Drei neue Arten desFestuca ovina-Formenkreises (Poaceae) aus dem Osten Österreichs

WithinFestuca ovina agg. two hexaploid taxa:F. brevipila andF. carnuntina, and one tetraploid:F. eggleri, are described as new species.F. pseudovina Hack. exWiesb. is better treated asF. valesiaca Schleich. exGaudin subsp.parviflora (Hack.), comb. nova.

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Drei neue Arten desFestuca ovina-Formenkreises (Poaceae) aus dem Osten Österreichs

     

Molecular (RAPD) analysis on Italian taxa of theOphrys bertolonii aggregate (Orchidaceae)

The RAPD technique has been used for analyzing the genetic material of seven Italian populations of theOphrys bertolonii aggregate (Orchidaceae). For comparison the related speciesO. sphegodes andO. fuciflora have also been analyzed. A high genetic variability has been observed within the same populations, and on the whole onlyO. bertolonii can be safely differentiated from the other taxa of the aggregate, which are undistinguishable with this method.     

Festuca curvula (Poaceae) — neu für Österreich

The octoploidFestuca curvula Gaudin (F. ovina agg.) is new for the Austrian flora. Epidermal structures indicate relationships toF. pallens. Its distribution ranges from the Pyrenees and French Massif Central through the Alps to western Austria.

     

Hybriden und hybridogene Sippen ausOphrys bertolonii undO. atrata (Orchidaceae)

Flower polymorphism in 15 ItalianOphrys populations (N = 282) was analysed in detail. Statistical and graphical evaluation of data demonstrates morphological relationships and amount of hybrid character coherence.O. bertolonii s. str. andO. atrata are stable parental taxa, but occasionally form hybrids and hybrid swarms. Much evidence suggests that it was through such homogamic hybridization, involving these and other species ofOphrys, that the stabilized taxaO. bertoloniiformis andO. promontorii (ecologically different, but sympatric with the parents on Gargano), andO. benacensis (geographically separated from the parents in the Insubric region) originated. These differentiation-hybridization cycles are linked to the break-down and new establishment of incomplete prezygotic crossing barriers (e.g. different flower biology, flowering time, habitat preference).

     

Populationsstatistische Analysen bestätigen die Heterogeneität vonOphrys „arachnitiformis“ (Orchidaceae)

Statistical population analyses have been carried out on so-calledOphrys arachnitiformis, partly sympatricO. sphecodes s. l. as well as on other species ofOphrys (O. holosericea, O. exaltata, O. apulica, O. cornuta, O. biscutella, O. bertolonii, O. bertoloniiformis.). Altogether 63 populations with 521 individuals were studied in regard to 54 morphological parameters. — It can be shown thatO. arachnitiformis is an extremely heterogeneous assemblage which includes taxa of most diverse origin, different degrees of stability and nomenclatorial rank. Only common trait is their very diverse relationships withO. specodes s.l.—The basionymO. arachnitiformis Gren. & Phil. refers to a colour variant ofO. sphecodes Mill. subsp.sphecodes only, and is not applicable to other segments of the assemblage.A late flowering, morphologically and biologically well isolated group of populations from southern France without any perceptible hybrid influence is established as a new species:Ophrys splendida Gölz & Reinhard. Populations from north and central Italy with distinct characters but hybrid influences fromO. holosericea/O. exaltata are calledO. tyrrhena Gölz & Reinhard.O. morisii, an endemic to Sardinia, represents an independent species of still obscure evolution. The hybrid interpretation ofO. catalaunica is statistically corroborated, but somewhat modified. For the remaining segments ofO. arachnitiformis investigated (from Castilia, Dalmatia, and the Monte Gargano) no formal taxonomic proposals are made, but some suggestions about their genesis are possible.

     

Labellum micromorphology in theOphrys bertolonii agg. and some related taxa (Orchidaceae)

Micromorphology and distribution of trichomes in six taxa of theOphrys bertolonii aggregate (O. bertolonii, O. bertoloniiformis, O. benacensis, O. aurelia, O. drumana, O. catalaunica) are described. A fairly good homogeneity between all the taxa, with a moderate degree of variability even within the same taxon, has been observed. Comparison of these characters with those ofO. promontorii andO. garganica corroborates the inclusion of the former species into theO. sphegodes aggregate. Trichome analysis of a hybrid is exemplified byO. ×chiesesica (O. drumana ×O. fuciflora).     

Neue Beobachtungen zur Bestäubung vonOphrys (Orchidaceae) in Südspanien,mit besonderer Berücksichtigung des FormenkreisesOphrys fusca agg.

