Ladinian radiolarian fauna from West Sichuan, China

A poorly preserved, but diversified radiolarian fauna was recovered from thin-bedded cherts occurring in the southeast of Daofu, Sichuan Province, southwestern China. Twenty-two radiolarian species belonging to 10 genera are identified and three new species (Paroertlispongus daofuensis n. sp., Falcispongus heinzi n. sp., Falcispongus pauliani n. sp.) are described. Falcispongus heinzi represents a transitional species between genera Oertlispongus inaequispinosus Dumitrica, Kozur and Mostler and Falcispongus Dumitrica. The fauna is divided into two assemblages, namely Muelleritortis cochleata and Oertlispongus inaequispinosus assemblages, which can be well correlated with the Ladinian radiolarian zones from Europe. The fauna indicates that basaltic rocks occurring in the Xianshuihe Belt were formed mainly during the Middle Triassic. Our results thus extend the previously known geological age of the Xianshuihe Belt to the Middle and Late Triassic interval.     

Late Triassic radiolarians of southern Cyprus

Radiolarians from the Upper Triassic of the allochthonous Mamonia Assemblage of southern Cyprus are considered. The Phasoula Formation, composed of basic volcanics, with lenses and interbeds of micritic limestones and cherts, contains (1) a Lower Norian assemblage with Capnodoce crystallina-Trialatus robustus, which also includes Capnodoce anapetes De Wever, Capnuchosphaera deweveri Kozur et Mostler, C. theloides De Wever, Deflandrecyrtium curvatum Kozur et Mostler, Icrioma cruciformis Tekin, Kahlerosphaera norica Kozur et Mock, Kinyrosphaera helicata Bragin, Mostlericyrtium sitepesiformis Tekin, Palaeosaturnalis latiannulatus Kozur et Mostler, Spongostylus tortilis Kozur et Mostler, Xiphotheca rugosa Bragin, and Zhamojdasphaera proceruspinosa Lahm; (2) a Middle Norian assemblage with Capnodoce sarisa accompanied by Loffa mulleri Pessagno, Nabolella trispinosa Bragin, and Praexehasaturnalis tenuispinosus (Donofrio et Mostler); and (3) an Upper Norian assemblage with Livarella densiporata-Lysemelas olbia accompanied by Pentactinocarpus sevaticus Kozur et Mostler, Praemesosaturnalis multidentatus (Kozur et Mostler), and others. This assemblage also occurs in clastic turbidites of the Vlambouros Formation. In the sections of southern Cyprus, radiolarian zones are recognized that correspond to the zones previously established in the Far East of Russia, which include Capnodoce crystallina (Lower and Middle Norian) and Lysemelas olbia (lower part of the Upper Norian). Radiolarians belonging to three orders, 24 families, 59 genera, and 101 species are described; of them 2 genera, 9 species, and 1 subspecies were previously described by the author; 14 new species and 1 new subspecies are established. The diagnoses of many genera and species are emended, the stratigraphic and geographical ranges of the majority of taxa are substantially expanded.     

Middle Triassic radiolarians from the Southeastern Pamirs (Republic of Tajikistan)

Triassic radiolarians were studied from a limestone/chert succession of the Dzhangisu and Boztere formations, Southeastern Pamirs, Republic of Tajikistan. The middle part of the Dzhangisu Formation is characterized by a Late Anisian assemblage with Eptingium nakasekoi Kozur and Mostler, Archaeospongoprunum bispinosum Kozur and Mostler, A. mesotriassicum mesotriassicum Kozur and Mostler, Paroertlispongus multispinosus Kozur and Mostler, P. rarispinosus Kozur and Mostler and Hozmadia rotunda (Nakaseko and Nishimura) that belongs either to the upper part of the Tetraspinocyrtis laevis Zone or to the Spongosilicarmiger transitus Zone (Kozur, 2003). The lowermost part of the Boztere Formation is characterized by a Late Ladinian assemblage with Muelleritortis cochleata (Nakaseko and Nishimura), M. expansa (Kozur and Mostler), Spongoserrula rarauana Dumitrica that corresponds to the M. cochleata Zone. Pamirian localities have a transitional position between the Western Tethys (Alpine–Mediterranean region) and the Eastern Tethys–Pacific junction (Southern China and Thailand). The assemblages are Tethyan in affinity.     

