ANGIOSPERM WOOD FROM THE UPPER CRETACEOUS OF CENTRAL CALIFORNIA: PART II

Three species of fossil wood representing two genera are described. The specimens are from a collection of woods from the Upper Cretaceous Panoche formation of central California. Tetracentronites panochetris sp. nov. resembles angiosperm wood in ray structure and vascular pitting but lacks vessels. Plataninium platanoides sp. nov. is similar to the wood of Platanus, and the evidence presented points to a direct relationship. The resemblance between Plataninium cali-fornicum sp. nov. and the woods of certain Icacinaceae is discussed, but evidence of relationship is inconclusive.     

ANGIOSPERM WOOD FROM THE UPPER CRETACEOUS OF CENTRAL CALIFORNIA: PART I

This paper is the first in a series about a collection of 40 Upper Cretaceous woods from central California. The wood is in the form of fragments of stems and roots embedded in nodules of collophane rich in microorganisms, pollen, and spores. Two new species are described: Ulminium pattersonensis and Ulminium mulleri. Evidence presented indicates that these two specimens are similar to woods of the Lauraceae and fall well within the range of variation of woods of that family. Because the taxonomy of the Lauraceae is in need of revision, meaningful comparisons of fossil plant parts with modern taxa were found to be unfeasible.     

PALEOCENE DICOTYLEDONOUS TREES FROM BIG BEND NATIONAL PARK,TEXAS: VARIABILITY IN WOOD TYPES COMMON IN THE LATE CRETACEOUS AND EARLY TERTIARY,AND ECOLOGICAL INFERENCES

Paraphyllanthoxylon abbottii n. sp. and cf. Plataninium haydenii Felix from the Paleocene Black Peaks Formation, Big Bend National Park, Texas, are the first Paleocene dicotyledonous woods described from North America. Both represent wood types common in the Cretaceous. There are 30 logs of Paraphyllanthoxylon abbottii; it is rare that a single locality has such as large number of petrified dicotyledonous logs with a similar structural pattern, and the variability in mature wood structure can be documented. Paraphyllanthoxylon abbottii has a combination of features that occurs in many dicotyledonous families, but it is most similar to genera of Burseraceae. The Big Bend Paraphyllanthoxylon trees lack distinct growth rings, which suggests they grew in a climate without marked seasonality; they have high (10–74) vulnerability indices; such high values occur in extant tropical trees. The type species of Paraphyllanthoxylon, P. arizonense Bailey was reexamined, and its quantitative features are described. Aplectotremas Serlin of Albian age from the Edwards Limestone has anatomy like Paraphyllanthoxylon, and most probably is wood from a tree. The wood designated cf. Plataninium haydenii Felix resembles extant Platanaceae but differs in having exclusively scalariform perforation plates. Comparison of this wood with other platanoid woods suggests that in platanoid woods there has been a shortening of vessel elements and a decrease in the frequency of scalariform perforation plates from the Cretaceous through the Tertiary. These changes are consistent with the Baileyan model for specialization in tracheary elements.     

New species of <Emphasis Type="Italic">Piceoxylon</Emphasis> Gothan (Pinaceae) from the Cretaceous and Paleogene of the northwestern Kamchatka Peninsula

New fossil species of Piceoxylon (Pinaceae), P. talovskiense sp. nov. and P. kamtschatkiense sp. nov., are described on the basis of wood anatomy from the Cretaceous and Paleogene (respectively) of the Talovka River basin in the northwestern Kamchatka Peninsula. The Late Cretaceous P. talovskiense sp. nov. shows wood characters of modern Picea. The wood anatomy of the Paleogene Piceoxylon kamtschatkiense sp. nov. is somewhat similar to those of modern Picea sitchensis and P. jezoensis. Fossil woods of Piceoxylon have been found in the Kamchatka Peninsula for the first time.     

Fungal diversity and deterioration in mummified woods from the ad Astra Ice Cap region in the Canadian High Arctic

Non-permineralized or mummified ancient wood found within proglacial soil near the ad Astra Ice Cap (81°N, 76°W), Ellesmere Island, Canada was investigated to ascertain the identification of the trees, current morphological and chemical characteristics of the woods and the fungi within them. These woods, identified as Betula, Larix, Picea and Pinus, were found with varying states of physical and chemical degradation. Modern microbial decomposition caused by soft rot fungi was evident and rDNA sequencing of fungi obtained from the samples revealed several species including Cadophora sp., Exophiala sp., Phialocephala sp., as well as others. Analytical ¹³C-labeled tetramethylammonium hydroxide thermochemolysis showed the lignin from the ancient wood was in a high degree of preservation with minor side chain alteration and little to no demethylation or ring hydroxylation. The exposure of these ancient woods to the young soils, where woody debris is not usually prevalent, provides carbon and nutrients into the polar environment that are captured and utilized by unique decay fungi at this Arctic site.     

