The American scale insect genus Udinia De Lotto (Coccidae)

Udinia, an Ethiopian coccid genus, is redefined and a key is given to the thirteen species now recognised: bruncki sp. n. , catori (Green) comb. n. [= Lecanium subhirsutum Newstead syn. n. ], farquharsoni (Newstead) [= U.exoleta (De Lotto) syn. n. ], glabra De Lotto, ikoyensis sp. n. , lobayana (Balachowsky & Ferrero) comb. n. , newsteadi sp. n. , nigeriensis sp. n. , pattersoni sp. n. , paupercula De Lotto, pterolobina De Lotto, scitula De Lotto, setigera (Newstead). Five species are described as new and three species are redescribed and for these lectotypes are designated. The morphological characters used are discussed and listed and intraspecific variations shown by two of the species are tabulated. The distribution of all the species is recorded on two maps and the literature dealing with associations between ants and Udinia and related coccid genera is briefly reviewed.     

Pararalichus gen.n. (Acarina, Pterolichidae) from New World parrots (Aves, Psittacidae)

Pararalichus gen.n. (Pterolichidae, Pterolichinae) is described; two species groups, from species of Brotogeris Vigors and Touit Gray, respectively, include two named and ten new species. From Brotogeris : the type species Pararalichus dilatatus (Favette & Trouessart) stat.n., comb.n. [= Protolichus ( Mesolichus ) hemiphyllus dilatatus ] from Brotogeris jugularis (Müller), Brazil, Guatemala, Panama; P. chrysopteri sp.n. from B. chrysopterus (L.). Guyana, Surinam; P. cyanopterae sp.n. from B. cyanoptera (Pelzeln), Bolivia, Peru; P. pyrrhopteri sp.n. from B. Pyrrhopterus (Latham), Ecuador, Peru; P. sanctithomae sp.n. from B. sanctithomae (Müller), Bolivia, Peru; P. tiricae sp.n. from B. tirica (Gmelin), Brazil; P. versicolori sp.n. from B. versicolorus (Müller), Brazil, Peru. From Touit: P. hemiphyllus (Mégnin & Trouessart) comb.n. [ =Pterolichus (P.) hemiphyllus ] from Touit batavica (Boddaert), Surinam, Trinidad; P. dilectissimae sp.n. thrn T. dilectissima (Sclater & Salvin), Colmbia, Costa Rica; P. huetii sp.n. from T. huetii (Temminck), Brazil, Guyana, Peru; P. parpuratae sp.n. from T. purpurata (Gmelin), Brazil, Guyana, Surinam; P. surdae sp.n. from T. surda (Kuhl), Brazil. All hosts are Central and South American parrots. Mesolichus Trouessart, 1899 is recognized as a monotypical genus.     

Phylogenetic Systematics and Biogeography of the Neoephemeridae (Ephemeroptera: Pannota)

A revision of species of the pannote Holarctic and Oriental mayfly family Neoephemeridae is presented. Three genera are recognized in a strictly phylogenetic classification. Potamanthellus [=Neoephemeropsis Ulmer syn. n.] includes P. caenoides (Ulmer) comb. n., P. amabilis (Eaton) [=N. cuaraoensis Dang syn. n.], P. ganges sp. n., P. chinensis (Hsu) [=P. rarus (Tiunova and Levanidova) syn. n.], P. edmundsi sp. n., and the Oligocene fossil Potamanthellus rubiensis Lewis. Neoephemera [=Leucorhoenanthus Lestage syn. n.] includes N. maxima (Joly), N. purpurea (Traver), N. youngi Berner, N. bicolor McDunnough, and N. compressa Berner. Ochernova gen. n., includes O. tshernovae (Kazlauskas) comb. n. Taxa are described, illustrated and keyed. Species cladistics and biogeography are presented.     

Rusty coloured species of the lichen genus Miriquidica (Lecanoraceae)

Lecidea atrofulva [= Miriquidica atrofulva comb, nov.] has been recognized as an obligatorily rusty coloured representative of the genus Miriquidica. M. garovaglii and Lecidea plumbeoatra [= M. plumbeoatra comb, nov.] have been found to develop facultatively rusty coloured thalli. Miriquidica lapponica sp. nov. is described as new to science.     