Latest Permian Palaeolithocycliidae (Radiolaria) from South China

The family Palaeolithocycliidae Kozur and Mostler was formerly thought to be ranging from Devonian to Carboniferous. However, a lot of radiolarians assignable to this family were discovered in the late Changhsingian (Latest Permian) of South China. Six species, including Palaeolithocyclia pilata n. sp., Palaeolithocyclia platta n. sp., Paramphibrachium woni n. sp., Klaengspongus spinosus Sashida, Klaengspongus formosus n. sp. and Klaengspongus umbilicatus n. sp., have been identified. In this paper, these species are illustrated and described.     

Triassic radiolarians of Kotel’nyi Island (New Siberian Islands,Arctic)

Triassic radiolarians from Kotel’nyi Island (New Siberian Islands, Arctic Region) are studied. Radiolarians occur in the Middle and Upper Triassic beds, which are well dated by ammonoids, nautiloids, and bivalves. In the Middle Triassic beds, which are composed of dark gray and black clays and claystones with interbeds of combustible schist, siltstone, clayey limestone, and many phosphatic concretions, the following two assemblages are recognized: (1) a Late Anisian assemblage, including Glomeropyle clavatum Bragin, sp. nov., G. boreale Bragin, G. manihepuaensis Aita, G. insulanum Bragin, sp. nov., Triassospongosphaera multispinosa (Kozur et Mostler), Tetraspongodiscus hibernus Bragin, sp. nov., T. borealis Bragin, sp. nov., Eptingium abditum Bragin, sp. nov., E. manfredi Dumitrica, Spongostephanidium japonicum (Nakaseko et Nishimura), and Ladinocampe vicentinensis Kozur et Mostler; (2) Late Ladinian assemblage, with Muelleritortis firma (Gorican), M. kotelnyensis Bragin, sp. nov., Tiborella nivea Bragin, sp. nov., Pseudostylosphaera goestlingensis (Kozur et Mostler), P. omolonica Bragin, Triassospongosphaera multispinosa (Kozur et Mostler), Sarla cincinnata Bragin, sp. nov., S. obscura Bragin, sp. nov., S. prava Bragin, sp. nov., and Eonapora stiriaca Bragin, sp. nov. The Upper Triassic beds, composed of gray and black clays and claystones with lenses of clayey limestones, with sideritic and phosphatic-calciferous concretions have yielded (1) an Early Carnian assemblage, with Pentactinocarpus colum Bragin, sp. nov., Glomeropyle cuneum Bragin, sp. nov., G. algidum Bragin, sp. nov., G. aculeatum Bragin, sp. nov., Kahlerosphaera isopleura Bragin, sp. nov., Stauracanthocircus folium Bragin, sp. nov., Tetraspongodiscus uncatus Bragin, sp. nov., Poulpus costatus (Kozur et Mostler), Eonapora robusta Kozur et Mostler, Planispinocyrtis kotelnyensis Bragin, sp. nov., Annulotriassocampe baldii (Kozur), and Pseudoeucyrtis annosus Bragin, sp. nov.; (2) a Middle-Late Carnian assemblage with Pseudostylosphaera glabella Bragin, sp. nov., P. gracilis Kozur et Mock, P. voluta Bragin, sp. nov., P. gelida Bragin, sp. nov., Kahlerosphaera unca Bragin, sp. nov., K. aspinosa Kozur et Mock, K. fuscinula Bragin, sp. nov., K. acris Bragin, sp. nov., Capnuchosphaera kuzmichevi Bragin, sp. nov., C. triassica De Wever, C. angusta Bragin, sp. nov., Sarla intorta Bragin, sp. nov., S. compressa Bragin, sp. nov., S. aequipeda Bragin, sp. nov., Betraccium irregulare Bragin, B. kotelnyensis Bragin, sp. nov., Spongotortilispinus carnicus Kozur et Mostler, S. subtilis Bragin, sp. nov., Dumitricasphaera simplex Tekin, D. aberrata Bragin, sp. nov., D. arbustiva Bragin, sp. nov., Zhamojdasphaera epipeda Bragin, sp. nov., Z. proceruspinosa Kozur et Mostler, Vinassaspongus subsphaericus Kozur et Mostler, Palaeosaturnalis triassicus Kozur et Mostler, Paronaella concreta Bragin, sp. nov., P. aquilonia Bragin, sp. nov., Tetraspongodiscus cincinnalis Bragin, sp. nov., Annulotriassocampe baldii (Kozur), Canoptum zetangense Wang et Yang, Whalenella speciosa (Blome), Syringocapsa turgida Blome, Droltus gelidus Bragin, sp. nov., and D. niveus Bragin, sp. nov.; (3) an Early Norian assemblage with Pseudostylosphaera glabella Bragin, sp. nov., P. gelida Bragin, sp. nov., P. voluta Bragin, sp. nov., Kahlerosphaera retunsa Bragin, sp. nov., Capnuchosphaera deweveri Kozur et Mostler, Sarla globosa Bragin, sp. nov., Palaeosaturnalis mocki Kozur et Mostler, Paronaella aquilonia Bragin, sp. nov., Syringocapsa turgida Blome, and Droltus gelidus Bragin, sp. nov.; (4) a Middle Norian assemblage with Sarla globosa Bragin, sp. nov., Sarla sp., Syringocapsa turgida Blome, Canoptum sp., and Laxtorum? sp. The fact that these assemblages include taxa that are recorded more southerly, including paleotropical localities, and support dating based on mollusks, enables the use of these data for Boreal-Tethyan correlation. The greatest similarity to Tethyan associations is observed in the Late Carnian and Early Norian, that is, the periods of the greatest penetration of thermophilic mollusks into this basin. A total of 69 radiolarian species of 3 orders, 18 families, and 29 genera are described; 44 species are newly described; the stratigraphic and geographical ranges of the majority of taxa are improved considerably.     

