Beetles that live with ants (Carabidae,Pseudomorphini, Pseudomorpha Kirby, 1825): A revision of the santarita species group

The Western Hemisphere genus Pseudomorpha Kirby 1825 was last revised by Notman in 1925 based on only a few known species (22) and paltry few specimens (73); other authors have added an additional six species represented by 53 additional specimens since 1925. Baehr (1997) assigned three species from Australia to this genus, albeit in a new subgenus, Austropseudomorpha Baehr 1997. A recent study of collections from throughout the Americas (1757 specimens) has revealed numerous new species that can be arrayed across 19 species groups based on a suite of attributes, some used by Notman and others newly discovered. A taxonomic revision of the species contained in one of these species groups, santarita, is provided herein, as well as a distributional synopsis of the remaining 18 species groups. New species described herein are as follows, each with its type locality: Pseudomorpha huachineras p. n., Arroyo El Cocono, Sierra Huachinera, Sonora, México; P. patagonia sp. n., Madera Canyon, Santa Rita Mountains, Arizona; P.penablanca sp. n., Peña Blanca Lake, Arizona; P. pima sp. n., Madera Canyon (lower), Santa Rita Mountains, Arizona; P. santacruz sp. n., Madera Canyon, Santa Rita Mountains, Arizona; and P. santarita sp. n., Santa Rita Ranch, Santa Rita Mountains, Arizona.     

Two new species of Euglossa from South America,with notes on their taxonomic affinities (Hymenoptera,?Apidae)

Two new species of the genus Euglossa Latreille, subgenus Glossurella Dressler are here presented. Euglossa (Glossurella) embera sp. n., from the Pacific lowlands of Colombia, and Euglossa (Glossurella) adiastola sp. n., from the Atlantic Forest of Brazil. Their taxonomic association and distinction are discussed, as well as the correct understanding of the subgenus Glossurella.     

Three new species of the genus Probles F?rster (Hymenoptera,Ichneumonidae, Tersilochinae) from?South Korea

Three closely related species of the genus Probles Förster, P. fulgida sp. n., P. korusa sp. n. and P. rukora sp. n., belong to the subgenus Euporizon Horstmann and differ from other Palearctic species of the genus by a combination of long and apically weakly sinuate ovipositor and short temple. These three species are assigned to a newly designated fulgida species-group, and a portion of the key for identification of this species-group is provided. Based on the shape of the ovipositor apex, the fulgida species-group resemble members of the subgenus Microdiaparsis Horstmann but are distinct in having a much shorter temple.     

Taxonomic status of cestodes of the genus Paracaryophyllaeus Kulakowskaja, 1961 (Caryophyllidea: Caryophyllaeidae)

A taxonomic revision of cestodes of the genus Paracaryophyllaeus Kulakowskaja, 1961, parasites of cobitid fishes, has been performed. Three species (Paracaryophyllaeus gotoi (Motomura, 1927), P. kulakowskae sp. n., and P. misgurni sp. n.) are considered valid species; two are described as new species. P. dubininorum Kulakowskaja, 1961 is a young synonym of P. gotoi. The taxonomic status for P. gotoi sensu Scholz, 1989, P. gotoi sensu Scholz et al., 2001, and Paracaryophyllaeus sp. 1 and Paracaryophyllaeus sp. 2 (described in present work) needs specification.     

Revision of the subgenus Ceratitis (Pardalaspis) Bezzi, 1918 (Diptera, Tephritidae, Ceratitini)

Abstract. The fruit fly genus Ceratitis (Diptera: Tephritidae) comprises several important pest species attacking a wide range of unrelated fruits. In this paper the subgenus Ceratitis ( Pardalaspis ) Bezzi is revised. Ten species are recognized of which five are described as new: C.(P.)hamata sp.n., C.(P.)munroi sp.n., C.(P.)serrata sp.n., C.(P.)semipunctata sp.n. and C.(P.)zairensis sp.n. A key to both sexes is provided. All species are restricted to the Afrotropical Region and distributional and known host plant data are given.     

Three new species of ants (genus <Emphasis Type="Italic">Aphaenogaster</Emphasis>, Hymenoptera: Formicidae) from Turkey

Three new species of the ant genus Aphaenogaster (A. aktaci sp. n., A. maculifrons sp. n., A. radchenkoi sp. n.) are described from Turkey and their taxonomic positions are discussed. Aphaenogaster aktaci sp. n. belongs to the obsidiana group, A. maculifrons sp. n. to subterranea group and A. radchenkoi sp. n. to pallida group of the Attomyrma subgenus. Key characteristics for identification of the species and data on their ecology are given.     

