Influence of habitat features and hunter behavior on white-tailed deer harvest

Sport hunting may help in controlling cervid populations over large areas. As with natural predators, several environmental factors can influence sport harvest. A better understanding of the environmental variables that limit the efficiency of sport hunting could provide guidelines for more efficient wildlife management using hunting. We studied white-tailed deer (Odocoileus virginianus) hunting on a high deer density island where hunting was the sole form of predation. Our objective was to study the behavior of sport hunters and determine the habitat characteristics (e.g., abundance of deer forage, visibility of the deer from the hunter's point of view, and accessibility of the territory to hunters) that are associated with a successful harvest. We collected movements and harvest site location data from 477 hunters equipped with handheld Global Positioning System (GPS) units. Harvest sites were visited and characterized, along with a paired random site, to determine the environmental conditions associated with a successful hunt. We also developed a model to predict the daily number of deer seen by hunters considering weather conditions, hunter characteristics (e.g., age, experience), and date of hunting. We used the mean number of deer seen per hunter per day as a relative index of local density in each hunted territory. At both the site and landscape scales, the combination of visibility and access had a positive effect on the distribution of harvested deer. Habitat types with less visual obstruction from vegetation enabled hunters to see more deer in a given day. At the site scale, harvested deer were located in areas with a lower density of access routes compared to areas where hunters travelled throughout the day. Using an innovative approach of studying hunter behavior with GPS technology, digital maps, and questionnaires, we highlighted the factors associated with hunter success. Our study suggests that habitat characteristics could be modified to increase harvest by improving accessibility and visibility near roads. Creating openings in mature and regenerating forest near access roads could make sport hunting a more efficient management tool, but the potential impact of increased forage availability in forest openings should not be overlooked. © 2012 The Wildlife Society.     

Big-game,small-game,and gamebird hunting in New Zealand: Hunting effort,harvest, and expenditure in 1988

Abstract

A postal survey of 8639 licensed firearm owners in 1989 indicated that an estimated 117 200 ± 6300 New Zealanders (3.5% of the total population) did some hunting in 1988. An estimated 33 100 former hunters did not hunt in 1988 but thought it likely that they would hunt again in future. The survey provided useful estimates of 1988 national totals for hunting effort (4.4 million hunter days), gross expenditure ($NZ100 million), and harvest (6.5 million animals).

Small-game hunting dominated, involving 81% of hunters, 59% of total hunting effort, and 86% of total numerical harvest: rabbits (Oryctolagus cuniculus) and possums (Trichosurus vulpecula) each comprised 40% of the national bag. Gamebird hunting involved 48% of hunters, 19% of effort, and 11% of total harvest: ducks comprised 73% of the gamebird harvest. Although big-game hunting attracted the fewest hunters (42%) it ranked second for hunting effort (21%). Big game formed 3% of the total numerical harvest: pigs (Sus scrofa), goats (Capra hircus), and red deer (Cervus elaphus scoticus) were the most commonly taken. Big-game were estimated to comprise 49% of total harvest biomass, followed by small-game (47%), and gamebirds (4%). Most deer (nearly 60%) were taken for recreation, with helicopter-based hunting accounting for only one-third the total deer harvest.

A quarter of those people hunting in 1988 hunted on five or fewer days that year, and a relatively small group of mainly professional hunters accounted for a disproportionately large share of the overall harvest Hunters reported spending an average of $851 each on hunting in 1988. Expenditure on big-game comprised 44% of the total, small-game 23%, and gamebirds 33%. Expenditure per animal harvested or per day hunted was greater for big-game animals (other than goats) than for gamebirds, which were more expensive to hunt than small-game. Total hunting effort was inversely related to the average expenditure per animal harvested, regardless of the type of game.     

Use of Bait to Increase Archery Deer Harvest in an Urban–Suburban Landscape

Abstract: Hunting is the primary tool for managing white-tailed deer (Odocoileus virginianus) populations. Effectiveness of hunting in suburban areas may be reduced due to limited hunter access to small properties, firearms-discharge laws, and public safety concerns. In Connecticut, USA, hunting over bait on private land was recently legalized to increase harvest opportunities. Our objective was to assess bow-hunter willingness to use bait and effects of bait type, hunter disturbance, time spent hunting, and property size on deer-harvest potential in a suburban landscape. We mailed a prebaiting survey in February 2002 and a postbaiting survey in February 2004 to the same group of hunters. Hunters using bait were more successful and harvested more deer than hunters using no bait. Hunters using bait on small properties observed similar numbers of deer within shooting range as hunters using bait on larger properties. Hunters using bait met their venison needs, whereas hunters using no bait did not meet their needs. Resource managers should implement strategies that increase hunter success when developing urban deer-management programs for communities.     