Norigondolella n. gen., eine neue obertriassische Conodontengattung

The development of the genus concept for platform-bearing gondolellid conodonts is discussed. The collective genusNeogondolella Bender & Stoppel s. l. must be subdivided into several genera separated mostly by morphological differences of their platform elements. Only for these genera, but not forNeogondolella s. l., differential diagnoses againstGondolella Stauffer & Plummer s. l. can be given.Norigondolella n. gen., species of which were formerly placed intoParagondolella Mosher s. l.,Gondolella Stauffer & Plummer s. l. orNeogondolella Bender & Stoppel s. l., is a typical gondolellid platform conodont genus of topmost Triassic (Norian-Rhaetian) pelagic sediments. It has evolved from Carnian representatives ofParagondolella, but the lower surface of the platform element shows considerable similarities (homoeomorphy) to the Middle Carboniferous to Middle Permian genusMesogondolella Kozur.     

On the status of the Permian radiolarian genus Multisphaera Nazarov and Afanasieva, 2000

Based on the detailed morphological study of the type species of the genus Multisphaera Nazarov and Afanasieva, the author emends the status and diagnosis of this genus and of the subfamily Multisphaerinae Nazarov and Afanasieva. He found that the initial spicule of the type species is not multi-rayed, as initially thought, but a tetrahedral spicule. Accordingly, the genus is assigned to the former tribe Tetrentactiniini Kozur and Mostler that is herein raised to the family rank and comprises all Upper Devonian to lower Permian genera with a tetrahedral initial spicule. Consequently, the subfamily Multisphaerinae is considered a junior synonym of the family Tetrentactiniidae. To better solve the taxonomic problems of this family the author compares the morphological diversity of its members with the morphological diversity of the lower Tithonian (uppermost Jurassic) tetrahedral spicule-bearing entactinarian Radiolaria occurring in the Solnhofen area, southern Germany. Although these genera have the same type of spicule as the Paleozoic ones, they seem to belong to a new family because between the lower Tithonian and the lower Permian genera there is a long time interval of about 120 Ma in which no radiolarians with a tetrahedral initial spicule have been recorded so far. This comparison shows once more the conservative character of the initial spicule and the wide morphological diversity of the extraspicular shell.     

