Phylogeny and classification of Corylophidae (Coleoptera: Cucujoidea) with descriptions of new genera and larvae

Abstract Phylogenetic relationships within the family Corylophidae were investigated. Twenty ingroup taxa and six outgroups were included in a cladistic analysis, based on 48 characters derived from adult and larval morphology. Phylogenetic analysis confirms that Corylophidae are monophyletic within the superfamily Cucujoidea and may be subdivided into two subfamilies: the Australian Periptycinae and the cosmopolitan Corylophinae containing 10 tribes: Foadiini trib.n. , Cleidostethini, Aenigmaticini, Parmulini, Sericoderini, Peltinodini, Orthoperini, Corylophini, Teplinini and Rypobiini. All currently recognized family‐group taxa are thoroughly diagnosed, and keys to their identification based on adults and larvae are provided. Two new genera and three species are described: Weirus gen.n ., containing only W. tozer sp . n . (Australia: Queensland), and Stanus gen.n. , with the two species S. bowesteadi sp.n . (New Zealand) and S. tasmanicus sp.n. (Tasmania). The larvae of Pakalukodes bimaculatus?lipiński et al. from Queensland and of Stanus bowesteadi sp.n. from New Zealand are described and illustrated for the first time.     

Rotoitidae, a curious new family of Chalcidoidea (Hymenoptera) from New Zealand

Abstract. A new chalcidoid family, Rotoitidae fam.n., is described from New Zealand. The probable systematic affinities of Rotoitidae are discussed and the type genus and a single included species, Rotoita basalis gen. et sp.n., are described and illustrated.     

Phylogenetic analysis of a group of palaeonemerteans (Nemertea) including two new species from Queensland and the Great Barrier Reef, Australia

Sundberg, P., Gibson, R. & Olsson, U. (2003). Phylogenetic analysis of a group of palaeonemerteans (Nemertea) including two new species from Queensland and the Great Barrier Reef, Australia. — Zoologica Scripta, 32, 279–296.
Based on 18S rDNA nucleotide sequences and morphological characters, we reconstruct the phylogeny for a group of palaeonemerteans estimated to be monophyletic. Two new palaeonemertean species from Queensland and the Great Barrier Reef, Australia are included in the phylogenetic analysis. The results confirm that one of the species, Cephalothrix queenslandica sp. n., is part of the Cephalothrix–Cephalotrichella–Procephalothrix group. These genera are redefined phylogenetically under the name Cephalothrix based on the cladistic analysis. The other species, Balionemertes australiensis gen. et sp. n., is placed in a new genus which forms a sister taxon to Cephalothrix . The morphology of both new species is described in detail.     

Systematics and biology of Penthaleus tectus sp. n. (Acari: Penthaleidae), a recently discovered pest of grain crops in eastern Australia

Abstract  A new species of pest mite, Penthaleus tectus sp. n. (Acari: Penthaleidae) is described from south-eastern Australia, where it attacks several grain crops including wheat, in which it can cause significant damage. It has previously been confused with the blue oat mite Penthaleus major (Dugès). Penthaleus tectus differs from P. major morphologically and genetically, and in geographical distribution, response to pesticides, and the timing of diapause. Previous records of ' P. major ' from Australia and other countries probably include a mixture of several species. Penthaleus minor (Canestrini) is recorded from South Africa for the first time.     

New fan worm genera and species (Polychaeta, Sabellidae, Fabriciinae) from the western Pacific, and cladistic relationships among genera

