On the relative frequencies of hominid maxillary and mandibular teeth and jaws as taphonomic indicators

In southern African samples of early hominid remains, maxillary and mandibular teeth (deciduous-plus-permanent) have a virtually equal chance of accumulating in the dolomitic limestone cave deposits, of being preserved therein and recovered therefrom. Thus, of 1066 fossil teeth ofAustralopithecus spp. plusHomo habilis, 51.9 per cent are maxillary and 48.1 per cent mandibular. On the other hand, the East African sample of 847 early hominid, deciduous-plus-permanent teeth, departs more strikingly from a 1∶1 ratio: it comprises 41.0 per cent maxillary and 59.0 per cent mandibular teeth. It is inferred that mandibular teeth have a somewhat better chance of accumulating and being preserved in, and being recovered from, the open, fluvial, lacustrine and deltaic sedimentary environments of the East African sites. The dental proportions are approximately matched by the proportions of jaws. For example, the maxilla: mandible proportions at Koobi fora in northern Kenya are 33.0∶67.0 for teeth and 21.6∶78.4 for jaws. In other words, the preponderance in favour of mandibular remains is somewhat more marked in the case of jaws than of teeth, this distinction doubtless reflecting the more fragile bony structure of the maxilla and the sturdier construction of the mandible. This first study known to the author of the differential distribution of maxillary and mandibular teeth of the Plio-Pleistocene hominids leads the author to hypothesize that, where environmental conditions at the place and time of the death of the hominids have been non-destructive, non-dispersive, relatively mild and protective, maxillae and mandibles may be expected to have been conserved and recovered in approximately equal proportions—and likewise of maxillary and mandibular teeth. On the other hand, the more brutal and destructive the sedimentary environment and other taphonomic influences have been, at the place and time when the hominid individuals died, the more likely it is that the maxillary and mandibular remains of jaws and teeth will deviate from equality of proportions, generally at the expense of the maxillae and upper teeth. Hence, it is proposed that the upper jaw/low jaw ratio (Mx/Mn jaw ratio) and the maxillary teeth/mandibular teeth ratio (Mx/Mn dental ratio) may serve as two useful new gauges of the rigour of palaeo-ecological and taphonomic conditions.     

Hominid evolution in Northwest Africa and the question of the North African "Neanderthaloids"

The North African evidence bearing on hominid evolution is summarized and reviewed in relation to its industrial associations and apparent chronological order. Three tacit suppositions of the past and their effects on taxonomy and thinking in evolutionary anthropology are examined, namely the concepts of a single and unique origin of Man, of the inherent and essential, distinction between physical and spiritual Man, and of physical resemblance as absolute proof of relationship. The consequent confusion and looseness of application which have characterized hominid taxonomy, especially concerning the term Neanderthal and its derivatives, are examined and discussed primarily in relation to the so-called Neanderthals of northern Africa. It is suggested that the Atlanthropus remains may represent not one but two hominid varieties. Further it is argued that none of the North African remains are demonstrably related to the European Neanderthals and that some at least of them seem clearly unrelated. In conclusion it is suggested that the hominid remains from Ternifine, the Littorina Cave (Casablanca), Témara, Rabat, Jebel Irhoud and Tangier may represent links in a North African chain of hominid evolution which developed locally and remained independent of all external influence during the period covered by those specimens.     

A mandible ofBranisella boliviana (Platyrrhini,Primates) from the oligocene of South America

A mandibular specimen from the Bolivian Early Oligocene is provisionally assigned toBranisella boliviana. The crown anatomy of the single preserved tooth, an M₂, indicates platyrrhine affinities and several details of the broken jaw are suggestive of symphyseal fusion. Like the African Oligocene parapithecids,Branisella contrasts with extant anthropoids in the relative shallowness of its mandible.Branisella is the most ancient, and seemingly the most primitive, fossil platyrrhine monkey, lacking any of the derived features of the two major clades of modern ceboids. Taxonomically, it is best regarded as family incertae sedis.     