Leaf-shape as an indicator of introgression between Betula pendula and B. pubescens

Summary

Leaf-shape comparisons were made between individual trees in a mixed wood containing B. pendula and B. pubescens, and two relatively pure stands. Data sets were subjected to analysis of variance, principal components and discriminant analyses in order to study within-tree variation, variation within each species and the relationships between them. Within-tree variation was found to be much less than that between trees and principal component analysis provided a separation of the material into species groups and indicated extensive variation within them. The employment of a discriminant function sharpened the separation of the pure woods and pointed to the occurrence of possible hybrids and back-crossed individuals within the mixed wood. The findings indicate that gene flow probably proceeds from the diploid to the tetraploid level.     

Local extinctions and recolonisations of passerine bird populations in small woods

This paper considers, for eight species of woodland bird, the factors that influenced both local extinctions and recolonisations in 145 woods over 3 years. In all species, probability of local extinction was inversely related to population size; most local extinctions occurred in woods containing one to three breeding pairs. However, considerable variation in extinction probabilities occurred between species and between years. In addition, the suitability of habitat within a wood (more extinctions in less suitable woods) was important for wren Troglodytes troglodytes, song thrush Turdus philomelos and blue tit Parus caeruleus; also, the structure of the surrounding landscape was important for blue tit, great tit Parus major, and chaffinch Fringilla coelebs (more extinctions in localities with less woodland). In only two species was the probability of recolonisation related to any of the measured variables. Wrens were more likely to recolonise larger woods, whereas song thrushes were more likely to recolonise woods with a high habitat suitability rating and those which are more isolated from other woodland     

Durch Protophytobia cupressorum gen. nov., sp. nov. (Agromyzidae,Diptera) verursachte Markflecke in einem Holz von Juniperoxylon aus dem Tertiär von Süd-Limburg (Niederlande) und der Nachweis von Markflecken in einer rezenten Callitris-Art

From a cupressaceous wood described by Kräusel & Schönfeld (1974) of the Tertiary of South-Limburg (Netherlands) Greguss (1970) selected such pieces showing remains of insect food. In a recent Callitris species from Australia he found the some phenomenon (1970). Indeed Greguss considered these phenomena as a natural specific wood element. Really these are pith flecks are caused by a cambium-mining insect. Both are described in the present paper. I name the pith flecks in the fossil coniferous wood of South-Limbourg Protophytobia cupressorum gen. nov., sp. nov. Under such name is considered an insect only represented by their characteristic food channals. In living angiospermous woods Phytobia Lioy (Agromyzidae, Diptera) is the only genus hitherto known. In coniferous woods recent species are unknown. The food channals in conifers in great part closed by changed tracheids.     

Fossil Robinia wood from the western United States

A collection of petrified wood from the Lower Pliocene Ogallala Formation in western Oklahoma was examined. All specimens appear to be of the same taxon and exhibit features of extant Robinia species. To date, four fossil wood species of Robinia have been described. The relationship of Robinioxylon zuriensis Falqui to Robinia is doubtful because of the lack of diagnostic critical features. The remaining three, Robinia alexanderi Webber, Robinia breweri Prakash, Barghoorn and Scott, and Robinioxylon zirkelii (Platen) Müller-Stoll and Mädel do show affinity to Robinia and all have been noted as structurally similar to R. pseudoacacia. The Oklahoma woods and these three fossil species show considerable overlap in quantitative features and are identical in qualitative features. Examination of different sections (and specimens) of extant Robinia pseudoacacia wood reveals quantitative and qualitative variation similar to that found amongst the petrified woods. Robinia alexanderi, Webber, R. breweri Prakash, Barghoorn and Scott, R. zirkelii (Platen) Müller-Stoll and Mädel, and the Oklahoma specimens are considered to be conspecific as the differences between these fossil wood species are no different from those accounted for by variation within a single living species, R. pseudoacacia.     

Anatomy and affinities of the petrified plants from the teriary of Chile (VI)

Three new species and one new genus of dicotyledonous woods,Chilechicoxylon microporosum gen. et sp. nov.,Proteoxylon patagonicum sp. nov. andLaurinium beilschmiedioides sp. nov., are described from the Tertiary (or in case of the first and last species from the Cretaceous-Tertiary) of Chile Chico, XI Region, Chile. Previously only wood of temperate species has been identified from Patagonia.Proteoxylon patagonicum (Proteaceae) is the first recorded tropical wood species from this geographical area. Consecutive number from the previous paper (M. Nishida, H. Nishida and T. Nasa, 1988). Contributions from the Laboratory of Phylogenetic Botany, Chiba University No. 120 and contributions from Chiba University Botanical Expedition to the Andes No. 20. Supported by a Grant-in-Aid for Overseas Scientific Research from the Ministry of Education, Science and Culture Nos. 61041015 and 63041034.     

