New insects (Insecta: Mecoptera,Grylloblattida) from the Middle Permian Chepanikha locality,Udmurtia

New scorpionflies, Asiachorista europaea sp. nov. and Petromantis udmurtica sp. nov. (Mecoptera: Permochoristidae), and new grylloblattids, Tshepanichoptera lacera gen. et sp. nov. (Grylloblattida: Aliculidae) and Miralioma urzhumica sp. nov. (Liomopteridae), are described from the Urzhumian of Udmurtia (Chepanikha locality). Liomopterites novissimus Aristov, 2004 (Liomopteridae) is redescribed.     

New fossil hangingflies (Insecta: Mecoptera: Raptipedia: Cimbrophlebiidae) from the Middle Jurassic of Inner Mongolia,China

Abstract: Two new genera with six new species of hangingflies (Mecoptera: Raptipedia: Cimbrophlebiidae) are described. They were collected from the Middle Jurassic nonmarine sedimentary strata of the Jiulongshan Formation in Daohugou, Inner Mongolia, north‐eastern China. This report of Cimbrophlebiidae from the Yanliao Biota provides detailed morphological information and evidence of a broad diversity of Cimbrophlebiidae during the Middle Jurassic. A key to the known species of Cimbrophlebiidae is provided.     

Middle Jurassic fossils of the genus Sharasargus from Inner Mongolia, China (Diptera: Archisargidae)

To date, only two species of the genus Sharasargus are known. These Upper Jurassic species were found in Mongolia and Kazakhstan, respectively. We herein describe two new species from the Middle Jurassic, which were found in Inner Mongolia, China. They are the oldest known fossils of this genus. A key to Sharasargus species is given.     

The earliest fossil record of Panorpidae (Mecoptera) from the Middle Jurassic of China

The early history of Panorpidae (Mecoptera) is poorly known due to sparse fossil records. Up to date, only nine fossil species have been described, all from the Paleogene, except the Early Cretaceous Solusipanorpa gibbidorsa Lin, 1980. However, we suggest S. gibbidorsa is too incompletely preserved to permit even family classification. A new genus with two new species, Jurassipanorpa impunctata gen. et sp. n. and Jurassipanorpa sticta sp. n., are described based on four well-preserved specimens from the late Middle Jurassic Jiulongshan Formation of Daohugou, Inner Mongolia, China. These two new species are the earliest fossil records of Panorpidae. The new genus is erected based on a combination of forewing characters: both R₁ and Rs₁ with two branches, 1A reaching posterior margin of wing distad of the forking of Rs from R_1, and no crossveins or only one crossvein between veins of 1A and 2A. In all four specimens, long and robust setae ranging from 0.09 to 0.38 mm in length and pointing anteriorly, are present on anal veins of forewings. The function of these setae is enigmatic.     

Tettigarctidae (Insecta: Hemiptera: Cicadoidea) from the Middle Jurassic of Inner Mongolia, China

Three new species of Tettigarctidae from the Middle Jurassic of Daohugou, China are described: Shuraboprosbole daohugouensis nov. sp., S. minuta nov. sp., and S. media nov. sp. The revised diagnostic characters of Shuraboprosbole Becker-Migdisova are provided based on well-preserved whole-bodied fossil tettigarctids. So far, only four definitive species of Tettigarctidae are recorded from the Mesozoic of China. The pronotum concealing major of mesonotum seems to be an autapomorphy of Tettigarctinae. The “cicadellid-like tarsi” found in Cicadellidae and Brazilian Tettigarctidae are probably a parallel evolution.     

中国内蒙古道虎沟中侏罗世蚊蝎蛉科(昆虫纲,长翅目)昆虫化石

描述蚊蝎蛉科1新属和3新种:新属Decoribittacus gen.nov.,新种美脉华美蚊蝎蛉Decoribittacus euneurus sp.nov.,斑点华美蚊蝎蛉Decoribittacus stictus sp.nov.和长翼卡拉蚊蝎蛉Karattacus longia latus sp.nov..同时修订了卡拉蚊蝎蛉属Karattaus的属征.所有的化石标本都来自内蒙古宁城县道虎沟中侏罗世九龙山组地层,模式标本现在保存于首都师范大学生命科学学院.     