Phylogenetic revision of the Taenogera Kröber genus-group (Diptera: Therevidae), with descriptions of two new genera

The phylogenetic relationships of the Taenogera genus-group, consisting of nine genera, are examined using cladistic methods. Twenty-one species representing nine genera and one outgroup were compared in a cladistic analysis across 99 states in 44 characters. The genus Taenogera Kröber is revised to contain only T. longa (Schiner), T. nitida (Macquart) and T. notatithorax Mann. Two new genera are described and figured: Actenomeros gen. n., with two species, A. corniculaticaudus sp. n. and A. onyx sp. n.; and Taenogerella gen. n., with four species, Ta. elizabethae sp. n., Ta. platina sp. n., Ta. schlingeri sp. n. and Ta . nigrapicalis (Mann) comb. n. Phylogenetic analysis also supports the removal of Nanexila gracilis (Mann) comb. n. from Taenogera .     

Capsalids (Platyhelminthes: Monogenea) from marine fishes off Okinawa in Japan with the proposal of two new genera

Molecular studies of the Capsalidae suggested that the genus Benedenia is polyphyletic, but a taxonomic organization of the genus that reflects molecular data has not yet been proposed. As a result of molecular analysis (28S rDNA, ITS1–5.8S-ITS2, and cox1 data) including specimens of Benedeniinae newly obtained from Okinawa-jima Island in Japan, two new genera and the revival of Tareenia independent to the genus Benedenia are proposed. Gracilobenedenia n. gen. is distinguished from the other genera of Benedeniinae based on morphological characteristics. This new genus comprises 6 species: G. lutjani n. comb. (type species), G. anticavaginata n. comb., G. rohdei n. comb., G. beverleyburtonae n. comb., G. kuremibai n. gen., n. sp., and G. hichi n. gen., n. sp. Armatobenedenia n. gen. is a monotypic genus for A. armata n. comb. The present molecular phylogenetic analysis showed the independence of Tareenia, and it can be morphologically separated from the other benedeniines. Four species including two new species obtained from Okinawa-jima Island are reported: G. kuremibai n. sp., G. hichi n. sp., G. lutjani n. comb., and Metabenedeniella parva. Furthermore, in the species identification and phylogenetic analysis of capsalids, the usefulness of not only the 28S rDNA but also ITS and the cox1 regions was suggested. These genes were evaluated the efficacy of those regions in DNA barcoding, and the ITS and cox1 regions shown to be useful for DNA barcoding in capsalids compared to the 28S rDNA sequence.     

Australian gall-inducing scale insects on Eucalyptus: revision of Opisthoscelis Schrader (Coccoidea, Eriococcidae) and descriptions of a new genus and nine new species

We revise the genus Opisthoscelis Schrader, and erect the genus Tanyscelisgen. n. with Opisthoscelis pisiformis Froggatt as its type species. Species of both genera induce sexually dimorphic galls on Eucalyptus (Myrtaceae) in Australia, with Opisthoscelis subrotunda Schrader also in Papua New Guinea. We synonymise the following taxa (junior synonym with senior synonym): Opisthoscelis fibularis Froggatt, syn. n. with Opisthoscelis spinosa Froggatt; Opisthoscelis recurva Froggatt, syn. n. with Opisthoscelis maculata Froggatt; Opisthoscelis globosa Froggatt, syn. n. (= Opisthoscelis ruebsaameni Lindinger) with Opisthoscelis convexa Froggatt; and Opisthoscelis mammularis Froggatt, syn. n. with Opisthoscelis verrucula Froggatt. We transfer seven Opisthoscelis species to Tanyscelis as Tanyscelis conica (Fuller), comb. n., Tanyscelis convexa (Froggatt), comb. n., Tanyscelis maculata (Froggatt), comb. n., Tanyscelis maskelli (Froggatt), comb. n., Tanyscelis pisiformis (Froggatt), comb. n., Tanyscelis spinosa (Froggatt), comb. n., and Tanyscelis verrucula (Froggatt), comb. n. We redescribe and illustrate the adult female of each named species of Opisthoscelis for which the type material is known, as well as the first-instar nymph of the type species of Opisthoscelis (Opisthoscelis subrotunda) and Tanyscelis (Opisthoscelis pisiformis). We describe four new species of Opisthoscelis: Opisthoscelis beardsleyi Hardy & Gullan, sp. n., Opisthoscelis thurgoona Hardy & Gullan, sp. n., Opisthoscelis tuberculataHardy & Gullan, sp. n., and Opisthoscelis ungulifinis Hardy & Gullan, sp. n., and five new species of Tanyscelis: Tanyscelis grallator Hardy & Gullan, sp. n., Tanuscelis megagibba Hardy & Gullan, sp. n., Tanyscelis mollicornuta Hardy & Gullan, sp. n., Tanyscelis tripocula Hardy & Gullan, sp. n., and Tanyscelis villosigibba Hardy & Gullan, sp. n. We designate lectotypes for Opisthoscelis convexa, Opisthoscelis fibularis, Opisthoscelis globosa Froggatt, Opisthoscelis maculata, Opisthoscelis mammularis, Opisthoscelis maskelli, Opisthoscelis pisiformis, Opisthoscelis recurva, Opisthoscelis serrata, Opisthoscelis spinosa, and Opisthoscelis verrucula. As a result of our taxonomic revision, Opisthoscelis has six species and Tanyscelis has 12 species. We describe the galls of females for all 18 species and galls of males for 10 species of Opisthoscelis and Tanyscelis, and provide photographs of the galls for most species. A key to the adult females of the species of both genera is included.     

