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1.
Grazing and browsing by sheep and goats has been an important anthropogenic influence on ecosystems in the Mediterranean region for centuries. This influence has changed significantly in recent decades, with a general shift from range grazing to the penning of animals. The intermediate disturbance hypothesis (IDH) proposes that perturbation - including anthropogenic disturbance - is the norm for Mediterranean ecosystems, and that higher species diversity is found under conditions of continuous, moderate disturbance. Here we test the IDH as it relates to grazing of scrub and open forest habitats in Cyprus, while also testing for the effects of fire. We carried out surveys of breeding birds and vegetation at 48 study sites in scrub and open woodland across Cyprus. We estimated relative grazing pressure (past and present) and fire history at these sites, and tested for associations between these factors, breeding bird species richness and perennial vegetation diversity. We found moderate current grazing reduces density of lower and middle level scrub, and a higher level of diversity of perennial vegetation at moderately over intensively grazed sites. We found that moderate historical grazing pressure had a positive influence on richness of breeding bird species in lower scrub habitat such as phrygana, but that this effect was weaker the taller and denser the habitat. By contrast, current grazing pressure had a negative effect on species richness in lower scrub, but species richness in grazed habitats was higher in denser, taller scrub habitat such as garrigue and maquis. Our study suggests grazing plays an important role in maintaining habitat heterogeneity but the impact on avian species richness depends on the density and height of the vegetation.  相似文献   

2.
The effects of two grazer species (cattle or sheep) and two flock types (single or mixed with goats) on vegetation and arthropod fauna were studied in a factorial design on eight plots which comprised two thirds of mechanically cleared heathland and one third of improved ryegrass-clover grassland. After six grazing seasons, the shrubland areas were dominated by gorse (Ulex gallii) in all treatments. Herbaceous cover was higher under mixed than under single grazing, and under sheep than under cattle grazing. Higher captures of Opiliones, Julida, Lithobiomorpha, Microcoryphia and Carabidae were recorded in shrublands than in grasslands, while the reverse was observed for Linyphiidae, Lycosidae and Hemiptera. Within shrublands, fauna responded to the flock type but not to the grazer species. More arthropod groups favoured the patchier areas with higher herbaceous biomass generated by mixed herds with goats. Within grasslands, species-specific responses to the grazer species were observed. Mixed grazing schemes which include goats within partially improved heathlands could contribute to maintain higher biodiversity levels in these marginal areas.  相似文献   

3.
Restored grasslands and shrublands are integral parts of the semi-natural landscape and are of major importance for biodiversity in the northern Loess Plateau. Determining the underlying factors that control the richness and composition of herbaceous species in restored grasslands and shrublands is urgently needed. Thus, the specific objective of this study was to evaluate the relative importance of soil, plant, and topographic explanatory variables affecting the richness and composition of herbaceous species in restored shrubland and grassland ecosystems in a typical watershed within the northern Loess Plateau. In this study, 27 restored grassland sites and 16 restored shrubland sites were sampled during September 2009. Using variation partitioning (partial canonical correspondence analysis), we determined the individual and shared effects of these three sets of explanatory variables on herbaceous biodiversity in the two restored habitats. Most of the explained variation in plant diversity was related to the pure effect of soil, plant, and topographic variables. Restored shrublands had significantly more species than grasslands, and abandoned dam farmlands had significantly more species than other grassland sites. Moreover, botanical diversity responded differently to the explanatory variables in different plant communities. The pure effects of soil properties, soil moisture in particular, accounted for the largest fractions of explained variation in species diversity in restored grasslands. Both plant and topographic variables had balancing pure effects on species diversity in restored shrublands, in particular the shrub density and slope angle. We conclude that the maintenance of a moderate density of shrubs (less than 3600 shrubs per ha), construction of check-dams, and grazing at a low stocking rate, taking conditions of soil and topographic site into account, may help to conserve biodiversity in the northern Loess Plateau.  相似文献   