Five new species are described from among the genera Fabriciola Friedrich, 1939 (one species from Okinawa), Novafabricia Fitzhugh, 1990 (two species from Australia and Papua New Guinea, respectively), and two new monotypic genera, Brifacia gen. n. (Australia), and Pseudoaugeneriella gen. n. (Okinawa). All species are known only from the intertidal. Fabriciola rubra sp. n. is a member of the species complex with red peristomial and pygidial eyes, inferior thoracic flagellate setae, and abdominal pin-head setae. Novafabricia labrus sp. n., and N. exiguus sp. n., are distinguished from other species based on the distribution of inferior thoracic pseudospatulate setae. Brifacia metastellaris sp. n., resembles Fabricia stellaris (Müller, 1774) with regard to anterior peristomial ring collar construction and distribution of pseudospatulate setae, but differs in having abdominal uncini with relatively short manubria. Pseudoaugeneriella unirama sp. n., most closely resembles species of Augeneriella Banse, 1956 in general form, but has unbranched vascularized ventral filamentous appendages. Revised keys to fabriciin genera and selected species are provided. Cladistic relationships of these new genera and species to other Fabriciinae are examined and the status of some characters are reviewed. Whereas previous analyses have supported a monophyletic Augeneriella on the basis of branched, vascularized ventral filamentous appendages, present results show Augeneriella paraphyletic relative to Fabricia Blainville, 1828, Novafabricia, Fabricinuda Fitzhugh, 1990 Parafabricia Fitzhugh, 1992 and Brifacia. Similarly, some topologies show Novafabricia, which has only been defined by the presence of poorly developed dorsal lips, to be paraphyletic relative to Fabricia, Fabricinuda, Parafabricia, and Brifacia. The cladistic analyses do, however, support recognition of Brifacia and Pseudoaugeneriella. With the addition of Brifacia, the presence of unbranched, vascularized appendages is plesiomorphic for the Fabriciinae, which has not been previously hypothesized.     

Review of Australian Asperoseius Chant, Euseius Wainstein, Okiseius Ehara and Phytoscutus Muma (Acari: Mesostigmata: Phytoseiidae) with a key to the genera of Australian Amblyseiinae and descriptions of two new species

Previous workers have used a conservative generic classification of the subfamily Amblyseiinae that treated most Australian Phytoseiidae as species in the genus Amblyseius Berlese. However, this lumping masks the occurrence of a number of genera recognised overseas, including some that contain important biological control agents. In this review, a key to the genera of Australian Amblyseiinae is provided and the Australian species in four amblyseiine genera are reviewed and keyed. Asperoseius Chant is represented by a single known Australian species, A. australiensis Fain and Krantz collected from a Culicoides midge in South Australia. Six species of Euseius Wainstein are recognised in Australia, including four endemics: E. dowdi (Schicha), E. elinae (Schicha), E. neovictoriensis (Schicha), and E. victoriensis (Schicha), and two Asia–Pacific species: E. noumeae (Schicha) and E. ovalis (Evans). Okiseius Ehara is represented in Australia by the previously described O. morenoi Schicha and O. domatorum (Schicha); the Asian species O. subtropicus Ehara (new record); and two new species, O. tribulation and O. cowbay described from tropical rainforest in Far North Queensland. Phytoscutus acaridophagus Collyer, a mite described from New Zealand, is reported in association with acarid mites ( Neotropacarus sp.) in Victoria and in Queensland.     

A new genus of mantidflies discovered in the Oriental region,with a higher‐level phylogeny of Mantispidae (Neuroptera) using DNA sequences and morphology

A remarkable new genus and two new species of Mantispidae (Neuroptera) are described from the Oriental region. Allomantispa Liu, Wu, Winterton & Ohl gen.n. , currently including A. tibetana Liu, Wu & Winterton sp.n. and A. mirimaculata Liu & Ohl sp.n. The new genus is placed in the subfamily Drepanicinae based on a series of morphological characteristics and on the results of total evidence phylogenetic analyses. Bayesian and Parsimony analyses were undertaken using three gene loci (CAD, 16S rDNA and COI) combined with 74 morphological characters from living and fossil exemplars of Mantispidae (17 genera), Rhachiberothidae (two genera) and Berothidae (five genera), with outgroup taxa from Dilaridae and Osmylidae. The resultant phylogeny presented here recovered a monophyletic Mantispidae with ?Mesomantispinae sister to the rest of the family. Relationships among Mantispidae, Rhachiberothidae and Berothidae support Rhachiberothidae as a separate family sister to Mantispidae. Within Mantispidae, Drepanicinae are a monophyletic clade sister to Calomantispinae and Mantispinae. In a combined analysis, Allomantispa gen.n. was recovered in a clade comprising Ditaxis McLachlan from Australia, and two fossil genera from the Palaearctic, ?Promantispa Panfilov (Kazakhstan; late Jurassic) and ?Liassochrysa Ansorge & Schlüter (Germany; Jurassic), suggesting a highly disjunct and relictual distribution for the family. This published work has been registered in ZooBank, http://zoobank.org/urn:lsid:zoobank.org:pub:464B06E8‐47E6‐482E‐8136‐83FE3B2E9D6B .     