Duck hunter activity,success, and satisfaction on public hunting areas

Because hunting disturbance can influence local distribution of ducks and their availability to hunters, managers often limit access to hunting areas to improve hunting success and satisfaction. Few studies have quantified the effectiveness of public area access restrictions on duck hunter activity, harvest, or hunters' satisfaction with their hunting experience. We used a cross-over design over 6 consecutive hunting seasons (2008–2009 through 2013–2014) on State Wildlife Areas (SWAs) in northeastern Colorado, USA, to compare the effects of restricted hunting access regulations and regulations without these restrictions on duck hunter activity, harvest success, harvest levels, and satisfaction. We also considered effects of SWA types, duck abundance, temperature, precipitation, use of equipment by duck hunting parties, and, for hunter satisfaction, hunting success, hunting parties' satisfaction with ducks seen, habitat conditions, crowding from other hunters, and SWA regulations. The number of days when duck hunters had access to restricted properties was about half that on unrestricted properties, and unrestricted properties were used by about twice as many duck hunting parties, but the mean number of hunting parties per available hunting day and mean party size were similar under the 2 types of regulations. Most (56%) duck hunting parties did not bag any ducks; hunting success (harvest of ≥1 duck by a hunting party) was best explained by a model that included a regulation type × hunting season interaction, a SWA type × month interaction, hunter density the previous day, an index of hunter investment (number of decoys used and whether dogs and calls were used), and temperature. Successful hunting parties harvested 1.92 ± 1.60 (SD) ducks/hunter/day (range = 0.1–7.0); the best model predicting the number of ducks harvested per hunter in successful parties included a regulation type × hunting season interaction, a SWA type × month interaction, hunter density the previous day, an index of hunter investment, temperature, and precipitation. Overall satisfaction of duck hunting parties with a day's hunt averaged 3.62 ± 1.20 based on a rank scale of 1 (very unsatisfied) to 5 (very satisfied), and was best explained by a model that included hunting party success; hunter investment; temperature; and satisfaction with duck numbers, habitat conditions, hunting regulations, and crowding. While greatly reducing days available for hunting, access restrictions were associated with improved chances of hunting parties successfully harvesting ≥1 duck in 5 of 6 hunting seasons, and substantially greater numbers of ducks harvested by hunters in successful parties in 1 of 6 hunting seasons. Restrictions did not have a strong direct effect on hunting parties' satisfaction with a day's hunt. Uncontrolled factors, including weather and use of equipment by hunters, had important influences on hunter success, harvest, and satisfaction. Managers should carefully assess hunting activity, hunter expectations and desires, and hunting area characteristics when considering access restrictions on public hunting areas.     

Effects of Earn-a-Buck and Special Antlerless-Only Seasons on Wisconsin's Deer Harvests

ABSTRACT Effective management of wildlife populations often requires motivating hunters to harvest sufficient numbers of animals of prescribed sex and age classes to meet management goals. For cervids, it is convenient to design harvest regulations relative to presence (male) or absence (young and female) of antlers because harvest of females has a larger effect on population growth. We used regression techniques to evaluate effects of 2 supplemental hunting programs based on additional days of hunting opportunity and an additional incentive used to complement additional days on harvest of antlered and antlerless deer in Wisconsin, USA. Earn-a-buck regulations, an incentive-based program that requires hunters to register an antlerless deer before being authorized to harvest an antlered deer, were associated with an average increase of 2.04 deer/km² in antlerless harvest and a 0.60 deer/km² decrease in harvest of antlered deer. Providing more opportunity for hunting of antlerless deer in the form of 4- and 8-day supplemental firearm seasons was associated with 1.10 deer/km² and 1.32 deer/km² increases, respectively, in antlerless harvest with trivial (0.02 deer/km² and 0.09 deer/km²) decreases in harvests of antlered deer. Our analysis suggests that extra days of hunting opportunity coupled with the earn-a-buck incentive was 56–88% more effective at increasing antlerless harvest relative to additional days of hunting without the incentive. Use of the earn-a-buck incentive resulted in decreased harvest of antlered deer and was disliked by many hunters. Quantifying these relationships is important for helping managers predict the costs and benefits of various hunting programs.     