4.
Question: What is the impact of grazing and/or afforestation on grassland diversity, species composition and cover parameters? Location: Semi‐arid Mediterranean grasslands of Jordan. Methods: Vegetation, litter, bare soil and rock cover were compared among four management types – free grazing and protected from grazing with three levels of tree cover. Species composition, plant cover, species richness and evenness were used to evaluate differences in vegetation among management types. Species composition differences among management types were also investigated. Results: Semi‐arid Mediterranean grasslands harbour appreciable levels of plant biodiversity. Grazing did not affect plant diversity, indicating the high resilience against and adaptation to grazing; however,grazing affected species composition and cover parameters. Afforestation seems to protect soil through higher litter cover but its impact on plant biodiversity was negative and markedly affected species composition. Conclusions: Neither protection from grazing or massive afforestation alone are sufficient for conserving biodiversity in this system. A management model is suggested where the landscape should be maintained as a mosaic of four management types: complete protection from grazing, grazing rotation, planting sparse trees in eroded areas and revegetating degraded areas using native, herbaceous and grazing tolerant species.  相似文献   

5.
Extensively managed pastures are of crucial importance in sustaining biodiversity both in local- and landscape-level. Thus, re-introduction of traditional grazing management is a crucial issue in grassland conservation actions worldwide. Traditional grazing with robust cattle breeds in low stocking rates is considered to be especially useful to mimic natural grazing regimes, but well documented case-studies are surprisingly rare on this topic. Our goal was to evaluate the effectiveness of traditional Hungarian Grey cattle grazing as a conservation action in a mosaic alkali landscape. We asked the following questions: (i) How does cattle grazing affect species composition and diversity of the grasslands? (ii) What are the effects of grazing on short-lived and perennial noxious species? (iii) Are there distinct effects of grazing in dry-, mesophilous- and wet grassland types? Vegetation of fenced and grazed plots in a 200-ha sized habitat complex (secondary dry grasslands and pristine mesophilous- and wet alkali grasslands) was sampled from 2006–2009 in East-Hungary. We found higher diversity scores in grazed plots compared to fenced ones in mesophilous- and wet grasslands. Higher cover of noxious species was typical in fenced plots compared to their grazed counterparts in the last year in every studied grassland type. We found an increasing effect of grazing from the dry- towards the wet grassland types. The year-to-year differences also followed similar pattern: the site-dependent effects were the lowest in the dry grassland and an increasing effect was detected along the moisture gradient. We found that extensive Hungarian Grey cattle grazing is an effective tool to suppress noxious species and to create a mosaic vegetation structure, which enables to maintain high species richness in the landscape. Hungarian Grey cattle can feed in open habitats along long moisture gradient, thus in highly mosaic landscapes this breed can be the most suitable livestock type.  相似文献   

6.
One of the most important conservation issues in ecology is the imperiled state of grassland ecosystems worldwide due to land conversion, desertification, and the loss of native populations and species. The Janos region of northwestern Mexico maintains one of the largest remaining black-tailed prairie dog (Cynomys ludovicianus) colony complexes in North America and supports a high diversity of threatened and endangered species. Yet, cattle grazing, agriculture, and drought have greatly impacted the region. We evaluated the impact of human activities on the Janos grasslands, comparing changes in the vertebrate community over the last two decades. Our results reveal profound, rapid changes in the Janos grassland community, demonstrating large declines in vertebrate abundance across all taxonomic groups. We also found that the 55,000 ha prairie dog colony complex has declined by 73% since 1988. The prairie dog complex has become increasingly fragmented, and their densities have shown a precipitous decline over the years, from an average density of 25 per ha in 1988 to 2 per ha in 2004. We demonstrated that prairie dogs strongly suppressed woody plant encroachment as well as created open grassland habitat by clearing woody vegetation, and found rapid invasion of shrubland once the prairie dogs disappeared from the grasslands. Comparison of grasslands and shrublands showed markedly different species compositions, with species richness being greatest when both habitats were considered together. Our data demonstrate the rapid decline of a grassland ecosystem, and documents the dramatic loss in biodiversity over a very short time period concomitant with anthropogenic grassland degradation and the decline of a keystone species.  相似文献   

7.
Abstract. Species richness in calcareous grassland is discussed against the background of historical dispersal processes associated with traditional land-use management such as grazing, but also the artificial establishment by hayseed. Important vectors in the traditionally man-made landscape were sheep and cattle or other livestock such as goats. Calcareous grasslands were not only connected to each other but also to other habitats such as villages, forests, arable fields and heathlands by these vectors which could cover large distances (e.g. transhumance shepherding), which is not the case in the current man-made landscape. Species richness after restoration management of abandoned and afforested calcareous grasslands was predicted by using characters of dispersability in space and time. These were the persistence of the species in the vegetation and the diaspore bank after abandonment or afforestation and the dispersal capacity through wind and sheep. The results reveal that reintroduction of sheep grazing is necessary to reestablish the original species richness. The first validation of the prediction of the succession on clear-cut sites and a comparison with data of species composition in abandoned quarries and the surroundings made it obvious that a species' own dispersal capacity in space is very low except for some well wind-dispersed species. Therefore, it is recommended to include and to simulate dispersal processes in future management to be able to restore the original species richness.  相似文献   