Five new species of subgenus Plesiominettia (Diptera,Lauxaniidae, Minettia) in southern China,with a key to known species

Five species of the subgenus Plesiominettia Shatalkin from the southern China are described as new to science: Minettia (Plesiominettia) flavoscutellata sp. n., Minettia (Plesiominettia) longaciculiformis sp. n., Minettia (Plesiominettia) nigrantennata sp. n., Minettia (Plesiominettia) tridentata sp. n. and Minettia (Plesiominettia) zhejiangica sp. n. One species, Minettia longistylis Sasakawa, is transferred to the subgenus Plesiominettia from Minettia s. str. A key to separate the known species of the subgenus is presented, along with a taxonomic list of species. The type materials of the new species are deposited in the China Agricultural University, Beijing, China (CAUC).     

New species of the bristletail family machilidae (insecta,microcoryphia) from Abkhazia

Six new species are described from Abkhazia: Charimachilis abchasica sp. n., Lepismachilis abchasica sp. n., Trigoniophthalmus longitarsus sp. n., T. nematocerus sp. n., T. abchasicus sp. n., and T. subalpinus sp. n. Species of the genus Lepismachilis and the subgenus Trigoniophthalmus s. str. (T. longitarsus sp. n.) are described from the Caucasus for the first time. A key to the species of the genus Trigoniophthalmus from the Caucasus is given.     

Integrative revision of the giant pill-millipede genus Sphaeromimus from Madagascar,with the description of seven new species (Diplopoda,Sphaerotheriida, Arthrosphaeridae)

The Malagasy giant pill-millipede genus Sphaeromimus de Saussure & Zehntner, 1902 is revised. Seven new species, S. titanus sp. n., S. vatovavy sp. n., S. lavasoa sp. n., S. andohahela sp. n., S. ivohibe sp. n., S. saintelucei sp. n., and S. andrahomana sp. n. were discovered, in one case with the help of sequence data, in the rainforests of southeastern Madagascar. The species are described using light- and scanning electron microscopy. A key to all 10 species of the genus is presented. All but one (S. andohahela) of the newly discovered species are microendemics each occurring in isolated forest fragments. The mitochondrial COI barcoding gene was amplified and sequenced for 18 Sphaeromimus specimens, and a dataset containing COI sequences of 28 specimens representing all Sphaeromimus species (except S. vatovavy) was analyzed. All species are genetically monophyletic. Interspecific uncorrected genetic distances were moderate (4–10%) to high (18–25%), whereas intraspecific variation is low (0–3.5%). Sequence data allowed the correct identification of three colour morphs of S. musicus, as well as the identity of a cave specimen, which although aberrant in its morphology and colouration, was genetically identical to the holotype of S. andrahoma.     

A new genus and eight new species of the subtribe Anillina (Carabidae,Trechinae, Bembidiini) from Mexico,with a cladistic analysis and some notes on?the?evolution of the genus

One new genus and eight new species of anilline carabids are described from southern Mexico. The new genus, Zapotecanillus gen. n., is established for Z. oaxacanus (type species) sp. n., Z. nanus sp. n., Z. iviei sp. n., Z. ixtlanus sp. n., Z. montanus sp. n., and Z. kavanaughi sp. n. from the Sierra Madre de Oaxaca, Z. pecki sp. n. from the Sierra Madre del Sur, and Z. longinoi sp. n. from the Sierra Madre de Chiapas. A taxonomic key for all described species of Zapotecanillus and a cladistic analysis, based on morphological data, are provided. Morphological, behavioral and biogeographical aspects of the speciation in the genus obtained from the resulting cladogram are discussed.     

On discovery of the weevil genus <Emphasis Type="Italic">Nanomicrophyes</Emphasis> Pic, 1908 (Coleoptera,Curculionidae) in China

A new subgenus Sinomicrophyes subgen. n. with four new species Nanomicrophyes belousovi sp. n. (type species), N. korolevi sp. n., N. oblongus sp. n., and N. kabaki sp. n. from Western China is erected in the genus Nanomicrophyes Pic, 1908 previously known from a single species distributed in the Caucasus. A map of the genus Nanomicrophyes range is provided with a brief discussion of the N. cyanipennis (Weise, 1880) distribution.     