New species of Crotonia (Acari: Oribatida) from South Africa

Four new species of oribatid mites of the genus Crotonia , ( C. pauropelor sp. nov., C. dicella sp. nov., C. lyrata sp. nov. and C. ecphylla sp. nov. ) are described from soil and plant litter from South Africa. Crotonia pauropelor, C. dicella and C. ecphylla are allied to the 'capistrata' species-group, previously known only from Queensland, Australia, in that they possess the full complement of c setae on the notogaster. They differ in having eight pairs rather than nine pairs of genital setae. In addition, C. ecpylla is unusual in the genus Crotonia in that it appears to possess two pairs of setae on each of the second epimera. Crotonia lyrata has only two pairs of c setae and is allied to the 'cophinaria' species-group. This is only the second published record of the genus Crotonia from South Africa, and the first of any named species. A key to the African Crotonia species is provided, which includes the only two previously known species C. alluaudi and C. rothschildi (Berlese, 1916).     

The family Capilloventridae (Annelida, Clitellata) in Australia, with descriptions of two new species of Capilloventer

Two new freshwater species of the small family Capilloventridae are recorded from rivers in south-eastern Australia. Capilloventer acheronensis sp. n. has all hair setae anteriorly and dissimilar crotchet setae without hairs posteriorly while Capilloventer longicapitus sp. n. has an elongate prostomium, very long hair setae anteriorly and all bifid setae posteriorly. Immature specimens from south-west Western Australia appear to represent a further species and are tentatively included within the family. Additional specimens of Capilloventer australis Erséus, 1993 have allowed some features (setae, segmental glands, genitalia and ventral buccal organ) to be described in more detail. The new C. australis specimens are from the upper reaches of Victorian rivers, showing that it is a freshwater species, rather than a marine incursion as the estuarine type locality suggested. There are now more freshwater species than marine species, although the origin and phylogenetic relationships of the family remain unclear.     

The family Ctenobelbidae (Acari,Oribatida), with description of a new species and discussion on systematic placement and taxonomic status of the genus Berndamerus Mahunka, 1977

The oribatid mite genus Berndamerus Mahunka, 1977 is transferred into the family Ctenobelbidae as the subgenus Ctenobelba (Berndamerus) Mahunka, 1977, stat. n. from the family Amerobelbidae. The known species of Berndamerus combined: C. (B.) bicostata (Berlese, 1910), comb. n., C. (B.) eremuloides (Berlese, 1910), comb. n., C. (B.) hellenica (Mahunka, 1977), comb. n. A new species, Ctenobelba (Berndamerus) bugiamapensis sp. n., is described from soil, Bu Gia Map National Park, southern Vietnam. It differs from the other species of the subgenus by the heterotrichy of notogastral setae, presence of adanal neotrichy and localization of adanal lyrifissures. Ctenobelbidae is recorded in Vietnam for the first time. A new diagnosis of the family Ctenobelbidae and the identification keys to the known subgenera of the genus Ctenobelba and species of the subgenus Ctenobelba (Berndamerus) are provided.     

Testing the monophyly of the New Zealand and Australian endemic family Conoesucidae Ross based on combined molecular and morphological data (Insecta: Trichoptera: Sericostomatoidea)

Conoesucidae (Trichoptera, Insecta) are restricted to SE Australia, Tasmania and New Zealand. The family includes 42 described species in 12 genera, and each genus is endemic to either New Zealand or Australia. Although monophyly has been previously assumed, no morphological characters have been proposed to represent synapomorphies for the group. We collected molecular data from two mitochondrial genes (16S and cytochrome oxidase I), one nuclear gene (elongation factor 1-α) (2237–2277 bp in total), and 12 morphological characters to produce the first phylogeny of the family. We combined the molecular and morphological characters and performed both a maximum parsimony analysis and a Bayesian analysis to test the monophyly of the family, and to hypothesize the phylogeny among its genera. The parsimony analysis revealed a single most parsimonious tree with Conoesucidae being a monophyletic taxon and sistergroup to the Calocidae. The Bayesian inference produced a distribution of trees, the consensus of which is supported with posterior probabilities of 100% for 15 out of 22 possible ingroup clades including the most basal branch of the family, indicating strong support for a monophyletic Conoesucidae. The most parsimonious tree and the tree from the Bayesian analysis were identical except that the ingroup genus Pycnocentria changed position by jumping to a neighbouring clade. Based on the assumption that the ancestral conoesucid species was present on both New Zealand and Australia, a biogeographical analysis using the dispersal-vicariance criteria demonstrated that one or two (depending on which of the two phylogenetic reconstructions were applied) sympatric speciation events took place on New Zealand prior to a single, late dispersal from New Zealand to Australia.     