8.
Montane grassy habitats in the Western Carpathians are relatively well preserved, maintain high species richness and are often important in accordance to the nature conservation policy in Europe. However, knowledge about the impact of farming on the habitat quality there is rather poor. The influence of various management types (permanent sheep pen, irregular grazing, mowing) on Orthoptera diversity and species determining assemblages of these habitats were analysed on 72 plots in Poľana Mts Biosphere Reserve. Altogether, 36 Orthoptera species (15 Ensifera, 21 Caelifera) were found, whereas the highest number of species was found on plots with irregular grazing (28 species), followed by plots with mown grass (17) and permanent sheep pens (14). All four measures of the assemblages’ diversity confirmed significant differences. Using Discriminant Function Analysis, correct classification rate of Orthoptera assemblages was unambiguous according to the type of management. Each form of the management harboured several characteristic species. Thus implications regarding the biodiversity conservation and grassland management were given.  相似文献   

9.
Managed grasslands are normally ecosystems with high species diversity. They are associated with traditional extensive livestock grazing. Land-use changes, in particular gradual abandonment or grazing intensification, lead to major changes in floristic patterns of these grasslands, some included in priority habitats of the EU Habitats Directive. In order to analyse these patterns of change, Mediterranean grasslands of eastern areas of mainland Portugal were studied aiming to: (1) establish ecological gradients underlying their floristic patterns; (2) examine the relative importance of both environmental and land-use factors on their floristic composition; (3) assess how floristic composition and species richness are affected by land-use factors. Vegetation sampling was carried out from 2008 to 2009 following phytosociological procedures. Canonical Correspondence Analysis was applied. Variation partitioning was used to assess the relative influence of land-use variables. Richness, legumes cover, endemic species and bryophytes cover were compared in four land-management regimes: unmanaged; extensive grazing by sheep; extensive grazing by cattle and sheep; frequent soil tillage. Significant differences were found in floristic and diversity patterns, revealing the importance of grazing as a management tool for maintaining or improving floristic diversity. From our findings slight extensive grazing can contribute to both maintenance of species diversity and to the increase in legumes cover and thus to palatability and economic value of studied grasslands. Interactions between herbivores and grasslands, considering also their influences on soil properties, have an important role in the equilibrium states which promote biodiversity and conservation of the services of these ecosystems.  相似文献   

10.
The target rate of afforestation in Ireland over the next 30 years is 20,000 ha per year, which would result in an increase of the forest cover from the current 10% to 17%. In order to promote sustainable forest management practices, it is essential to know the composition and conservation value of habitats where afforestation is planned and the effects of subsequent planting upon biodiversity. The objectives of this study were to investigate changes in vegetation composition and diversity of grasslands 5 years after afforestation with Sitka spruce (Picea sitchensis) and determine the primary ecological and management factors responsible for these changes. Species cover, environmental and management data were collected from 16 afforested and unplanted improved and wet grassland site pairs in Ireland. Our results indicate that 5 years after tree planting, there were significant changes in richness, composition, and abundance of species. Competitive and vigorous grasses were more abundant in planted than in unplanted sites, as were generalist species found in both open and wooded habitats, while small-stature shade-sensitive species were less abundant. Vascular plant species richness and Shannon’s diversity index were higher in unplanted wet grassland, than in the planted sites. Bryophyte species richness was higher in planted improved grassland than in unplanted sites. The differences were primarily the result of the exclusion of grazing, ground preparation, changes in nutrient management and drainage for afforestation. Drainage ditches provided a temporary habitat for less competitive species, but the overall effect of drainage was to reduce the diversity of species dependent on wet conditions. Variance partitioning showed differences in the relative influences of environmental and management variables on biodiversity in the two habitats, probably due to the greater pre-afforestation grazing pressure and fertilisation levels in improved grasslands. The differences in biodiversity between planted and unplanted grasslands indicate that afforestation represents a threat to semi-natural habitats where distinctive and highly localised plant communities could potentially occur.  相似文献   