Review of the Permian family Permulidae nomen novum pro Aliculidae Storozhenko, 1997 (Grylloblattida)

A new replacement name Permulidaenom. n. is proposed for the Permian family Aliculidae Storozhenko, 1997 (Insecta: Grylloblattida). A review of Permulidae is given. All genera and species are redescribed and illustrated. A key to genera of Permulidae is given. New taxa of Permulidae are described: Permula edemskiisp. n., Sojanopermula rasnitsynisp. n., Kazanalicula reductagen. et sp. n., Mezenalicula connatagen. et sp. n., all from the Soyana locality (Middle Permian, Kazanian Stage; Arkhangelsk Region, Russia), and Mezenalicula conjunctasp. n. from the Isady locality (Upper Permian, Severodvinian Stage; Vologda Region, Russia). New combinations are proposed: Permula minor (Aristov, 2004), comb. n. for Sojanopermula minor Aristov, 2004; Permula tshekardensis (Aristov, 2004), comb. n. for Sojanopermula tshekardensis Aristov, 2004. A new genus Acropermulagen. n. (type species: Permula acra Kukalová, 1964, from the Lower Permian of Czech Republic) is established in the family Ideliidae. Neraphidia Novokshonov & Novokshonova, 1997 is transferred from Aliculidae to the order Eoblattida as a genus of uncertain taxonomic position inside this order.     

Synonymy of Plotococcus Miller & Denno with Leptococcus Reyne, and description of a new species from Colombia (Hemiptera: Pseudococcidae)

Plotococcus Miller & Denno is synonymized with Leptococcus Reyne (Coccoidea: Pseudococcidae). The genus is redescribed and the adult female of the type species, L. metroxyli Reyne, is redescribed and illustrated. All species hitherto included in Plotococcus are transferred to Leptococcus as L. capixaba (Kondo) comb. nov., L. eugeniae (Miller & Denno) comb. nov., L. hambletoni (Kondo) comb. nov., L. minutus (Hempel) comb. nov., and L. neotropicus (Williams & Granara de Willink) comb. nov. A new species of Leptococcus, L. rodmani Kondo sp. n., from leaves of Guarea guidonia (Meliaceae) from Colombia, is described and illustrated based on the adult female. A revised key to adult females of all species in the genus is provided.     

A new genus of oak gallwasps, Zapatella Pujade-Villar & Melika, gen. n., with a description of two new species from the Neotropics (Hymenoptera, Cynipidae, Cynipini)

A new genus of cynipid oak gallwasp, Zapatella Pujade-Villar & Melika, gen. n. (Hymenoptera: Cynipidae: Cynipini), with two new species, Zapatella grahami Pujade-Villar & Melika, sp. n. and Zapatella nievesaldreyi Melika & Pujade-Villar, sp. n., is described from the Neotropics. Zapatella grahami,known only from the sexual generation,induces galls in acorns of Quercus costaricensis and is currently known only from Costa Rica. Zapatella nievesaldreyi, known only from the asexual generation, induces inconspicuous galls in twigs of Quercus humboldtii, and is known only from Colombia. Diagnostic characters for both new species are given in detail. Five Nearctic species are transferred from Callirhytis to Zapatella: Zapatella cryptica (Weld), comb. n., Zapatella herberti (Weld), comb. n., Zapatella oblata (Weld), comb. n., Zapatella quercusmedullae (Ashmead), comb. n.,Zapatella quercusphellos (Osten Sacken), comb. n. (= Zapatella quercussimilis (Bassett), syn. n.). A key based on adults for the species belonging to Zapatella is also given. Generic limits and morphological characteristics of Zapatella and closely related genera are discussed.     