Trimeridianellidae nov. fam., a Triassic spumellarian radiolarian family with simple initial skeleton and pyloniacean mode of growth

Based on several samples of Middle and Late Triassic age, the authors describe a new family of spumellarian Radiolaria – Trimeridianellidae Dumitrica and Tekin, characterized by a three-fold symmetry, a first shell of triangular prism type with 6 primary rays originated at the edges of two bases of the prism, or 3 meridian arches with 3 equatorial rays, and growth of pyloniacean type. The family comprises 4 monospecific new genera (Enneacladus Dumitrica, Endoprisma Dumitrica, Trimeridianella Dumitrica and Tekin, and Tristylopyle Dumitrica), 4 new species and one left in open nomenclature. In spite of their spotty occurrences and low diversity, the authors attempt to trace the evolution of the family, which is considered to be the result of heterochrony in ontogenetic growth, mostly of paedomorphic type. The family seems to be related to the Middle Triassic family Patruliidae Dumitrica with which it shares in common the simple microsphere with 6 primary rays and the pyloniacean mode of growth.     

New fossil insects from the Anisian (Lower to Middle Muschelkalk) from the Central European Basin (Germany and The Netherlands)

The Palaeozoic–Mesozoic transition is characterized not only by the most massive Phanerozoic mass extinction at the end of the Permian period, but also its extensive aftermath and a prolonged period of major biotal recovery during the succeeding Middle to Late Triassic. Particularly, Anisian insect species from units of the Lower to Middle Muschelkalk from the Central European Basin are rare. The specimens described here originated from the Anisian Wellenkalk facies (Lower Muschelkalk), Vossenveld Formation of the Winterswijk quarry, The Netherlands, and from the orbicularis Member (lowermost Middle Muschelkalk, Anisian) of Esperstedt near Querfurt (Saxony-Anhalt). Thus, the described insect remains from Winterwijk and Esperstedt expand our knowledge about Middle Triassic terrestrial arthropod communities and their palaeodiversity. A new species of Chauliodites (C. esperstedti sp. nov) is introduced.     

广西南部二叠纪长兴期放射虫动物群

广西南宁南部柳桥区的长兴组上部硅质岩中存着精美的长兴期晚期放射虫动物群,共有24属39例,其中3个新种,其中以Latentifistularia最为常见,其次是Albaillllidae和Entactiniidae,属于二叠纪晚期的Neoalbaillella optima组合。半生生物有有孔虫、海绵骨针、腕足动物等。多数属在接近二叠纪－三叠世界线附近消失,但生物群中已出现少量三叠纪分子。     

Pleuromeia discovered from the Middle Triassic Linjia Formation of Benxi,Northeast China

A new species Pleuromeia shaolinii Zhang et Wang n. sp. was established under the family Pleuromeiaceae, based on eight specimens of stems and leaves, cones, rhizophores, and adventitious roots that were recently collected from the Middle Triassic Linjia Formation in Benxi, Northeast China. P. shaolinii n. sp. is similar to the type species P. sternbergii, in the morphology of the stem and rhizophore, but differs from it by the transversely rounded sporophylls organized in a heterosporous cone, and the two conspicuous triangular scars present on both sides of the vascular bundle scar. The Linjia flora, an only reliable Middle Triassic flora in Northeast China, was discovered 40 years ago. Although the Linjia Formation can be compared with the Middle Triassic Ermaying Formation from another area of the North China plate, its age was questionable due to the absence of the Early to Middle Triassic index fossil Pleuromeia. Based on the flora similarity, the zircon dating, lithologic characteristics, presence of a disconformity, and the occurrence of the Middle Triassic–Early Cretaceous insect fossil Sogdoblatta, the age of the Linjia Formation can be confirmed as Middle Triassic. The new species not only supplements our knowledge in biology and taxonomy of Pleuromeia, but also supports the Middle Triassic age assignment of the Linjia flora.     