An investigation of pseudocopulation behaviour in species ofOphrys from southern Spain confirms the close relationship betweenCampsoscolia ciliata (Scoliidae) andOphrys speculum, and betweenEucera nigrilabris (Apoidea) andOphrys tenthredinifera. It could be demonstrated thatEucera barbiventris is the pollinator ofOphrys scolopax subsp.scolopax, whereas other species ofEucera andTetralonia which are active at the same time show no interest at all for the flowers of this species. Special attention was paid to the forms of the taxonomically confusedOphrys fusca group:O. fusca s. str.,O. iricolor, O. omegaifera andO. atlantica. WhileO. fusca s. str. is widespread, small-flowered and has late anthesis,O. iricolor has very large flowers and early anthesis. Each of the four members ofO. fusca agg. in S. Spain is pollinated by a different bee, and selective experiments show that each of these four species of bees is specifically attracted only to one of theOphrys species:Andrena flavipes is the pollinator ofO. fusca s. str.,Colletes cunicularius infuscatus ofO. iricolor, Anthophora atroalba ofO. omegaifera, andChalicodoma parietina ofO. atlantica. These four pollinators belong to 4 different bee families (Andrenidae, Colletidae, Anthophoridae, Megachilidae). As this type of pollination represents a very effective pregamic isolation mechanism, these four taxa ofOphrys fusca agg., at least in southern Spain, behave and should be regarded as genuine species.

     

Orchis prisca (Orchidaceae), eine neue Art aus Kreta

Orchis prisca Hautzinger is described as a new diploid species from Crete, related toO. patens andO. spitzelii.

     

Neue Taxa vonTillandsia subgenusDiaphoranthema (Bromeliaceae) aus Bolivien und Argentinien

5 new taxa are described and illustrated; their position within the subgenus is discussed:T. hirta W. Till & L. Hromadnik,T. cotagaitensis L. Hromadnik,T. caliginosa W. Till, andT. gilliesii Baker subsp.polysticha W. Till & L. Hromadnik are members of a group includingT. myosura Grisebach exBaker,T. mandonii E. Morren exMez in DC.,T. retorta Grisebach exBaker em.Grisebach andT. andicola Gillies exBaker. T. brealitoensis L. Hromadnik is related toT. angulosa Mez in DC. but distinct and possibly of hybrid origin.

     

Systematik,Verbreitung und Karyologie derFestuca violacea-Gruppe (Poaceae) im Ostalpenraum

The biosystematics of theFestuca violacea group (F. rubra subsp.violacea sensuHackel) in the Eastern Alps is studied.F. picturata Pils (=F. picta Kit. exSchultes nonJ. F. Gmelin) is confirmed as diploid, whereas the chromosome numbers forF. nitida (diploid),F. puccinellii (hexaploid), andF. norica (diploid, tetraploid and hexaploid) are reported for the first time. Details of morphology, leaf anatomy, and epidermal structures now allow a better separation of these species; new maps illustrate their distribution. Karyological, chorological, morphological, and anatomical data form the basis for a discussion of the phylogeny of theF. violacea group and its position withinF. rubra s. latiss.

     

Über zwei diploide,in Südwest-Anatolien endemische Arten aus dem Komplex derVeronica cymbalaria (Scrophulariaceae)

Veronica lycica Lehm. is a distinct, diploid member of theVeronica cymbalaria group, endemic in Lycia (S.W. Anatolia). Closely related is the newV. stamatiadae M. Fischer etW. Greuter which is also diploid and seems to be restricted to the small Greek island Ro close to the South coast of Lycia. The chromosome numbers for both species are reported for the first time (2n = 18).

Untersuchungen über den PolyploidkomplexVeronica cymbalaria agg., II. — Der erste Beitrag dieser Serie:Fischer (1975).     

A taxonomical revision ofThymus sect.Teucrioides (Lamiaceae)

TheThymus teucrioides Boiss. & Spruner aggregate is revised and the following new taxa, all from the alpine zone in the Greek mountains, are described:Th. leucospermus Hartvig from the calcareous mountains of Pindhos and Mt Parnassos in Sterea Ellas,Th. rechingeri Hartvig with the subsp.macrocalyx Hartvig from calcareous mountains in Sterea Ellas and N Peloponnissos, andTh. teucrioides subsp.alpinus Hartvig from the serpentine areas of N Pindhos. In the variableTh. teucrioides s. str. many characters have turned out to be markedly geographically correlated and many local populations can be distinguished by a particular combination of characters.Dedicated to Prof.K. H. Rechinger on the occasion of his 80th birthday.     