Ontogenetic features of <Emphasis Type="Italic">Dolichopithecus</Emphasis> (Primates,Colobidae) from the Pliocene of Pridnestrovie

In the permanent dentition of the extinct genus Dolichopithecus, M₁, I₁, and I₂ were the first to erupt, followed by M₂, canine, P₄, and P₃. M₃ was the last permanent tooth to erupt. At the stage of eruption of P₄ and P₃, M₃ was incompletely mineralized. The difference from the extant Cercopithecoidea is the loss of all deciduous teeth after eruption of incisors and M1 and the similarity is observed in the succession of eruption of permanent teeth. In Dolichopithecus, the lower jaw body retained constant in thickness after eruption of M₂. The lower jaw increased in length and depth, as the horizontal ramus grew with the formation and eruption of M₃.     

Recent human evolution in East Asia and Australasia.

In both East Asia and Australasia arguments for evolutionary continuity between middle-late Pleistocene hominid populations and modern Homo sapiens are of long standing. In both regions, however, problems of chronological distribution, dating and preservation of hominid skeletal materials provide an effective barrier to extending regional sequences back to 'archaic' Homo sapiens or Homo erectus. The earliest securely dated modern Homo sapiens in East Asia are currently represented by Zhoukoudian Upper Cave at a minimum of 29 ka BP. In Australia skeletal remains of modern Homo sapiens have been dated to 26 ka BP, with archaeological materials at 38 to 50 ka BP. Late Pleistocene human skeletons from sites like Coobool Creek are morphologically and metrically outside the range of recent Australian Aboriginal populations. Similarly Liujiang and the Upper Cave crania can be distinguished from recent East Asian 'Mongoloids'. Evolutionary change within the Holocene needs to be taken into consideration when the evidence for regional evolutionary continuity is considered.     

Hominid molars from a Middle Stone Age level at the Mumba Rock Shelter, Tanzania

Three hominid molars were recovered from a depth of 7.0-7.1 meters in the Mumba Shelter at Lake Eyasi, northern Tanzania. Geological context of the finds and archaeological data indicate that people with a Middle Stone Age technology were using the Mumba locality intermittently whenever retreat of lake waters allowed access to the site. Uranium series dates suggest an age on the order of 130,000 years bp for the teeth and stone tools. Based on morphological analyses, the dental remains probably belonged to one individual and appear to be the crowns of two upper permanent M2s and one lower permanent M2. Crown areas are very small, even in comparison to the variation exhibited by recent African populations. Crown patterns have no archaic features. These teeth are smaller than any verifiable archaic Homo sapiens examples; thus, they may represent early anatomically modern Homo sapiens.     

江苏镇江莲花洞动物化石铀系年代

莲花洞位于镇江市南约 6ｋｍ的白龙岗山北坡 (原在丹徒县境内 ) ,地理座标 119°2 4′Ｅ ,32°10′Ｎ。白龙山为宁镇山脉东支 ,走向近东西 ,顶峰海拔 118ｍ。溶洞发育于三迭纪青龙组灰岩 ,洞口标高约 35ｍ。 1981年 ,南京博物院等单位[1 ] 对该地点进行发掘 ,获哺乳动物化石16种 ,其中人牙化石 1枚。洞内堆积自下而上被分为 4层 ,其中第 2层为棕红色亚粘土 ,含石灰岩碎块、铁锰质结核和大量动物化石。发掘者根据动物化石中有 3个绝灭种 ,将该地点定在“晚更新世或晚更新世早期” ,人牙化石被认为属于智人。韩德芬等[2 ] 认为莲花洞在我国晚…     

Australopithecus, Meganthropus and Ramapithecus

The South African members of Australopithecus form a single group, trending from earlier, more gracile or smaller forms, to later, more robust or larger forms, in accordance with the “Law of Cope”. This is supported by the evidence of the lower first deciduous molar: it is only slightly molarized in the earlier, smaller forms and astonishingly heavily molarized in the later, robust forms, such as that of Kromdraai. Hence, Robinson's view that two different genera are represented by the gracile and robust forms is not supported here. A. robustus is seen as a late offshoot of the Australopithecus stem. The resemblance of the Taung dm₁ to that of Sinanthropus, except for the more differentiated talonid of the Taung specimen, suggests that the separation of the Australopithecinae and Homininae must have taken place at an earlier stage than that represented by the oldest South African Australopithecinae. The lower jaw of Meganthropus of Java combines certain characteristics of A. africanus with those of A. robustus: Meganthropus might provisionally be called “Australopithecoid”. The geographically intermediate India has yielded a hominid, Ramapithecus punjabicus, but the author does not consider “Kenyapithecus” to be a hominid. “K. wickeri” is a pongid species of its own and “K. africanus” a Proconsul. Ramapithecus sensu stricto is known only from the Indian Siwaliks and the author suggests that the transition from Ramapithecus to a still unknown Australopithecus took place in the same region prior to their migration into Africa and Southeast Asia.     