New Ephialtitidae (Insecta: Hymenoptera) from the Jurassic Daohugou Beds of Inner Mongolia,China

Five new species are described from the Middle–Upper Jurassic Daohugou Beds of Inner Mongolia, China, and assigned to the genus Karataus Rasnitsyn, 1977 in the subfamily Symphytopterinae of Ephialtitidae. They are illustrated as Karataus daohugouensis n. sp., K. strenuus n. sp., K. vigoratus n. sp., K. exilis n. sp., and K. orientalis n. sp., and a key to species of Karataus is presented.     

内蒙古道虎沟中侏罗世中生蝎蛉科(昆虫纲,长翅目)昆虫化石新发现

描述中生蝎蛉科Mesopanorpodidae Tillyard,1918化石2新属3新种:宽翅柱状蝎蛉Stylopanorpodes eurypterus gen.et sp.nov.,华美纺锤蝎蛉Netropanorpodes decorosus gen.et sp.nov.,多刺纺锤蝎蛉Netropanorpodes sentosus gen.et sp.nov..所有化石标本均采于内蒙古宁城道虎沟中侏罗世九龙山组地层,模式标本现保存于首都师范大学生命科学学院.     

The first agetopanorpine mecopteroid insect from Middle Permian of China (Insecta: Mecoptera: Permochoristidae)

Sinoagetopanorpa permiana n. gen., n. sp., first Chinese Permian Agetopanorpinae, is described and compared to the other genera of this group.     

New ommatids from the Late Jurassic of western Liaoning, China （Coleoptera： Archostemata）

A new genus Amblomma gen. nov. of fossil beetles is erected and can be assignedto the family Ommatidae because its two procoxal cavities are contiguous and the articulations of the abdominal ventrites are abutting. The new genus is similar to Zygadenia Handlirsch, 1906 (=Notocupes Ponomarenko, 1964), Tetraphalerus Waterhouse, 1901,Rhobdocupes Ponomarenko, 1966 and Sinocupes Lin, 1976, but can be distinguished from other genera according to the following characters: the second segment of antennae is shorterthan the third one in length; the posterior tarsi with the basal segment is obviously shorter than the three following taken together in length; the antennae reach the posterior ridge of prothroax in length, and the sides of the prothroax with serrulate margin. Four new species of the new genus are described and figured: Amblomma psilata gen. et sp. nov., Amblomma rudis gen. et sp. nov., Amblomma epicharis gen. et sp. nov., and Amblomma stabilis gen. etsp. nov. A key to species within this new genus is provided. All the specimens are collected from the Late Jurassic Yixian Formation of western Liaoning and are now housed in the College of Life Science, Capital Normal University, Beijing, China.     

蝎蛉属Panorpa一新种记述(长翅目:蝎蛉科)

记述采自河南省王屋山的蝎蛉属Panorpa Linn．1新种：王屋山蝎蛉Panorpa wangwushana,sp．nov．。模式标本分别保存在西北农林科技大学昆虫博物馆和郑州大学生物系。     

New archisargids from China (Insecta: Diptera)

The extinct family Archisargidae, which was rich in the Mesozoic, has been found in China, Kazakhstan and Mongolia. By now, six genera and 14 species have been found in Inner Mongolia and Liaoning Province of China. In this paper, two new species of the genus Mesosolva Hong, 1983 and a new genus Brevisolva gen. nov., represented by a new species belonging to the Archisargidae, are described from the Middle Jurassic deposits of Daohugou, Inner Mongolia. A key to the species of Mesosolva is given.     

Four new species of hangingflies (Insecta,Mecoptera, Bittacidae) from the Middle Jurassic of northeastern China

Two new species of Mongolbittacus Petrulevičius, Huang & Ren, 2007, Mongolbittacus speciosus sp. n. and Mongolbittacus oligophlebius sp. n., and two new species of Exilibittacus Yang, Ren & Shih, 2012, Exilibittacus foliaceus sp. n. and Exilibittacus plagioneurus sp. n., in the family Bittacidae, are described and illustrated based on five well-preserved fossil specimens. These specimens were collected from the late Middle Jurassic Jiulongshan Formation of Daohugou, Inner Mongolia, China. These new findings enhance our understanding of the morphological characters of early hangingflies and highlight the diversity of bittacids in the Mid Mesozoic ecosystems.     