Encarsia or Encarsiella? – redefining generic limits based on morphological and molecular evidence (Hymenoptera,Aphelinidae)

Abstract.  The genus Encarsiella ( syn.n. ) (Hymenoptera, Chalcidoidea, Aphelinidae) is synonymized with Encarsia on the basis of molecular and morphological characters. A phylogenetic analysis was conducted of the D2 expansion region of the 28S ribosomal DNA including thirty-eight Encarsia species. Our phylogenetic analysis revealed a polyphyletic origin of Encarsiella and confirmed the results of earlier analyses indicating the paraphyly of Encarsia . Four species are described as new ( Encarsia bifasciata sp.n. , E. caelata sp.n. , E. obliqua sp.n. and E. pilosa sp.n. ). The following species are transferred from Encarsiella to Encarsia : Encarsia aleurodici ( Girault, 1916 ) comb.n. , E. amabilis ( Huang & Polaszek, 1998 ) comb.n. , E. boswelli ( Girault, 1915 ) comb.n. , E. magniclava ( Girault, 1915 ) comb.n. , E. narroi Gómez & García comb.n. , E. nepalensis ( Polaszek & Hayat, 1992 ) comb.n. , E. noyesi ( Hayat, 1983 ) comb.n. , E. pithecura ( Polaszek, 1999 ) comb.n. , E. tachii ( Polaszek & Hayat, 1992 ) comb.n. , E. taiwanensis ( Chou & Chou, 1994 ) comb.n. and E. tamaulipeca ( Myartseva & Coronado-Blanco, 2002 ) comb.n. The generic limits of Encarsia are re-defined based on a re-assessment of morphological characters, and the results are discussed with respect to the current classification of the aphelinid subfamily Coccophaginae.     

First report of Litomosa spp. (Nematoda: Filarioidea) from Malagasy bats; review of the genus and relationships between species

The presence of the filarial genus Litomosa in Malagasy bats is demonstrated by the finding of L. goodmani n. sp. from Miniopterus gleni and Litomosa sp. (male unknown) from M. manavi, both in the Special Reserve of Ankarana. These materials are compared to the 22 Litomosa species, including two Indian species originally placed in the genus Litomosoides, L. fotedari (Gupta and Trivedi, 1989) n. comb. and L. tewarii (Gupta and Trivedi, 1989) n. comb., and the new taxon L. seurati n. sp. (= L. beaucournui Bain, 1966 pro parte), type-host Rhinolophus ferrum-equinum, Algeria, distinguished by the narrow area rugosa and the female caudal extremity with two conspicuous points, instead of several small ones. The Malagasy material belongs to a group of species close to the type, L. filaria, which have a male area rugosa composed of cuticular bosses and microfilariae folded within the sheath, and which are parasitic in Vespertilionidae, Hipposideridae and Rhinolophidae from Africa and Europe. The two Malagasy species resemble L. seurati n. sp., L. beshkovi Jancev, 1971, L. chiropterum Ortlepp, 1932, L. adami Petit, 1980 and L. ottavianii Lagrange et Bettini, 1948, with the enlarged third segment of the buccal capsule. L. goodmani n. sp. is distinct with its small size and female caudal extremity with a single point, which is suppressed in old mature worms; the females of Litomosa sp. have two conical points. Relationships among Litomosa species appear to be dependent upon both the chiropteran host groups and the geographical region.     