Paranthropus boisei: An example of evolutionary stasis?

Of the presently recognised early hominid species, Paranthropus boisei is one of the better known from the fossil record and arguably the most distinctive; the latter interpretation rests on the numbers of apparently derived characters it incorporates. The species as traditionally diagnosed is distributed across approximately one million years and is presently confined to samples from East African sites. The hypodigm has been examined for evidence of intraspecific phyletic evolution, with particular emphasis on the areas best represented in the fossil record, namely the teeth and mandible. The results of this examination of 55 mandibular and dental variables, which uses the Γ test statistic for the detection of trend, and nonparametric spline regression (Loess regression) for investigating pattern and rate of temporal change, show that within Paranthropus boisei sensu stricto most evidence of temporally related morphological trends relates to the morphology of the P₄ crown. There is little or no evidence of any tendency to increase in overall size through time. Fossils from the Omo Shungura Formation and West Turkana which have been recovered from a time period earlier than the P. boisei sensu stricto hypodigm resemble the latter taxon in some features, but differ from it in aspects of cranial morphology. There is insufficient fossil evidence of the earlier taxon to tell whether it changes through time, but when trends of 47 mandibular and dental variables are assessed across the combined East African “robust” australopithecine sample, there is evidence for a relatively abrupt change around 2.2–2.3 Myr in approximately 25% of the dental and mandibular remains. Some of these changes correspond with the temporal trends within P. boisei sensu stricto, but others, such as mandible height, do not. If the earlier material is ancestral to P. boisei sensu stricto, evidence from the teeth and jaws is consistent with a punctuated origin for the latter taxon. © 1994 Wiley-Liss, Inc.     

Dental anomalies and dental variations in the red foxVulpes vulpes in the Czech Republic

Dental abnormalities including polydonty, oligodonty, extra roots, different root morphotype, root fusion, different crown morphotype, crown reduction, partial crown eruption, supernumerary cusp, irregularities in the position of the teeth, and malocclusion were studied in a set of 785 red foxVulpes vulpes (Linnaeus, 1758) skulls (401 males, 273 females, and 111 individuals of unknown sex) from the Czech Republic. Three hundred sixty one cases of deviations from normal were found in 170 specimens (21.7%). Most of the deviations were variants within a genetically determined range. The prevalent dental variants included an extra root of M¹ (5.7% specimens), and different root morphotype of P₁ (1.9% specimens). On the other hand, the real dental anomalies, eg polydonty, occurred seldom within the population. P₁ ¹ and M₃ were missing significantly more frequently among females than among males unlike the other deviations, which were divided equally between the sexes. No differences were found between the left and right side of the jaw. Irregularities in the position of the teeth and oligodonty (excluding P¹ ₁, M₃) appeared significantly more abundantly on mandible, whereas extra roots and polydonty were more common on the maxilla. There was no relationship between the incidence of dental abnormalities and the relative mandible and rostrum length.     

Re-analysis of the hominid radii from Cave of Hearths and Klasies River Mouth, South Africa

Two of the few postcranial fragments from the late Early Stone Age and/or the Middle Stone Age of southern Africa are the proximal radii from the Cave of Hearths and Klasies River Mouth. The Cave of Hearths fossil is metrically indistiguishable from both archaic (e.g., Neandertals) and recent humans, and presents a mosaic of primitive and modern features. The primitive include a relatively slender neck and thick cortical bone (the latter of which distinguishes recent humans from archaic, Early Modern, and Upper Paleolithic hominids); the modern includes an anteromedially (rather than medially) facing radial tuberosity. Its extreme collo–diaphyseal angle is unusual, although it can be matched by modern homologues. The neck–shaft angle of some Neandertal and Early Modern radii also appears to match that of the Cave of Hearths specimen. The Klasies River Mouth radius also has thick cortical bone of the neck. It is morphologically indistinguishable from Early Modern and Neandertal homologues. These, and other fossils, suggest a mosaic pattern of evolution in the postcranial skeleton of the late Early Stone Age and/or Middle Stone Age inhabitants of sub-Saharan Africa.     

