.), hyoliths (Conotheca brevica), chancelloriids (Archiasterella cf. pentactina and Allonnia sp.) and tommotiids (Lapworthella rete). L. rete is recorded in upper Meishucunian (Cambrian Stage 3) strata of the Yangtze Platform (South China) where it is used as index fossil of the Cambrian Stage 3 Sinosachites flabelliformis–Tannuolina zhangwentangi Assemblage Zone. Therefore, the presence of this tommotiid provides evidence that the studied carbonate beds and lenses are Cambrian Age 3 (Atdabanian according to the Siberian chart). The upper part of the Marcory Formation in the Mélagues slice, dated as Cambrian Stage 3, might represent a lateral equivalent of the mixed (carbonate-siliciclastic) Pardailhan Formation defined in the southern Montagne Noire.     

Flume studies using 1:1 scale models of Series 2 and basal Series 3 Cambrian gogiid eocrinoids from Guizhou Province,China to determine feeding posture and mode of attachment

In Series 2 and 3 Cambrian of Guizhou Province, China, most echinoderms inhabited deeper/quieter water and were attached directly to siliciclastic substrate or biodetritus by biogluing (extrusion of extensible collagen). Feeding postures of abundant long stalked gogiids (e.g., Sinoeocrinus) from these beds were interpreted to have heeled over in the current from the thin flexible distal end of the stalk, with the brachioles streaming in a loose bundle, down current from the theca. To test these and other feeding posture assumptions, 1:1 scale models (holdfast, stalk, and theca) of three genera were carved from soft rubber and brachioles were modeled from braided fishing line. By varying current velocities long stalked flume models did not significantly heel over. Brachioles, both straight and spiraled, extended vertically from the theca in an (elliptical) cone and distally curved downstream. Disrupted flow around straight brachioles (Sinoeocrinus) kept them somewhat evenly spaced. Spiraled brachioles (Guizhoueocrinus, Globoeocrinus) are initially straight and angle outwards so that each proximal end defines a sector over the theca; this spacing keeps the brachioles free from tangling distally. Biogluing the animal to the bottom or to biodetritus seems to be correctly interpreted from the morphological evidence. Superglue was used as the proxy gluing agent for the models, success was limited. The dewatered, siliciclastic, non-bioturbated, seafloor could be only partly reconstructed and the somewhat viscous glue did not deeply penetrate the illite substrate. It is probable that bioglue had low viscosity, penetrated the sediment easily, and was able to agglutinate a large three dimensional anchoring body of sediment without (as is commonly observed) disrupting bedding.     

Crypto-helical body plan in partially disarticulated gogiids from the Cambrian of South China

All living echinoderms have a pentaradial symmetry that is unique within the Bilateria. However, the Cambrian origin of echinoderm radial/pentaradiate symmetry is a long-standing problem. During the Cambrian (542–488 Ma), gogiids were the most common stalked echinoderm characterized by an “irregularly” plated body. Based on recently discovered material from the Balang Formation (Cambrian Series 2), eastern Guizhou, China, three unusual, partially disarticulated specimens of Guizhoueocrinus have clear evidence for a helical body plan. This helical plating is only evident in partially disarticulated specimens, thus a crypto-helical body construction is present. Crypto-helical construction in a gogiid raises the possibility of a phylogenetic connection among helicoplacoids, gogiids, and Helicocystis. The crypto-helical body construction may be an important evolutionary innovation among pre-radiate echinoderms.     

英文贵州中寒武世凯里生物群中宏观藻类化石新材料(英文)

贵州省台江县中寒武世凯里生物群含有丰富的非钙质藻类和具有软躯体后生动物化石 ,它为布尔吉斯页岩型生物群在世界广泛分布提供了更有力的证据。在生物群的宏观藻类中描述了 5个属 5个种 ,包括 2个新属。它们是MarpoliaspissaWalcott、AcinocricusstichusConwayMorrisandRobison、UdotealgaerectaYang、EosargassumsawataYang和RhizophytonzhaoyuanlongiiYang ,并且将凯里生物群中的宏观藻类化石组合与加拿大布尔吉斯页岩生物群中的宏观藻类进行了对比 ,发现两个生物群不仅具有相似的动物化石组成 ,而且宏观藻类化石组成也很相似。     

龙凤山藻在澄江早寒武世生物群中的发现

主要报道了产自云南昆明海口耳材村早寒武世筇竹寺组玉案山段澄江生物群中可能为漂浮生长的宏观藻类化石-心型龙凤山藻（新种）（Longfengshania cordata sp.nov.)和中华豆芽藻（新属、新种）（lantulaformis sinensis gen.et sp.nov)。这些化石的发现进一步显示了澄江生物群种物多样性,为揭示该生物群爆发性演化提供了新的证据。通过对龙凤山藻属亲缘关系的深入比较研究,进一步证明该类化石应归属宏观藻类以及它们遗传上的稳定性、演化上的保守性和环境上的适应性。     

Aspects of recent advances in the Ordovician stratigraphy and palaeontology of China

The continuous and richly fossiliferous Ordovician succession of China (particularly South China) comprises a heterogeneous suite of litho- and bio-facies, which has been a main focus of stratigraphical and palaeontological research in recent years. Among the seven GSSPs established in China, three are within the Ordovician System, and the GSSP of the Darriwilian Stage at Huangnitang, Changshan County, western Zhejiang Province, was the first “golden spike” in China and the first for the Ordovician System. A series of case studies have revealed that: (1) the Ordovician radiation of some fossil groups on the Upper Yangtze Platform (e.g., brachiopods and graptolites) reached their first α-diversity acme in the Didymograptellus eobifidus Biozone, four zones earlier than the global trend; (2) the β-diversity peak was attained 3–4 zones later than the α-diversity peak; (3) many brachiopod communities or faunas first occurred in the central part of the Upper Yangtze Platform and subsequently expanded to both more offshore and near-shore facies; (4) diachroneity existed in many aspects of the radiation. The end-Ordovician mass extinction was a severe event in South China. Two pulses of the extinction are recognized for a number of major fossil groups, some being most strongly affected during the first pulse whereas the others suffered during the second pulse. Macroevolution during the Ordovician–Silurian transition has been investigated in detail, and the role of the Lazarus effect has been found to be less important than previously believed.     

Unique lowermost Cambrian small shelly fossils (SSFs) from eastern Yunnan,South China: implications for the early diversification history of animals

With a particular focus on the earliest Cambrian diversification of small shelly fossils (SSFs), stratigraphic analysis was conducted on the lower Cambrian Zhongyicun Member at the Hongjiachong section in the Chengjiang area, Yunnan, China. From ca. 3-m-thick bedded phosphorites (Unit A) in the lower part of the member, we recovered unique SSFs. This interval with 4 SSF-bearing horizons is characterized by the dominance in blade-shaped SSFs, including Halkieria spp., Brushenodus prionodes, Sinosachites delicatus, and Pteronus sp., and the absence of typical tube/cap-shape SSFs common in the Fortunian. This interval tentatively named “Halkieria-dominant interval” is stratigraphically positioned ca. 3 m below the previously known level of the oldest mollusks in the continuous bedded phosphorite sequence between the Anabarites trisulcatus-Protohertzina anabarica Assemblage Zone and the overlying Paragloborilus subglobosa-Purella squamulosa Assemblage Zone of the Fortunian (the earliest Cambrian). The “Halkieria-dominant interval” yields some SSFs likely affiliated with ostracods in view of size and morphology, suggesting that arthropod body fossils appeared in the Fortunian, considerably earlier than previously believed.