Cancrinella and Costatumulus (Brachiopoda) from the Permian of South Mongolia and South China: Their morphology,biostratigraphy and distribution

Two brachiopod genera, Cancrinella Fredericks and Costatumulus Waterhouse, are studied from the Permian of South Mongolia and South China. Many of the specimens previously described as Cancrinella are revised and assigned to Costatumulus. The new data presented in this paper indicate that these two genera are similar to each other, but are distinguishable by different features in relation to their morphology. Several species of both genera are described here, providing critical new information on the morphology and taxonomy of Cancrinella and Costatumulus. The reported biostratigraphical information reveals that species of Cancrinella are present from the Lower Carboniferous (Tournaisian) to Middle Permian (Capitanian), whereas species of Costatumulus are mostly restricted to the Lower Permian (Sakmarian) to Upper Permian (Changhsingian). An analysis of the palaeogeographical distribution of Cancrinella and Costatumulus reveals that Cancrinella was more commonly distributed in the Northern Transitional Zone and the Boreal Realm, in contrast to Costatumulus, which tended to be more common in the Southern Transitional Zone and Gondwanan Realm.     

Taxonomic study of spongy spumellarian Radiolaria with three and four coplanar spines or arms from the middle Carnian (Late Triassic) of the Köseyahya nappe (Elbistan, SE Turkey) and other Triassic localities

The present article is a taxonomic study of all spongy spumellarian radiolarian taxa with three and four coplanar spines or spongy arms occurring in the middle Carnian from the Köseyahya section, near the town of Elbistan, SE Turkey. This fauna is characteristic of the Tetraporobrachia haeckeli radiolarian Zone, and comes from an 8 m thick succession of clayey-cherty limestones occurring at the lower part of the section. In addition, a few species from the Middle and Upper Triassic from other areas have been also included in this study to improve some generic diagnoses, and to better comprehend the diversity and evolutionary trends of some genera, subfamilies and families. The taxonomy at the generic and suprageneric levels is based primarily on the types of microsphere. This new approach allowed new taxonomic arrangements of genera and suprageneric units, and suggested new phylogenetic relationships among these radiolarians and between them and younger radiolarians. The authors discuss and describe 69 species, of which 37 are new, and 14 genera, of which three are new (Paraparonaella, Pseudangulobracchia, and Ropanaella). The genus Triassoastrum and others are reinterpreted. All genera studied are assigned to five subfamilies, of which two are new (Tetrapaurinellinae and Triassocrucellinae), and two families (Tritrabidae and Veghicycliidae). Nine species in open nomenclature are also illustrated.     

Permian and Triassic radiolarians from chert breccia in the Nong Prue area,western Thailand: its origin and depositional setting in the Paleotethys

Bedded chert and siliceous shale successions previously regarded as the Silurian–Devonian rock units, distributed in the Nong Prue area, northwest of Kanchanaburi, western Thailand, yielded Lopingian (upper Permian) and Lower–Middle Triassic radiolarians. We found chert breccia layers in northern Nong Prue area, mainly consisting of angular to sub-angular chert clasts with matrices of silt-sized chert grains and clay minerals. We discriminated uppermost Pennsylvanian–Lopingian (upper Carboniferous–upper Permian) and Middle Triassic radiolarian-bearing chert clasts from four different levels of the chert breccia; 28 species of 15 genera with one radiolarian gen. et sp. indet. are identified. On the basis of sedimentary characteristics of the chert breccia, we suggest that the chert breccia is of sedimentary origin. The radiolarian assemblages reported here, together with previously known lithological and paleontological evidence, further indicate that the chert breccia was deposited in the Paleotethys with chert clasts derived from fine grained siliceous rocks in the continental margin to deep ocean basin of the Sibumasu Terrane.     