The mixed dentition and associated skull fragments of a juvenile fossil hominid from Sterkfontein,South Africa

In April–May 1983, the late A.R. Hughes and his field team recovered more than 40 bone fragments and teeth from a single solution pocket of the Sterkfontein Formation. After preparation and reconstruction by JMC, it was recognised that these fragments represent a single juvenile individual (Stw 151), consisting of more than 40 cranial and dental parts, with mixed dentition. It constitutes the most complete set of jaws and teeth of an early hominid child since the Taung child was recovered in 1924. In this paper, the morphological and metrical features of the individual teeth are described. The other associated skull fragments (right ramus of the mandible, left petrous bone, right glenoid region) are also described. Comparisons are made with other South (and East) African fossil hominids. The beautiful preservation simultaneously of most of the deciduous teeth and of the permanent teeth exposed in their crypts allows an accurate analysis of the developmental sequence. A report on the dental developmental status of this juvenile is presented. On the basis of the microanatomical study of the developing permanent teeth, the estimated age at death is 5.2–5.3 years. Reconstructions of the maxillary and mandibular arcades are also offered. The morphological and metrical features of Stw 151 raise the possibility that it may represent a hominid more derived towards an early Homo condition than the rest of the A. africanus sample from Member 4. Am J Phys Anthropol 106:425–465, 1998. © 1998 Wiley-Liss, Inc.     

贵州兴义猫猫洞出土的人类化石

本文主要是对贵州兴义猫猫洞遗址出土的人化石进行的初步研究,并对哺乳动物化石、地层、堆积物作叙述。该遗址的人类化石包括下颌骨4件,股骨3段,是中国南方洞穴遗址中发现的数量较多,系统分类地位较清楚的晚期智人的化石。人类化石同层的鹿牙作铀系法测年结果为14500±1200 BP,相当于更新世的末期。     

云南西畴仙人洞动物化石铀系年代

本文报道晚期智人地点云南西畴仙人洞动物牙化石的铀系测年结果。4个样品的230Th234U年龄范围为47—105ka,测定了其中2个样品的227Th/230Th年龄,结果与230Th/234U法的一致。在没有进一步的地层和年代证据的情况下,将西畴人牙化石的年代暂定为47—105ka或许是合理的。但洞穴地点骨化石铀系年代总体偏年轻,上述年代范围很可能被低估。本文结果与我们已有研究的广西通天岩、前洞等地点一致,是晚期智人在我国出现比原认为的早得多的又一例证。     

Taxonomic affinities and evolutionary history of the Early Pleistocene hominids of Java: dentognathic evidence

Temporal changes, within-group variation, and phylogenetic positions of the Early Pleistocene Javanese hominids remain unclear. Recent debate focused on the age of the oldest Javanese hominids, but the argument so far includes little morphological basis for the fossils. To approach these questions, we analyzed a comprehensive dentognathic sample from Sangiran, which includes most of the existing hominid mandibles and teeth from the Early Pleistocene of Java. The sample was divided into chronologically younger and older groups. We examined morphological differences between these chronological groups, and investigated their affinities with other hominid groups from Africa and Eurasia. The results indicated that 1) there are remarkable morphological differences between the chronologically younger and older groups of Java, 2) the chronologically younger group is morphologically advanced, showing a similar degree of dentognathic reduction to that of Middle Pleistocene Chinese H. erectus, and 3) the chronologically older group exhibits some features that are equally primitive as or more primitive than early H. erectus of Africa. These findings suggest that the evolutionary history of early Javanese H. erectus was more dynamic than previously thought. Coupled with recent discoveries of the earliest form of H. erectus from Dmanisi, Georgia, the primitive aspects of the oldest Javanese hominid remains suggest that hominid groups prior to the grade of ca. 1.8-1.5 Ma African early H. erectus dispersed into eastern Eurasia during the earlier Early Pleistocene, although the age of the Javanese hominids themselves is yet to be resolved. Subsequent periods of the Early Pleistocene witnessed remarkable changes in the Javanese hominid record, which are ascribed either to significant in situ evolution or replacement of populations.     