Evaluating Differences in Harvest Data Used in the Sex-Age-Kill Deer Population Model

Abstract: A steady increase in archery hunting participation and frequent changes in hunter regulations led to an evaluation of harvest data used in a common white-tailed deer (Odocoileus virginianus) population model. Our goal was to determine if model parameters and population estimates traditionally estimated solely by firearm harvest data were biased with respect to altered sex and age ratios brought about by increases in archery hunting and harvest success. The sex-age-kill (SAK) model, commonly used by state agencies, was developed in the mid-1900s when deer numbers were low and firearm harvest was predominant. Management actions were concentrated on increasing deer numbers, and model assumptions relied heavily on a stable age distribution and a minimal antlerless deer harvest. We evaluated the reliance of SAK in a modern hunting scenario using a 10-year dataset obtained from Michigan, USA, that encompassed a variety of climatic regions, hunting seasons, and regulation scenarios. We found that firearm and archery harvest sex and age ratios differed among 5 geographic groups and study years for males, females, and fawns (P<0.001, P = 0.001, and P = 0.037, respectively). Also, the addition of archery harvest data increased population estimates but did not alter overall trends. We recommend that managers reassess harvest-based population estimates in 2 situations: 1) if regulation changes affect antlerless deer harvest, and 2) when trends in hunter success rates cause fluctuations in harvest data.     

Spatial distribution and landscape associations of large-antlered deer

For more than a century, the Boone and Crockett Club has kept records of harvest locations for 38 categories of North American big game, which includes measurement details and sizes of enduring characteristics (e.g., antlers, skulls, horns). All white-tailed deer (Odocoileus virginianus) entered in the Club records (i.e., record deer) have large antlers because the measurements must meet or exceed a minimum size to qualify for entry. We used the records from the Club to test the hypothesis that the spatial distribution in the harvest of record deer was related to the number of antlered deer harvested, variations in land productivity, soil mineral content, and attributes of land cover for 9 states in the midwestern United States. We used a Bayesian theoretical framework to develop a spatial model that examined the influence of explanatory variables on the number of record deer harvested in each county. In our study area, 3,658 record deer were harvested in 692 of the 856 counties (80.8%). More record deer were harvested in counties that had more high-contrast edges, less contiguous land cover, and greater variation in soil productivity. These results provide information for managers and hunters to better understand the spatial distribution of record deer and factors that are correlated with their distribution. © 2019 The Wildlife Society.     

Interaction effects between weather and space use on harvesting effort and patterns in red deer

Most cervid populations in Europe and North America are managed through selective harvesting, often with age‐ and sex‐specific quotas, with a large influence on the population growth rate. Less well understood is how prevailing weather affects harvesting selectivity and off‐take indirectly through changes in individual animal and hunter behavior. The behavior and movement patterns of hunters and their prey are expected to be influenced by weather conditions. Furthermore, habitat characteristics like habitat openness are also known to affect movement patterns and harvesting vulnerability, but how much such processes affect harvest composition has not been quantified. We use harvest data from red deer (Cervus elaphus) to investigate how weather and habitat characteristics affect behavioral decisions of red deer and their hunters throughout the hunting season. More specifically, we look at how sex and age class, temperature, precipitation, moon phase, and day of week affect the probability of being harvested on farmland (open habitat), hunter effort, and the overall harvest numbers. Moon phase and day of week were the strongest predictors of hunter effort and harvest numbers, with higher effort during full moon and weekends, and higher numbers during full moon. In general, the effect of fall weather conditions and habitat characteristics on harvest effort and numbers varied through the season. Yearlings showed the highest variation in the probability of being harvested on farmland through the season, but there was no effect of sex. Our study is among the first to highlight that weather may affect harvesting patterns and off‐take indirectly through animal and hunter behavior, but the interaction effects of weather and space use on hunter behavior are complicated, and seem less important than hunter preference and quotas in determining hunter selection and harvest off‐take. The consideration of hunter behavior is therefore key when forming management rules for sustainable harvesting.     

Is hunting an effective tool to control overabundant deer? A test using an experimental approach

Overabundant populations of cervids have induced drastic negative effects on plant communities in several regions worldwide. Antlerless deer harvest by sport hunters has been proposed as a potential solution to overabundance because the philopatric behavior of female deer is expected to limit recolonization of hunted zones. The efficiency of this method, however, has rarely been tested in the wild. Using a large-scale experimental design, we reduced white-tailed deer (Odocoileus virginianus) density within 5 20-km² areas on Anticosti Island (Québec, Canada). Our objective was to harvest 50% of antlerless deer in each site during the first year of the study in 2002, and 30% from 2003 to 2006. We monitored deer density, vegetation abundance and growth as well as deer life-history traits during 6 years in these experimental sites and in 5 control sites where harvest rate was 5–7%. Overall, we achieved 93% of harvest objectives. Contrary to our expectations, however, deer density, vegetation abundance and growth, and deer life-history traits did not vary differently in experimental and control sites during the study period. They rather varied stochastically but synchronously. We discuss several alternative hypotheses that may explain these results, including 1) compensatory mechanisms, 2) biases in density estimates, 3) limited access to territory for hunters, 4) large target areas for localized management, 5) low hunter density, 6) recolonization by surrounding deer, 7) slow plant response under canopy cover, and 8) bottom-up mechanisms. Given the large efforts invested in this study, we conclude that the local control of abundant cervid populations through sport hunting may be difficult to achieve in many natural environments. © 2012 The Wildlife Society.     