西藏申扎县木纠错地区晚三叠世海参骨片化石的发现及其地质意义

本文描述的海参骨片化石采自西藏申扎县木纠错西的多布日一带,主要有Theelia planorbicula Mostler,Theelia uariabilis alouakensis Kzour et Mock等晚三叠世诺利期标准分子。充分显示了海参骨片化石的重要的生物地层意义。该动物群的发现首次确定了西藏自治区北部冈底斯一腾冲地层区永珠蛇绿岩带以南晚三叠世海相地层的存在,并为中特提斯海的演化历史,古地理轮廓的确定,以及青藏高原的形成演变历史提供了新的资料。     

Middle Permian Ostracods from Tak Fa Limestone, Phetchabun Province, Central Thailand

This paper presents the first Permian ostracod fauna discovered in Thailand. The ostracods are recovered from the Tak Fa Limestone (Middle Permian) in Phetchabun province, central Thailand. The ostracods belong to eight genera and 15 species. Four species are newly described: Sargentina phetchabunensis nov. sp., Geffenina bungsamphanensis nov. sp., Reviya subsompongensis nov. sp. and Bairdia takfaensis nov. sp. The ostracod assemblages characterize a shallow marine, near shore environment at the time of deposition. Except for one species, which shows palaeobiogeographical links between Central Thailand and South China, all the other species are endemic.     

High organic carbon content and a decrease in radiolarians at the end of the Permian in a newly discovered continuous pelagic section: A coincidence?

The greatest mass extinction occurred at the end of the Permian. Most records of the mass extinction are not from pelagic sediments, but from shallow-marine and terrestrial sediments. Although several pelagic sections that span the end-Permian mass extinction have been found, these sections contain few index fossils and are often discontinuous because of small faults. We found the index fossils Albaillella cf. triangularis (Radiolaria) in siliceous claystone beds, Hindeodus parvus (Conodont) in the overlying black claystone beds, and Neospathodus cf. cristagalli and Ns. waageni (Conodont) in the subsequent siliceous claystone beds in Akkamori section-2 in northern Japan. These fossils suggest that this section ranges from the late Permian to the Early Triassic, including the early Induan and Olenekian stages. Furthermore, the lithological changes in the section, i.e., starting from bedded chert through siliceous claystone and black claystone to siliceous claystone, are concordant with those of well-known Permian–Triassic pelagic sequences in Japan. There is no gap between each lithofacie of the Akkamori section-2. Critical lithological continuity between Upper Permian siliceous claystone beds and uppermost Permian to lowermost Triassic black claystone beds of the Akkamori section-2 was recognized by observing hand-polished specimens and thin sections. Such paleontological and sedimentological evidence implies that the Akkamori section-2 is a continuous pelagic section that records the end-Permian mass extinction event. The carbonaceous black claystone beds have high total organic carbon (TOC) concentrations (1.06–3.31 wt.%), suggesting oceanic anoxia at least deep and probably stable primary productivity. A decrease in radiolarian abundance from 26–563 to 0.27–20 specimens/cm² coincided with an increase in TOC content from 0.01–0.16 to 1.06–3.31 wt.% at the boundary of the siliceous claystone and the overlying black claystone beds near the top of the Permian, implying that radiolarian extinction occurred at the end of the Permian coinciding with oceanic anoxia. Although TOC contents decreased in the early Olenekian (Smithian), radiolarian abundance did not increase at that time, indicating that radiolarian recovery was delayed by > 1.5 m.y.     