The family Ctenobelbidae (Acari,Oribatida), with description of a new species and discussion on systematic placement and taxonomic status of the genus Berndamerus Mahunka, 1977

The oribatid mite genus Berndamerus Mahunka, 1977 is transferred into the family Ctenobelbidae as the subgenus Ctenobelba (Berndamerus) Mahunka, 1977, stat. n. from the family Amerobelbidae. The known species of Berndamerus combined: C. (B.) bicostata (Berlese, 1910), comb. n., C. (B.) eremuloides (Berlese, 1910), comb. n., C. (B.) hellenica (Mahunka, 1977), comb. n. A new species, Ctenobelba (Berndamerus) bugiamapensis sp. n., is described from soil, Bu Gia Map National Park, southern Vietnam. It differs from the other species of the subgenus by the heterotrichy of notogastral setae, presence of adanal neotrichy and localization of adanal lyrifissures. Ctenobelbidae is recorded in Vietnam for the first time. A new diagnosis of the family Ctenobelbidae and the identification keys to the known subgenera of the genus Ctenobelba and species of the subgenus Ctenobelba (Berndamerus) are provided.     

Contribution to the knowledge of the weevil genus Otiorhynchus (Coleoptera, Curculionidae) from northeastern Turkey and Transcaucasia

Otiorhynchus (Podonebistus) korotyaevi sp. n., (Proremus) karsavurani sp. n., and O. (Proremus) temeli sp. n. from northeastern Turkey are described. O. kirschi is transferred from the genus Simo (= Homorhythmus) to the genus Otiorhynchus (subgenus Proremus); O. madari, from the genus Cirrorhynchus to the genus Otiorhynchus (subgenus Motilacanus); and O. kataevi, from the subgenus Nilepolemis to the subgenus Otismotilus. New data on the morphology and distribution of O. cataphractus, O. latinasus, O. terrifer, O. pipitzi, O. angustifrons, O. jarpachlinus, and O. sculptirostris are given. Lectotypes of O. lederi, O. erivanensis, O. diotus, O. terrifer, O. kirschi, and O. madari are designated.     

Two new species of the genus Philaenus (Homoptera, Aphrophoridae) from Iran

Two new species of Philaenus from Iran are described. Philaenus elbursianus sp. n. belongs to the Ph. signatus species group from the nominotypical subgenus. Philaenus iranicus sp. n. differs considerably from all the known representatives of the genus and was separated in a new subgenus Gyrurus subgen. n. The occurrence of this highly peculiar species in Iran may indicate that the center of origin of the genus Philaenus lies in this territory and not in the Mediterranean region as it was considered before.     

New and little-known crickets of the subfamily Phalangopsinae (Orthoptera,Gryllidae). 10. The genus <Emphasis Type="Italic">Parendacustes</Emphasis> (Part 1)

The composition of the genus Parendacustes Chop. and of its two subgenera is briefly discussed. Nine new taxa of the nominotypical subgenus are described: P. berezini sp. n., P. arachne sp. n., P. pahangi borealis subsp. n., P. sylvestris sp. n., P. tkatshevae sp. n., P. tawau sp. n., P. tiomani sp. n., P. ectoparameralis sp. n., and P. magnispeculum sp. n. New data on the distribution of P. p. pahangi Gor. and considerations on the generic position of P. latifrons Chop. and P. lifouensis Desutter-Grandcolas are also provided.     

Revision of the Neotropical bark mantis genus Liturgusa Saussure, 1869 (Insecta,Mantodea, Liturgusini)