Ophrys helenae (Orchidaceae), eine lange verkannte Sippe der Balkanhalbinsel

Population analyses, scatter diagrams and field observations demonstrate thatOphrys helenae is morphologically (and ecologically) well separated from the partly sympatricO. mammosa andO. ferrumequinum in N. Greece. Hybrids with the distantly relatedO. mammosa are very rare.

     

Stachys beckeana (Labiatae) in Albanien und Jugoslawien

Morphological and cytological investigations were carried out inStachys beckeana Dörfler & Hayek andS. recta L. subsp.sarajevensis sensuHayek. These two taxa have the same morphological characteristics (shape and indumentum of leaves; morphology of sepals, petals and fruits) and both have 2n = 2x = 34 chromosomes. They cannot be considered as different taxa;S. beckeana Dörfler & Hayek is the valid name (synonyms:S. recta L. subsp.sarajevensis sensuHayek,S. hayekii Malý,S. recta L. subsp.hayekii Malý).

Erster Beitrag zur Neubearbeitung der Artengruppe derStachys recta L.     

Contributions of numerical taxonomy toOrnithogalum subg.Beryllis (Hyacinthaceae) in Morocco

In North Africa,Ornithogalum L. subg.Beryllis Baker is represented by three taxa (O. narbonense L.,O. pyrenaicum L., andO. sessiliflorum Desf.), often difficult to identify. 90 individuals from 4 localities were subjected to a biometric study based on 19 morphological characteristics. Several types of multivariate analyses allow a perfect identification of the three species. Studies on the reproductive biology, cytotaxonomy, and ecology supplement these data.     

Giemsa karyotypes and their evolutionary significance inScilla bifolia,S. drunensis,andS. vindobonensis (Liliaceae)

Quantitatively evaluated C-banded and conventional karyograms are presented for the related speciesScilla bifolia subsp.danubialis Speta,S. drunensis (Speta)Speta, andS. vindobonensis Speta. On the basis of banding patterns and karyotype structureS. bifolia subsp.danubialis (2n = 18, 2×) andS. drunensis (2n = 36, 4×) are quite similar, whileS. vindobonensis (2n = 18, 2×) is entirely different. There is a moderate degree of karyotypic variation withinS. bifolia subsp.danubialis andS. drunensis. However, inS. vindobonensis karyotypes and banding patterns are almost completely stable over a geographical range of about 500 km. The present results confirm the recent taxonomic separation ofS. vindobonensis fromS. bifolia, and suggest a considerable phylogenetic distance between these two diploid species. The results are discussed with reference to the morphological characters of the species and their geographical distribution.Evolution ofScilla and Related Genera, II.Dedicated to Univ.-Prof. Dr.Elisabeth Tschermak-Woess on the occasion of her 60th birthday.     

Veronica allahuekberensis (Scrophulariaceae), eine neue Art aus der Türkei

Veronica allahuekberensis A. Öztürk, spec. nova, probably endemic to the Allahuekber mountain in N.E. Anatolia, belongs toV. orientalis group of sectionVeronica and resemblesV. cuneifolia D. Don subsp.cuneifolia, endemic to S.W. Anatolia.

     

Tillandsia marconae — eine neue Art aus der peruanischen Küstenwüste

T. marconae is described as a new species from the Peruvian coastal desert. Its systematic position betweenT. paleacea Presl em.L. B. Smith andT. purpurea Ruiz & Pavon is discussed.

     

Zwei neue Arten der GattungSaponaria (Caryophyllaceae) aus Afghanistan und Pakistan

Saponaria stenopetala sp.n. in Eastern Afghanistan is close toS. pachyphylla Rech. f. andS. subrosularis Rech. f.—The nearest allies ofS. makranica sp.n. from Western Pakistan and Southeastern Iran areS. kermanensis Bornm. andS. floribunda (Kar. & Kir.)Boiss.

Flora Iranicae praecursores 36–37. — Praecursores praecurrentes in Pl. Syst. Evol.139, 313–317 (1982).     

Fünf neue Arten der GattungAnthemis (Compositae) aus Iraq und Afghanistan

Five new species ofAnthemis sect.Anthemis from the Flora Iranica region are described:A. gillettii (subsect.Anthemis) from NW. Iraq and adjacent Iran is allied toA. damascena.—A. kurdica (subsect.Anthemis) also grows in Iraqi Kurdistan.—A. hamrinensis (subsect.Rascheyana; akin toA. plebeia) is distributed in the Jabal Hamrin region at the extreme outer margin of the Zagros chains.—A. kandaharica (subsect.Anthemis) andA. freitagii (subsect.Rascheyana) are distributed in Afghanistan.

Anschrift des Herausgebers: Hofrat Univ.-Prof. Dr.Karl Heinz Rechinger, Beckgasse 22, A-1130 Wien, Österreich.