An unusually diverse mollusc fauna from the earliest Triassic of South China and its implications for benthic recovery after the end-Permian biotic crisis

A single carbonate coquinoid lens from the Griesbachian (Early Triassic) of Shanggan, South China, yielded 11 bivalve species described in this study in addition to four gastropod and one ammonoid species reported elsewhere. This makes the Shanggan fauna one of the richest mollusc faunas from the early post-extinction interval after the end-Permian mass extinction event. Four of the present genera are long-term survivors, five are holdovers that went extinct at the end of the Griesbachian or later in the Early Triassic, and seven first appear in the Griesbachian. Three new bivalve species are described: Myalinella newelli nov. sp., Scythentolium scutigerulus nov. sp., and Eumorphotis shajingengi nov. sp. The genus Astartella, previously assumed to have vanished at the end of the Permian, is reported for the first time from the Early Triassic, which also removes Astartidae from Early Triassic Lazarus taxa. The small growth size of the Astartella specimens supports an earlier hypothesis that many of the Early Triassic Lazarus taxa did not survive in unknown refuges but were simply overlooked due to the scarcity of easily observable large-sized specimens. Ecologically, a comparatively high proportion of infaunal bivalve species (4/11) is remarkable for the early post-extinction interval, supporting the impression of a relatively advanced recovery state. Moreover, abundance-data of the bivalve-gastropod community reveal a remarkably low dominance index (D = 0.17) that is suggestive for advanced recovery and stable environmental conditions. It is proposed that the Shanggan fauna represents a late Griesbachian benthic recovery event that coincided with the appearance of similarly diverse benthic faunas in Oman and Primorye. A high proportion of genera that have previously not been reported from the Early Triassic indicate that the prevalence of poor preservation conditions is a major obstacle in identifying early phases of recovery from the greatest crisis in the history of metazoan life. The early recovery of benthic faunas reported in this study questions previous claims of a prolonged lag phase as a consequence of the extraordinary extinction magnitude or the persistence of adverse environmental conditions.     

A Remarkable New Family of Jurassic Insects (Neuroptera) with Primitive Wing Venation and Its Phylogenetic Position in Neuropterida

Background

Lacewings (insect order Neuroptera), known in the fossil record since the Early Permian, were most diverse in the Mesozoic. A dramatic variety of forms ranged in that time from large butterfly-like Kalligrammatidae to minute two-winged Dipteromantispidae.

Principal Findings

We describe the intriguing new neuropteran family Parakseneuridae fam. nov. with three new genera and 15 new species from the Middle Jurassic of Daohugou (Inner Mongolia, China) and the Early/Middle Jurassic of Sai-Sagul (Kyrgyzstan): Parakseneura undula gen. et sp. nov., P. albomacula gen. et sp. nov., P. curvivenis gen. et sp. nov., P. nigromacula gen. et sp. nov., P. nigrolinea gen. et sp. nov., P. albadelta gen. et sp. nov., P. cavomaculata gen. et sp. nov., P. inflata gen. et sp. nov., P. metallica gen. et sp. nov., P. emarginata gen. et sp. nov., P. directa gen. et sp. nov., Pseudorapisma jurassicum gen. et sp. nov., P. angustipenne gen. et sp. nov., P. maculatum gen. et sp. nov. (Daohugou); Shuraboneura ovata gen. et sp. nov. (Sai-Sagul). The family comprises large neuropterans with most primitive wing venation in the order indicated by the presence of ScA and AA1+2, and the dichotomous branching of MP, CuA, CuP, AA3+4, AP1+2. The phylogenetic position of Parakseneuridae was investigated using a phylogenetic analysis of morphological scoring for 33 families of extinct and extant Neuropterida combined with DNA sequence data for representatives of all extant families. Parakseneuridae were recovered in a clade with Osmylopsychopidae, Prohemerobiidae, and Ithonidae.

Conclusions/Significance

The presence of the presumed AA1+2 in wings of Parakseneuridae is a unique plesiomorphic condition hitherto unknown in Neuropterida, the clade comprising Neuroptera, Megaloptera, Raphidioptera. The relative uncertainty of phylogenetic position of Parakseneuridae and the majority of other families of Neuroptera reflects deficient paleontological data, especially from critical important periods for the order, earliest Triassic and latest Triassic/earliest Jurassic.