11.
Scattered trees are considered keystone structures and play an important role in Mediterranean sylvopastoral systems. Such systems are associated with high biodiversity and provide important natural resources and ecosystem services. In this study, we measured the contribution of scattered trees and different grazing management (cattle, sheep and wildlife only) to the diversity of the grassland sward in a dehesa (open holm oak woodland) located in Central Spain. We analyzed alpha and beta diversity through measurement of species richness, Shannon-Wiener, and Whittaker indices, respectively; and the floristic composition of the herb layer using subplots within two adjacent plots (trees present vs. trees absent) under three different grazing management regimes, including wildlife only, during a year. We found a 20–30% increment in the alpha diversity of wooded plots, compared to those without trees, regardless of grazing management. All beta indices calculated showed more than 60% species turnover. Wooded plots were occupied by different herbaceous species in different heterogeneous microsites (under the canopy, in the ecotone or on open land) created by the trees. Livestock grazing modified species composition (e.g. more nitrophilous species) compared to wildlife only plots. In addition to all their other benefits, trees are important to maintaining grassland diversity in Mediterranean dehesas.  相似文献   

12.
In grazing ecosystems, mature seeds fall directly to the soil to form the soil seed bank (SSB), or are ingested by grazing livestock to become part of the dung seed bank (DSB; i.e., seed circulation). Both the SSB and DSB form the basis for the natural regeneration of vegetation. However, little is known about the relationships between the SSB, DSB, and aboveground vegetation (AGV) community under different stocking rates (SRs). This study investigated the relationships between the SSB, seeds in Tan sheep (Ovis aries) dung, and AGV at different SRs (0, 2.7, 5.3, and 8.7 sheep ha–1) in a semiarid region of the Loess Plateau in China. We found that Tan sheep grazing increased the species richness heterogeneity of grassland vegetation, and negatively influenced the density of AGV. Under natural conditions, 17 species from soil‐borne seeds and 10 species from Tan sheep dung germinated. There was low species similarity between the soil and DSBs and AGV. Sheep SR and the seed banks (soil and dung) were negatively correlated with AGV. Seeds are cycled from herbage to livestock to soil during cold season grazing; the seasonal nature of this seed dispersal is an adaptation to harsh, semiarid environments. Increased seed bank diversity under sheep grazing facilitates grassland regeneration on the Loess Plateau, similarly to other semiarid regions globally.  相似文献   

13.
The study described changes in floristic and vegetation structure in relation to livestock grazing intensity in a conservation area in the Succulent Karoo, South Africa. Grazing by goats and sheep is allowed in the Richtersveld National Park (a contractual National Park. which is also an area of high floristic richness and endemism. We used goat faecal pellet density, degree of trampling and percentage bare-ground at distances from the stock posts as surrogates for a gradient in grazing pressure. A stock post is the place where farmers keep, in most cases in an enclosure called a ‘kraal’, their animals at night and to which they return every evening after the day’s herding. Twenty-seven stock posts were located in the Richtersveld National Park; nine stock posts on flats, foot– slopes and mountain each. We measured plant species richness and diversity, and mean percentage cover of the various plant growth forms (including the number of species falling into each growth form category. in each of the five 10 m . 10 m plots (each 200 m apart. demarcated along a transect of one kilometre length from the centre of each stock post. The results showed that distance from the stock post does reflect grazing intensity use because densities in faecal pellets rapidly declined with increasing distances away from the stock post for all habitats studied. Faecal density was positively correlated with stocking density. Plant species richness and diversity was at a minimum near stock posts. Plants able to endure the effects of heavy grazing occurred near stock posts where declines in palatable plant species, assumingly sensitive to heavy grazing and trampling, were recorded. Grazing increased vegetation patchiness up to 800 m from the stock post for all the habitats. The degree to which this change in species composition occurred did not depend on stocking densities, suggesting that both grazing and landscape variability were responsible for vegetation changes in rangelands of that area of the Succulent Karoo biome.  相似文献   

14.
The assessment of the effects of long-term management practices is relevant in understanding the current patterns of plant assemblages in semi-natural ecosystems. We hypothesized that the variety of management practices across different farming systems under the same ecological conditions directly and indirectly shapes these patterns via the long-term changes induced in soil features. The aims of this paper were to evaluate the influence of two sets of variables describing long-term management practices and soil features on plant assemblages and their importance in the context of Mediterranean silvo-pastoral systems. The analysis of variance revealed that richness and grazing value were not affected at all by grazing livestock species and soil tillage frequency and that they both showed relatively high absolute values for the specific context under study. Trifolium subterraneum was a key species in contributing to grassland grazing value and habitat biodiversity. The Canonical Correspondence Analysis highlighted the influence of management practices and soil features on plant assemblage composition, which was significantly affected by grazing livestock species and stocking rate and by soil pH and K content. The Redundancy Analysis showed that soil pH and related features were in turn affected by stocking rate, supporting our hypothesis that management practices influenced plant assemblage composition directly and indirectly via their long-term effects on soil features. The results also highlighted that a systemic analytical perspective applied at a grazing system scale can be effective in addressing sustainable grassland management issues in Mediterranean silvo-pastoral systems.  相似文献   