WOOD OF GINKGO IN THE TERTIARY OF WESTERN NORTH AMERICA

Scott , R. A., E. S. Barghoorn , and U. Prakash . (U.S. Geol. Sur., Denver, Colo.) Wood of Ginkgo in the Tertiary of western North America . Amer. Jour. Bot. 49(10): 1095–1101. Illus. 1962. —Woods of Ginkgo and extinct related genera are very rare in the fossil record in contrast to the numerous ginkgoalean leaves. Ginkgo wood may be distinguished from other gymnosperms by a combination of anatomical features herein described. Ginkgo wood from beds of Miocene age at Vantage, Washington, first identified by Beck, is assigned to a new species, G. beckii. Ginkgo wood from the upper Eocene Clarno Formation, John Day Basin, Oregon, is described as G. bonesii sp. nov. Scarcity of fossil ginkgoalean woods may reflect unusual susceptibility to degradation of their cell walls in contrast to the greater chemical resistance to degradation which features many coniferous woods.     

The urban woods of Rome (Italy)

ABSTRACT

A phytosociological study of the urban woods of Rome is described. Six syntaxonomical types were identified: 1) Orno-Quercetum ilicis, an edaphic-xerophytic variant of the natural potential vegetation of the city (deciduous Quercus sp. pl. woods); 2) Aquifolio-Fagetum carpinetosum betuli, in the northern slopes of alluvial valleys; 3) Q. suber and Q. pubescens wood, ranked in Quercetalia pubescenti-petraeae order 4) Quercetum ilicis galloprovinciale suberetosum; 5) coppice of Q. cerris and Q. frainetto, which belongs to the Teucrio siculi-Quercion cerridis alliance; 6) mixed mesophile wood of Q. cerris and Ostrya carpinifolia which can be considered a transition between the Doronico-Fagion and the Teucrio siculi-Quercion cerridis alliances. All of these six woody vegetation types are characterised by the large penetration of Mediterranean species belonging to the Quercetea and Quercetalia ilicis and the middle-European ones of the Querco-Fagetea. This mosaic of species exemplifies the bioclimatic characteristics of Rome, which is situated in a Mediterranean transitional region.     

The fairy shrimp genus Branchinella Sayce (Crustacea: Anostraca: Thamnocephalidae) in Western Australia,including a description of four new species

The genus Branchinella is diverse in Western Australia, with 18 species, including four new species described here. B. complexidigitata n. sp. is characterized by an intricate frontal appendage, unlike any within the genus. The other three are less remarkable; B. halsei n. sp. is like B. lyrifera, B. kadjikadji n. sp. belongs to the B. affinis group and B. nana n. sp. is similar to B. simplex; similarities are based on frontal appendage and to a lesser extent on the second antennae and penis. Only two species, B. affinis and B. longirostris, are common and widespread; many of the remainder being localized endemics. The high diversity in Western Australia is explained by broad latitudinal range, habitat diversity, and great age and isolation of the landscape.     

FLORA OF THE LOWER CRETACEOUS CEDAR MOUNTAIN FORMATION OF UTAH AND COLORADO. PART III: ICACINOXYLON PITTIENSE N. SP

Icacinoxylon pittiense, a new species of angiospermous wood from the Lower Cretaceous Cedar Mountain Formation of Utah is described and compared with similar fossil and modem woods. It is distinguished from other species of Icacinoxylon by its thick-walled fiber-tracheids with their walls making up at least 50% of the total diameter of the cells, conspicuous bordered pits with obliquely crossing extended apertures on both the tangential and radial walls of its fiber-tracheids, scalariform perforation plates with as few as four or greater than 30 bars, transitional opposite to scalariform pitting on its vessel walls, thick-walled ray cells, and distinct sheath or border cells in its rays. Icacinoxylon pittiense is the first species of this genus to be reported from Cretaceous sediments. This wood is of special interest because very few angiosperm woods have been reported from lower Cretaceous strata.     