Remarkable Rhinophoridae in a growing generic genealogy (Diptera: Calyptratae,Oestroidea)

Four new genera (Apomorphyto gen.n. from Costa Rica, Bixinia gen.n. from Australia, Rhinodonia gen.n. from New Caledonia, Rhinopeza gen.n. from Papua New Guinea) and nine new species (Apomorphyto inbio sp.n. , Bixinia collessi sp.n. , B. solitaria sp.n. , B. spei sp.n. , B. variabilis sp.n. , B. winkleri sp.n. , Rhinodonia antiqua sp.n. , R. flavicera sp.n. , Rhinopeza gracilis sp.n.) of Rhinophoridae (Diptera: Calyptratae, Oestroidea) are described. All new species were included in a morphology‐based phylogenetic analysis to provide arguments for the justification and monophyly (when nonmonotypic) of the new genera and for including these in the Rhinophoridae. The New Caledonian Rhinodonia is a candidate sister taxon to all other rhinophorids, and the Australasian ‘axiniine’ species emerge inside a clade of all Neotropical taxa thus suggesting migration from South America across Antarctica into Australia. This published work has been registered in ZooBank, http://zoobank.org/urn:lsid:zoobank.org:pub:51C1F448‐DDD0‐4F14‐8173‐B8C687F7E841 .     

Anzygina, a new genus for some Australasian microleafhopper species formerly placed in the genus Zygina Fieber (Cicadellidae: Typhlocybinae: Erythroneurini)

Species of Erythroneurini (Cicadellidae: Typhlocybinae) currently placed in the genus Zygina and found in Australia, New Zealand and some neighbouring islands are transferred to the new genus Anzygina , type species Erythroneura sidnica Kirkaldy, following comparison with the type species of the genus: Typhlocyba nivea Mulsant and Rey. New combinations are Anzygina sidnica (Kirkaldy), Anzygina honiloa (Kirkaldy), Anzygina melanogaster (Kirkaldy) and Anzygina sativae (Evans) from Australia, Anzygina toetoe (Cumber), Anzygina agni (Knight), Anzygina dumbletoni (Ghauri) and Anzygina ramsayi (Knight) from New Zealand, Anzygina zealandica (Myers) from Australia and New Zealand, Anzygina jowettae (Knight) from Norfolk Island and Anzygina medioborealis (Ghauri) from Papua New Guinea. Lectotypes are designated for Erythroneura honiloa Kirkaldy and E. sidnica Kirkaldy. Anzygina billi sp.n. is described from SE Qld, and Anzygina barrattae sp.n. is described from the South Island of New Zealand. A. agni is a new record for Australia and is presumed to be Australian in origin. A. dumbletoni has a distribution which suggests that it also is introduced to New Zealand although its origins are not known. A. ramsayi, A. barrattae and A. toetoe , all of which appear to be New Zealand endemics, show affinity with each other based on aedeagal structure. A key to these species, based on males, is provided. The lack of male syntypes for Erythroneura honiala Kirkaldy and Erythroneura lubra Kirkaldy precludes establishment of their identities relative to other species of the genus, and both names are regarded as having nomen dubium status. Australian species not transferred to Anzygina are Zygina evansi (Ross) and Zygina ipoloa (Kirkaldy), both of which belong elsewhere.     