15.
Grasslands being used in sheep farming systems are managed under a variety of agricultural production, recreational and conservational objectives. Although sheep grazing is rarely considered the best method for delivering conservation objectives in seminatural temperate grasslands, the literature does not provide unequivocal evidence on the impact of sheep grazing on pasture biodiversity. Our aim was therefore to review evidence of the impacts of stocking rate, grazing period and soil fertility on plant communities and arthropod populations in both mesotrophic grasslands typical of agriculturally improved areas and in native plant communities. We therefore conducted a literature search of articles published up to the end of the year 2010 using ‘sheep’ and ‘grazing’ as keywords, together with variables describing grassland management, plant community structure or arthropod taxa. The filtering process led to the selection of 48 articles, with 42 included in the stocking rate dataset, 9 in the grazing period dataset and 10 in the soil fertility dataset. The meta-analysis did not reveal any significant trends for plant species richness or plant community evenness along a wide stocking rate gradient. However, we found frequent shifts in functional groups or plant species abundance that could be explained by the functional properties of the plants in the community. The meta-analysis confirmed that increasing soil fertility decreased plant species richness. Despite the very limited dataset, plant species richness was significantly greater in autumn-grazed pastures than in ungrazed areas, which suggests that choosing an appropriate grazing period would be a promising option for preserving biodiversity in sheep farming systems. Qualitative review indicated that low grazing intensity had positive effects on Orthoptera, Hemiptera (especially phytophagous Auchenorrhyncha) and, despite a diverse range of feeding strategies, for the species richness of Coleoptera. Lepidoptera, which were favoured by more abundant flowering plants, also benefited from low grazing intensities. Spider abundance and species richness were higher in ungrazed than in grazed pastures. In contrast, there are insufficient published studies to draw any firm conclusions on the benefits of late grazing or stopping fertilization on insect diversity, and no grounds for including any of this information in decision support tools at this stage.  相似文献   

16.
青藏高原高寒灌丛植被对长期放牧强度试验的响应特征   总被引:1,自引:1,他引:0  
在青藏高原中国科学院海北高寒草甸生态系统定位研究站对金露梅高寒灌丛草场植被开展了长期不同放牧强度试验,分别在短期(4年)、中期(11年)和长期(18年)放牧阶段研究不同放牧干扰强度对草地植物物种多样性、群落结构、地上生物量和草场质量的影响.研究表明,在不同放牧阶段,随着放牧强度增加植物群落的高度和盖度都降低.在中期放牧干扰阶段,物种多样性数和均匀度指数随着放牧强度增加呈现典型的单峰曲线模式;在长期放牧干扰阶段,随着放牧强度增加,占优势地位的灌木和禾草被典型杂类草替代,其中的重度放牧干扰简化了高寒灌丛植被群落结构,减少了地上现存生物量,特别是可食优良牧草生物量.植被对放牧的响应除了与放牧强度和放牧时间阶段密切相关外,还与该地区水热条件的变化有一定的相关性.针对长期放牧干扰的反应特性可将金露梅灌丛草场中植物划分为增加型、敏感型、忍耐型和无反应型4种类型.除了丰富度指数、多样性指数和均匀度指数外,其它一些特征参数并不支持著名的中度干扰假说.本研究发现,长期重度放牧促进了青藏高原高寒草地退化,适度放牧有利于高寒灌丛草场的生物多样性保护和牧草利用;"取半留半"的放牧原则在青藏高原草场放牧管理实践中值得推荐,它将有利于防止草场退化,提高牧草利用率和维持较高的生物多样性.  相似文献   