The praying mantis genus Liturgusa Saussure, 1869 occurs only in Central and South America and represents the most diverse genus of Neotropical Liturgusini (Ehrmann 2002). The genus includes bark dwelling species, which live entirely on the trunks and branches of trees and run extremely fast. All species included within the genus Liturgusa are comprehensively revised with a distribution stretching from central Mexico, the island of Dominica to the southeastern regions of Brazil and southern Bolivia. All known species are redescribed to meet the standards of the new treatment of the genus (11 species). Three new genera are described including Fuga gen. n., Velox gen. n., and Corticomantis gen. n. for species previously included in Liturgusa as well as Hagiomantis. Liturgusa mesopoda Westwood, 1889 is moved to within the previously described genus Hagiomantis Audinet Serville, 1838. A total of 19 species are newly described within Liturgusa, Fuga, and Velox including L. algorei sp. n., L. bororum sp. n., L. cameroni sp. n., L. cura sp. n., L. dominica sp. n., L. fossetti sp. n., L. kirtlandi sp. n., L. krattorum sp. n., L. manausensis sp. n., L. maroni sp. n., L. milleri sp. n., L. neblina sp. n., L. purus sp. n., L. stiewei sp. n., L. tessae sp. n., L. trinidadensis sp. n., L. zoae sp. n., F. grimaldii sp. n., and V. wielandi sp. n. Four species names are synonymized: Liturgusa peruviana Giglio-Tos, 1914, syn. n. = Liturgusa nubeculosa Gerstaecker, 1889 and Hagiomantis parva Piza, 1966, syn. n., Liturgusa sinvalnetoi Piza, 1982, syn. n., and Liturgusa parva Giglio-Tos, 1914, syn. n. = Mantis annulipes Audinet Serville, 1838. Lectotypes are designated for the following two species: Liturgusa maya Saussure & Zehntner, 1894 and Fuga annulipes (Audinet Serville, 1838). A male neotype is designated for Liturgusa guyanensis La Greca, 1939. Males for eight species are described for the first time including Liturgusa cayennensis Saussure, 1869, Liturgusa lichenalis Gerstaecker, 1889, Liturgusa guyanensis La Greca, 1939, Liturgusa maya Saussure & Zehntner, 1894, Liturgusa nubeculosa Gerstaecker, 1889, Fuga annulipes (Audinet Serville, 1838), Corticomantis atricoxata (Beier, 1931), and Hagiomantis mesopoda (Westwood, 1889). The female of Fuga fluminensis (Piza, 1965) is described for the first time. Complete bibliographic histories are provided for previously described species. The spelling confusion surrounding Liturgusa/Liturgousa is resolved. Full habitus images for males and females are provided for nearly all species. Habitus and label images of type specimens are provided when possible. Diagnostic illustrations of the head and pronotum for males and females are provided for all species when possible. Illustrations of male genital structures are provided for all species for which males are known. Measurement data, including ranges and averages, are provided for males and females of all species. Combined male and female genus and species level dichotomous keys are provided with a Spanish translation. A complete table of all examined specimens lists label data, museum codes, repositories, and other specimen specific information. A KML file with all georeferenced locality records is downloadable from mantodearesearch.com for viewing in Google Earth. Natural history information is provided for species observed by the author.     

Five new species and a new subgenus of the mite genus <Emphasis Type="Italic">Premicrodispus</Emphasis> (Acari,Heterostigmata, Microdispidae) from Turkmenistan

A new monotypic subgenus, Premicrodispulus subgen. n., with the type species Premicrodispus reductus sp. n., and four new species of the nominative subgenus of the mite genus Premicrodispus, P. (Premicrodispus) paradoxus sp. n., P. (P.) heterocaudatus sp. n., P. (P.) obtusisetosus sp. n., and P. (P.) incisus sp. n., collected in soils of Turkmenistan are described. The subgenus Premicrodispulus subgen. n. differs from the nominative subgenus in the presence of 3 setae on genu I (seta l″ absent) and 1 seta on genu II (setae d and l″ absent). Premicrodispus (Premicrodispus) paradoxus sp. n. differs from all the species of the genus in the presence of solenidion on tarsus III. P. (P.) heterocaudatus sp. n. is most similar to P. longisetosus (Mahunka, 1970), but differs in the setae ps3 distinctly longer than ps ₁ (in P. longisetosus, ps ₃ and ps ₁ are similar in length). P. (P.) obtusisetosus sp. n. is most similar to P. montanus Khaustov, 2006, but differs in obtuse setae c ₁, e, and f (in P. montanus, these setae are pointed); P. (P.) incisus sp. n. is most closely related to P. longisetosus (Mahunka, 1970), but differs in the presence of distinct excavations on the posterior margin of tergites C and D (in P. longisetosus, excavations are absent).     

Taxonomy of the genus Peyerimhoffia Kieffer from Mainland China,with a description of seven new species (Diptera,Sciaridae)

The taxonomy of the genus Peyerimhoffia Kieffer in China was studied. Eight species were recognized, including seven new species that are herein described and illustrated: P. hamata sp. n., P. obesa sp. n., P. sparsula sp. n., P. longiprojecta sp. n., P. brachypodua sp. n., P. yunnana sp. n., and P. shennongjiana sp. n. In addition, P. vagabunda (Winnertz, 1867) is reported for the first time from China. A key to these Chinese species is provided.