Species area-relationships and frequency — Floristical data analysis of 44 isolated woods in northwestern Germany

44 isolated ancient woods (9,3–8579 ha) in southeastern Lower Saxony (northwestern Germany), where the Tertiary hilly country meets the Pleistocene plain, were investigated. Complete lists of 273 vascular plant species showing a more or less strong preference for woodlands were made for each wood including all the species of the groups Querco-Fagetea, Trifolio-Geranietea, Galio-Calystegietalia (selection of species) and Epilobietea. The majority of character species of woods show either a low or a high frequency, whereas fewer ones have medium frequencies. Most of the rarer species have their main occurrence in the larger woods. The number of species of all of the four groups increases with area of the wood and the correlation between the number of species and the log of area is related by a highly significant linear regression. Comparison of a single wood with two smaller woods of the same total area reveals that the two smaller woods on average have the greater number of wood species. Considering this and the fact that the rarer species have their main occurrence in the larger woods, when discussing questions of nature conservation a simple comparison of number of species and area is problematical.     

Characteristics of woods used recently and historically by Lesser Spotted Woodpeckers Dendrocopos minor in England

Lesser Spotted Woodpecker Dendrocopos minor numbers have declined greatly in England since the early 1980s for reasons that are not yet fully understood. It has been suggested that the species’ decline may be linked to the increase in Great Spotted Woodpeckers Dendrocopos major, changes in woodland habitat quality (such as deadwood abundance) and landscape‐scale changes in tree abundance. We tested some of these hypotheses by comparing the characteristics of woods in southern England where the species is still relatively numerous with those of woods used in the 1980s before the major decline. In each time period, habitat, predator and landscape information from woods known to be occupied by Lesser Spotted Woodpeckers was compared with those found to be unoccupied during surveys. Before the main period of decline, Lesser Spotted Woodpeckers used oak‐dominated, mature, open woods with a large amount of standing deadwood. Habitat use assessed from recent data was very similar, the species being present in mature, open, oak‐dominated woodlands. There was a strong relationship between wood use probability and the extent of woodland within a 3‐km radius, suggesting selection for more heavily wooded landscapes. In recent surveys, there was no difference in deadwood abundance or potential predator densities between occupied and unoccupied woods. Habitat management should focus on creating and maintaining networks of connected woodlands in areas of mature, open woods. Finer‐scale habitat selection by Lesser Spotted Woodpecker within woodlands should be assessed to aid development of beneficial management actions.     

Bird species turnover and stochastic extinction in woodland fragments

Year-to-year turnover in bird species composition was recorded across, the whole size range (0 02-30 ha) of 146 woods studied The mean number of resident breeding species both lost and gained per wood between consecutive breeding seasons was 2 (range 0-8) No relationship was found between this absolute turnover rate and woodland area, or any other of 24 predictor variables (describing woodland structure, isolation, connectedness and surrounding land use) Extriction and colonisation rates (in terms of numbers of species lost and gained) were also unrelated to woodland area In all sizes of woods, the species most likely to show local extinctions and colonisations were those with small populations within those woods, but the identity of the species concerned changed as woodland area increased In the smallest woods, the majority of turnover involved common species, such as wren and dunnock, which occurred in only small numbers in these small woods As woodland area increased, these species attained sufficient numbers to usually avoid stochastic extinction The majority of turnover was then due to more specialist (and less numerous) woodland species, such as great-spotted woodpecker and marsh tit, which were usually lacking in small woods In Britain, much existing broadleaved woodland falls within the size range studied Thus the numbers of many bird species are liable to be small enough for yearly turnover in woodland bird communities to be appreciable, and for the long-term persistence of individual species in particular woods to depend on dispersal     

Small Celastraceae and Polygonaceae twigs from the Upper Cenozoic (Ituzaingó Formation) of the La Plata Basin,Argentina

Two new wood types from the Late Cenozoic of the Ituzaingó Formation, La Plata Basin, Northeast Argentina add to our knowledge of South American Cenozoic plants. The materials were preserved by siliceous cellular permineralization, and they were prepared for microscopic examination by surface polishing and in thin sections. The anatomy of these new species was described. The relationship and comparison with the nearest living relatives (NLRs) are discussed. Maytenoxylon perforatum Franco gen. and sp. nov. is described as the first fossil wood referable to Celastraceae from South America. This new fossil species is related to extant Maytenus Molina. The other fossil twig, Ruprechtioxylon breae Franco sp. nov., has features of the Polygonaceae family and particularly resembles the extant specie Ruprechtia laxiflora Meisn. The occurrence of these fossil woods in south-eastern South America suggests that a relatively warm and dry to seasonally dry climate prevailed over this region of Gondwana during the Upper Cenozoic. It also provides new evidence for the hypothesis of the more wide distribution of Seasonally Dry Tropical Forest (SDTF) during the Upper Cenozoic.