Giselinidae fam. nov., a new monophyletic group of cyclopoid copepods (Copepoda, Crustacea) from the Atlantic deep sea

Four new species of Cyclopoida from deep-sea waters are described and placed in two new genera: Giselina gen. n. and Sensogiselina gen. n. The new genera and species belong to a new monophyletic group within the cyclopinid cyclopoids. A new name, Giselinidae, is proposed for this monophylum. The new family is characterised by the combination of the following characters: (1) tergite of leg 1 fused to cephalosome dorsally, but incompletely fused laterally, (2) absence of aesthetascs on ancestral antennulary segments XVI, XXI and XXV, (3) absence of antennary exopodal setae, (4) presence of only three spines on distal exopodal segment of leg 1, (5) absence of inner setae on first endopodal segments of legs 1–4, (6) outer terminal and distal inner elements of distal endopodal segment of leg 4 transformed into spines, (7) distal outer element of leg 5 exopod transformed into a spine, (8) leg 6 with only one seta, and (9) furcal setae I and III located on dorsal margin. Received in revised form: 16 June 2000 Electronic Publication     

A review of Buccipagoda Ponder, 2010 and descriptions of new species and a new genus (Gastropoda: Neogastropoda: Buccinoidea: Buccinidae)

This revision addresses two Southern Hemisphere genera in the family Buccinidae. Buccipagoda kengrahami (Ponder, 1982) and B. ponderi n. sp. are recognised from off southern Australia and B. achilles n. sp. from off New Zealand. Sagenotriton n. gen. is introduced for S. ajax n. sp. from off New Zealand, and S. bathybius (Bouchet & Warén, 1986) and S. bonaespei (Barnard, 1963) from off South Africa.     

Thinempis, a new genus from Australia and New Zealand (Diptera: Empididae), with notes on the tribal classification of the Empidinae

The genus Thinempis (Diptera: Empididae) is newly described and includes six species, four from Australia – T.austera sp.n., T.esperance sp.n., T.minuta sp.n., and T.turimetta sp.n., and two from New Zealand – T.otakouensis (Miller) comb.n., and T.takaka sp.n. It occurs in coastal habitats and is defined by a unique synapomorphy, a modified triangular female oviscapt (tergum 10). However, the genus shows variable expression in characters considered to be of higher taxonomic importance in the Empidinae. The tribes Hilarini and Empidini are discussed and redefined.     

<Emphasis Type="Italic">Foxtosognus rarus</Emphasis> gen. n., sp. n.—A new genus and species of copepods (Copepoda: Calanoida) from the abyssopelagic zone of the Kuril-Kamchatka Trench

A female of Foxtosognus rarus gen. n., sp. n., a new genus and species of copepods, is described from the abyss of the Kuril-Kamchatka Trench. The new genus is placed in the family Arctokonstantinidae, whose diagnosis is emended and supplemented with the following characters: mandibular basis with one to two setae; mandibular endopod one without setae; maxillular distal basal endite plus endopod with two to four setae or without setae. The features that distinguish Foxtosognusgen. n. from other representatives of this family are as follows: mandibular palp with a long endopod segment one, endopod segment two with seven setae, and exopod segment five with two setae; maxillula with distal basal endite lacking setae and separated from endopod bearing three setae; maxillular precoxal arthrite with seven to eight setae and exopod with four setae; maxilliped syncoxa without setae on precoxal endites and endopod of three segments. The genera Sognocalanus and Foxtonia earlier included in the Spinocalanidae are herein transferred to the Arctokonstantinidae.     

A new oribatid mite species of the genus Protoribates (Acari,Oribatida, Haplozetidae) from Ethiopia

A new oribatid mite species, Protoribates aethiopicus sp. n., from Ethiopia is described. This species is morphologically similar to Protoribates dentatus (Berlese, 1883), but differs from it in the longer lamellae and notogastral setae, shorter adanal setae ad ₁ and smaller notogastral porous areas, as well as in the presence of two claws on all legs.     

Three new genera of Braconini from Australasia and Malaysia (Insecta, Hymenoptera, Braconidae)

Ancilibracon gen.n. (type species: Ancilinracon townesi sp.n.). Ancilibracon bakeri sp.n., Gelasinibracon gen.n. (type species: Gelasinibracon sedlaceki sp.n.), Gelasinibracon simplicicaudatus sp.n., Esengoides gen.n. (type species: Esengoides crentdatus sp.n.) and Esengoides fulvus sp.n. are each described and illustrated. Ancilibracon towensi is from Malaysia. A. bakeri is from Borneo, both Gelasinibracon species are from Guinéa, E. crenulatus is from the solomon Islands and E. fulvus is from Australia. These three new genera belong to the Plesiobrscon Cameron group of the Braconina. Their relationship are discussed.