Revision of the Alamirinae (Diptera: Phoridae)

Abstract. The genus Couturiera Disney, 1979, only known in the female sex, is synonymized with Perissa Borgmeier, 1967, only known in the male sex, a syn.n., thus solving the problem of the 'missing' males of the Alamirinae. Perissa kensmithi (Disney) comb.n., P.lewisi (Disney) comb.n. and P.orientalis (Disney & Peterson) comb.n. are transferred from Couturiera. Perissa couturieri Disney is synonymized under P. lewisi (Disney) and C.palawanensis Disney under P.georgei Disney. Perissa latiptera sp.n., P.oligoseta sp.n., P.tinglei sp.n. and Perittophora couturieri gen.n., sp.n., are described from Zimbabwe. A key to the twelve species of Alamirinae is provided. It is hypothesized that the Alamirinae may be paraphyletic, by virtue of exclusion of the Termitoxeniinae. The latter is probably polyphyletic, in that each Alamirinae genus is probably the sister group of a different Termi-toxeniin clade.     

Ten new species of Haliotrema (Monogenoidea: Dactylogyridae) from Australian fish and a revision of the genus

Ten new species of Haliotrema from Australian fish are described and figured: H. cteno-chaeti sp. n. from Ctenochaetus strigosus; H. falcanalis sp. n. from Triacanthus falcanalis; H. lineate sp. n. from Acanthurus lineatus; H. chrysotaeniae sp. n. from Lutjanuschrysotaenia; H. cromileptis sp. n. from Cromileptis altivelis; H. epinepheli sp. n. from Epinephelus merra and E. fasciatus; H. holocentri sp. n. from Holocentrus ruber; Haliotrema chrysostomi sp. n. from Lethrinus chrysostomus and Plectorhinchus pictus; H. fleti sp. n. n. from L. fletus and L. chrysostomus; H. scari sp. N. from Scarus fasciatus.
H. dempsteri (Mizelle & Price, 1964) comb. n. from Acanthrus mata, A. dussumieri and A. xanthopterus: H. johnii (Tripathi, 1959) comb. n. from Lutjanus johni and L. fulviflamma: H. parahaliotrema (Mizelle & Price, 1964) comb. n. from Zebrasoma veliferum and A. grammoptilus: and H. obesa (Caballero, Bravo Hollis & Grocott, 1955) comb. n. from Tetraodon hispidus are redescribed and transferred from the genera Parahaliotrema Mizelle & Price, 1964, Ancyrocephalus Creplin, 1839, Parahaliotrema , and Tetrancistrum Goto & Kikuchi, 1917 respectively.
H. brevis (Mizelle & Price, 1964) comb, n. , H. canescens (Mizelle & Price, 1964) comb. n. and H. zanclus (Mizelle & Price, 1964) comb. n. are transferred from Pseudohaliotrema Yamaguti, 1953; H. eilatica (Paperna, 1965) comb. n. , H. teuthis (MacCallum, 1915) comb. n. , H. triacantha (Tripathi, 1959) comb. n. and H. lethrini (Yamaguti, 1937) comb. n. are transferred from Ancyrocephalus Creplin, 1839.
The generic diagnosis is emended to include the above-mentioned species and the taxonomy of the genus is discussed and the formation of six species groups is proposed.     

A new genus of Paramacronychiinae (Diptera: Sarcophagidae), argued from a genus-level cladistic analysis

Primorya gen.n. is described as a new genus of the flesh fly subfamily Paramacronychiinae, including the type species and sole member Primorya ussuriensis sp.n. from the Russian Far East. Arguments for erecting a new genus are obtained from a phylogenetic analysis of all paramacronychiine genera. Goniophyto suenagai (Kurahashi), comb.n. is transferred from Wohlfahrtiodes Villeneuve and included in a broadened concept of Goniophyto Townsend, and Agria cicadina (Kato) , comb.n. is transferred from Angiometopa Brauer & Bergenstamm. A key to all paramacronychiine genera is provided.     

Revision of the Nearctic species of the genus Stiropius Cameron (=Bucculatriplex Auct.) with the description of a new related genus (Hymenoptera: Braconidae)

Abstract. The New World rogadine genus Stiropius Cameron is revised for the Nearctic region and a new related genus from the Americas, Viridipyge , is decribed. Bucculatriplex Viereck is found to be a junior subjective synonym of Stiropius Cameron. Three Nearctic species are recognized: S.bucculatricis (Ashmead) comb.n., S.californicus sp.n. and S.wagneri sp.n. A single Nearctic specis of Viridipyge is described as new: V.prunicola , and a Neotropical species, V.letifer (Mann) comb.n., is found to be congeneric. The biology and systematic relationships of both genera are discussed; the two genera appear to represent a relatively basal branch of the Rogadini lineage.