17.
Livestock grazing is a widespread grassland disturbance and can negatively impact biodiversity. Pollinators constitute a vital component of grassland ecosystems, but the impact of grazing on pollinator diversity has seldom been evaluated in North America. We assessed vegetation structure, and pollinator and flowering plant abundance, richness, diversity, and community composition in four pairs of spring-grazed/ungrazed sites in south-central British Columbia, Canada. We also investigated whether pollinator or floral communities differed between the two threatened shrubsteppe habitat types we sampled—antelope-brush and big sagebrush shrubsteppe. Pan-trapping surveys captured 5907 bees, flies, beetles, wasps and butterflies constituting 253 species. We found that the percent cover of shrubs and bare soil increased with grazing, while the height of grasses and forbs decreased. In contrast, pollinator and flowering plant abundance, richness, diversity, and community composition were not significantly affected by grazing. Flowering plant and pollinator community composition did differ significantly between shrubsteppe habitats. Our results indicate that grasslands in North America, when managed responsibly, can maintain pollinator and flowering plant diversity under grazing pressure. The continued effort of land managers to balance ecological integrity and economic viability will be important for the conservation of grassland pollination systems.  相似文献   

18.
Balancing food production and biodiversity conservation is a global challenge today. Livestock grazing is one of the main activities triggering habitat degradation and land-use change around the world. Its effects on biodiversity have been widely explored, with birds being the most studied vertebrates. However, its impact seems to be contradictory given the disparity of the results. To understand the influence of livestock grazing on birds, we conducted a meta-analysis exploring the effects of several grazing characteristics on bird abundance and species richness. Our results showed that livestock grazing has a significant negative effect on bird abundance (mean effect size -0.422 ± 0.140), and species richness (mean effect size -0.391 ± 0.141). Livestock grazing affected negatively the bird abundance in riparian habitats in contrast to the other habitat types. Species richness was negatively affected by grazing in woody habitats and Afrotropical and Neotropical regions. Grazing by cattle was more detrimental for both bird richness and abundance than sheep grazing or a mixture of domestic livestock. Moreover, intermediate grazing intensity seems appropriate to maintain bird abundance and richness, as high grazing intensity dropped both bird abundance and species richness substantially, and low grazing intensity reduced bird species richness. This pattern supposes a non-linear effect of grazing intensity on birds. Therefore, the management of grazing intensity and type of livestock could help to reduce the negative effect on bird abundance and richness, as moderate grazing intensities and mix of livestock types appear to have a minor or null impact on bird abundance and richness. Future studies should explore in-depth the effect of moderate grazing intensities on bird diversity and composition to provide better management recommendations to enhance avian conservation in rangelands.  相似文献   

19.
The Loess Plateau is a special natural–cultural unit in northern China. Intensive land use in the past has had, and forestation and grass planting at present will have inevitable impacts on plant biodiversity in the Loess Plateau. Based on the analysis of floristic features within three sampling sites with different land use practices and analysis of species richness among different land use types, we discuss impacts of land use on species richness and floristic features in the Northern Loess Plateau. The results drawn from this case study are as follows: (1) It appears that forestation and grass planting have had a positive influence on the local species diversity, but they have contributed little to the native vegetation in terms of conserving its floristic features. (2) Caragana intermedia shrubland, Pinus tabulaeformis forestland, and natural grassland have made important contributions to supporting indigenous species and maintaining local plant biodiversity. (3) There is a significant positive correlation between land use diversity and species richness. These results imply that practicing biodiversity conservation in situ is feasible and the suitable choice for the Loess Plateau. Concrete measures for biodiversity conservation in the area can include setting up small nature reserves and diversifying land use patterns to maintain as much habitat as possible for native vegetation. The artificial Hippophae rhamnoides shrubland should not be further promoted, considering its negative influence on biodiversity conservation.  相似文献   

20.
“Landscape modulators” are ecosystem engineers that have an impact on community structure by creating patches in the landscape mosaic. Our aim was to study the effect of evergreen-trees, as landscape modulators, on herbaceous plants in a Mediterranean maquis system in northern Israel. We examined the effects of canopy removal and cattle grazing on species richness, plant functional types, and rare plant species in two patch-types: (1) woody—under tree canopy (or the location of a removed canopy); (2) herbaceous—in open areas with no tree canopy. Patch-type and tree removal affected species richness and plant functional types. The extreme negative effect of the woody patch-type on species richness disappeared soon after the removal of the landscape modulator canopy. We conclude that the dominant effect of the evergreen woody landscape modulators can be regulated by canopy removal and grazing for maintaining patch-type and landscape diversities, and consequent high species richness in Mediterranean ecosystems, which is a main goal of global nature conservation policy.  相似文献   

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