Three distinct but rare kovalevisargid flies from the Jurassic Daohugou biota,China (Insecta,Diptera, Brachycera,Kovalevisargidae)

Abstract: Three nearly complete male specimens of kovalevisargid flies are described as Kovalevisargus macropterus sp. nov., K. brachypterus sp. nov. and Kerosargus sororius sp. nov. (family Kovalevisargidae) from the Callovian–Oxfordian Daohugou biota in Inner Mongolia, China. These extend the range of the family Kovalevisargidae outside of Central Asia for the first time, reveal new morphological details about kovalevisargid flies and offer new evidence for the biostratigraphic correlation of nonmarine sedimentary strata of both the Karabastau and Daohugou Formations. Close similarities in the composition of insect taxa from both entomofaunas imply not only the geological age but also the sedimentary environment at that time being the same, or nearly so. Familial and generic diagnoses of kovalevisargid flies are supplemented based on information derived from these new species.     

Pronemouridae fam. nov. (Insecta: Plecoptera), the stem group of nemouridae and notonemouridae,from the Middle Jurassic of Inner Mongolia,China

Abstract: A new family, Pronemouridae fam. nov., with a new genus, Pronemoura gen. nov., and five new species of fossil stoneflies are described from Daohugou village (Middle Jurassic), Inner Mongolia, China: P. shii sp. nov., P. angustithorax sp. nov., P. longialata sp. nov., P. minuta sp. nov. and P. peculiaris sp. nov. Wing venation of pronemourinids presents some or considerable variety in the fore and hind wings of the same or different individuals. Pronemourinidae, retaining plesiomorphic characters (short, multisegmented cerci and CuA with forks), should be treated as the stem group of Nemouridae and Notonemouridae. Based on fossil data, we propose a model in which the extant plecopterid distribution began in the Early Cretaceous. Nemouridae and Notonemouridae must have occurred in Asia and begun to migrate from here by at least the Early Cretaceous.     

Two new species of Sinosmylites Hong (Neuroptera, Berothidae) from the Middle Jurassic of China, with notes on Mesoberothidae

Two new species of the genus Sinosmylites Hong are described from the Middle Jurassic locality at Daohugou (Inner Mongolia, China): Sinosmylites fumosussp. n. and Sinosmylites rasnitsynisp. n. This is the oldest known occurrence of the family Berothidae. The berothid affinity of this genus is confirmed by examination of the hind wing venation characteristic of the family. The Late Triassic family Mesoberothidae may represent an early group of Berothidae.     

First record of Eretmophyllum (Ginkgoales) with well-preserved cuticle from the Middle Jurassic of the Ordos Basin,Inner Mongolia,China

The genus Eretmophyllum, a characteristic and widespread taxon of the order Ginkgoales, is restricted in the Mesozoic floras of Eurasia. In China, although some specimens were assigned to Eretmophyllum based on gross leaf morphology, none have epidermal structure, which is essential for species delimitation within this genus. In this paper, we describe a new species, Eretmophyllum neimengguensis n. sp., from the Middle Jurassic Yan’an Formation of the Ordos Basin, China. This is the first record of the genus Eretmophyllum (Ginkgoales) from the Ordos Basin as well as from China based upon the gross leaf morphology and the epidermal structure. The new record significantly extends the stratigraphic and geographic distribution of Eretmophyllum in China and Eurasia. It also improves our understanding of its leaf morphology, epidermal structure, and past diversity. The taxonomic position and the stratigraphic and geographical range of the genus Eretmophyllum are discussed. Briefly, leaf morphological and epidermal characters of the new species, associated plant assemblages, and sedimentology indicate that the regional climate of the Ordos Basin was warm and humid with seasonal temperature and precipitation fluctuations in warm-temperate zone during the Middle Jurassic. Furthermore, given the relationship between the distribution/diversity of the genus and the climate, the genus Eretmophyllum may be confined to the warm-temperature climate, is intolerant of heat and frost, and can be considered as an indicator of humid and warm climatic conditions.