First record of Macaca mulatta (Cercopithecidae: Papionini) from the Taedong River Basin,the Democratic People’s Republic of Korea

The macaque fossils, including mandible fragments, were discovered for the first time from the Taedong River Basin around Pyongyang (TRBP) (the Chongphadae Cave, Hwangju County; the Taehung Cave, Sangwon County; the Maeri Cave, Sungho County), the Democratic People’s Republic of Korea, which represent the first recognized occurrences of fossil M. mulatta in DPRK. Although the only available skull specimens are the mandible including the teeth, the materials display some mandibular and dental feature combinations that might belong to the known species of Macaca mulatta. This discovery indicates that TRBP may be the richest area of Macacina fossils and an evolutionary center for macaques in DPRK. Radiometric dating of the Chongphadae Cave sediment Layer 12 and Layer 13 yielded 34770–27800 and 24980–21340 cal yr BP respectively.     

古人类遗址湖北郧西黄龙洞的铀系年代

湖北郧西黄龙洞为近年来发现的晚期智人遗址。本文报道该地点具明确层位意义的洞穴次生碳酸盐岩和骨化石样铀系测年的结果。含文化堆积下伏一局部钙板的年代为约100ka BP, 其中部偏上一局部钙板为约77ka BP, 表层钙板形成于27—57ka BP间。人类化石和石制品出土于文化堆积的底部, 其年代应在57—100ka BP, 并很可能在77—100ka BP间。与人牙化石同层的四枚犀牛牙化石在35—72ka BP间, 与基于次生碳酸盐岩的年代框架没有冲突。本文结果为中国现代人类的早期出现和距今40—100ka BP间有人类活动提供了有力证据。     

广西崇左木榄山智人洞的鼠科化石

最近在广西崇左木榄山智人洞遗址采集到1件具有现代智人解剖特征初始状态的下颌骨和丰富的哺乳动物化石。啮齿类中的鼠科化石有锡金小鼠(Mus pahari)、中华姬鼠(Apodemus draco)、黑线姬鼠(Apodemus agrarius)、似德氏狨鼠(Hapalomys cf.H.delacouri)、笔尾树鼠(Chiropodomys gliroides)、社鼠(Niviventer confucianus)、针毛鼠(Niviventer fulvescens)、爱氏巨鼠(Leopoldamys edwardsi)、印度板齿鼠(Bandicota indica)、褐家鼠(Rattus norvegicus)和黄毛鼠(Rattus losea)共8属11种。这种全部由当地或邻近地区现生种类构成的组合显然比当地早更新世的三合大洞和川黔地区中更新世歌乐山期的鼠类组合进步, 而与广西田东雾云洞的鼠类组合相似, 其动物群的时代为晚更新世早期, 与不平衡铀系法得到的测年结果(距今约11万年)相吻合。智人洞鼠科动物的组合明显具有东洋界热带-亚热带动物群的特点, 其中林灌和草地型所占比例较大表明当时的森林面积可能减少、林灌和草地面积可能增加, 反映出当时的气候相对干旱。     

中国早期智人牙齿化石

中国目前较为可靠的早期智人牙齿材料应包括在巢县、许家窑、长阳、周口店新洞和桐梓发现的人类牙齿化石,代表一类与早期组早期智人相当或十分相近的古人类。丁村人类牙齿化石可能是代表晚期组早期智人也可能是代表解剖学上的现代智人。就牙齿材料而言,尚无充分的证据能表明在中国曾同时存在过两种类型的早期智人。     

The current state of korean paleoanthropology

The hominid fossil and Paleolithic archaeology records from the Korean Peninsula are extensive, but relatively little is known about the Korean human evolutionary record outside this region. The Korean paleoanthropological record is reviewed here in light of major research issues, including the hominid fossil record, relative and chronometric dating, lithic analysis, hominid subsistence, and the presence of bone tools, art and symbolism. Some of the major conclusions drawn from this review include: (1) hominid fossils have been found in nine separate sites on the Korean Peninsula; (2) possible Homo erectus fossils are present in North Korea; (3) Ryonggok Cave, in North Korea, has exposed the remains of at least five archaic Homo sapiens individuals; (4) a possible burial of an anatomically modern Homo sapiens child, discovered in Hungsu Cave in South Korea, has been tentatively dated to roughly 40,000 years ago; (5) handaxes and cleavers have been found at a number of sites near Chongokni and they appear to date to at least 100,000 years ago; and (6) taphonomic studies are necessary for addressing issues related to determining the nature of hominid-carnivore interaction over similar resources (e.g. carcasses and shelter); and the presence/absence of Early Paleolithic bone tools, art, and symbolism in Korea.