Reduction and Maintenance of a White-Tailed Deer Herd in Central Massachusetts

Abstract: Controlled public hunts have been used in a variety of settings to reduce overabundant white-tailed deer (Odocoileus virginianus) herds. We present the results of a large-scale (160 km²) controlled hunt at Quabbin Reservation (QR) in central Massachusetts, USA. The QR was divided into 5 hunt zones. Hunting was initiated in each zone from 1991 to 1994 and continued through 2004. The management goal was to achieve posthunt deer densities of4 deer/km². Initial estimated deer densities in each zone ranged from 11.4 deer/km² to 27.6 deer/km². The management goals were reached in each zone after 2-4 years of hunting. Posthunt populations were maintained at or below the goal even though total hunter effort was reduced. Hunters were not required to harvest antlerless deer, but antlerless deer comprised 55-83% of the harvest each year. We simulated the effects of 5 years without hunting on deer populations. The simulated deer population exceeded management goals after 2 years. Our results demonstrate that controlled public deer hunts can effectively reduce deer populations and maintain them at desired levels over large areas with minimal hunter restrictions. Managers should prepare stakeholders during the hunt planning process for the need to continue overall harvest rates of >30% during the maintenance phase of a deer management program.     

Evaluating how hunters see and react to telemetry collars on white-tailed deer

Fates of individuals outfitted with radiotransmitters commonly are used for estimating survival rates in populations of large animals that are hunted. Despite precautions, this practice may be subject to complex biases associated with hunter reaction to presence of radiotransmitters. To assess this potential bias we conducted an experiment using artificial deer (i.e., decoys) to measure hunters' abilities to see deer and determine if deer seen were wearing radiocollars. We used logistic regression to quantify probabilities that seeing deer and subsequently seeing radiocollars might be influenced by distance, percent visual obstruction, body orientation, hunter experience, and antler characteristics of deer. Additionally, we evaluated how experience and antler characteristics of deer might influence a hunter's decision to harvest a radiocollared deer. We found that 25.8% of the potentially observable collared deer (n = 663) were subsequently observed by hunters. Odds of observing deer and radiocollars increased 95% and 230%, respectively, for each additional log(yr) of hunting experience. Willingness to harvest radiocollared deer increased 89% for each additional log(yr) of hunting experience and 144% for large-antlered deer relative to antlerless deer. When hunting is an important source of mortality, analysts need to understand how potential biases associated with observing deer are associated with hunters' reactions to and subsequent decisions to harvest radiocollared animals. Our study suggested that presence of radiocollars may influence a deer's potential risk of being harvested and in turn bias telemetry-based estimates of survival, given that hunting mortality is the largest component of total mortality in hunted deer populations. Collar-based telemetry is used nearly universally by wildlife managers and researchers throughout North America and elsewhere to estimate and monitor the survival of big game populations that are managed through hunting. Our findings demonstrate that these estimates are likely subject to complex and systematic biases that managers should consider when evaluating future population-level effects of managed hunting. © 2011 The Wildlife Society     

The role of ethics in the management of New Zealand's wild mammals

Abstract

Wild deer are used to illustrate the conflicts that arise when an animal is both a pest and a valued resource. At present there are c. 250 000 wild deer in New Zealand. These are pursued by c. 37 000 hunters who spend more than $20 million annually to harvest c. 70 000 deer. Nearly two-thirds of the harvest is taken by recreational hunters; the remainder forms the basis of a small industry for venison export. Despite their recreational and commercial value, wild deer continue to be a significant conservation pest. Conservationists generally would like wild deer to be eradicated, or, failing that, their densities held at the lowest levels possible with the control resources available. Hunters, however, would like deer to remain, and at higher rather than lower densities, to provide better prospects for hunting success. We discuss ways in which these apparently incompatible aspirations can be partially reconciled.     

Hunting Patterns,Ban on Baiting,and Harvest Demographics of Brown Bears in Sweden

Abstract We analyzed harvest data to describe hunting patterns and harvest demography of brown bears (Ursus arctos) killed in 3 geographic regions in Sweden during 1981–2004. In addition, we investigated the effects of a ban on baiting, instituted in 2001, and 2 major changes in the quota system: a switch to sex-specific quotas in 1992 and a return to total quotas in 1999. Brown bears (n=887) were harvested specifically by bear hunters and incidentally by moose (Alces alces) hunters. Both hunter categories harvested bears 1) using dogs (37%), 2) by still hunting (30%), 3) with the use of bait (18%), and 4) by stalking (16%). The proportion of bears killed with different harvest methods varied among regions and between bear- and moose-oriented hunters. We found differences between male (52%) and female bears (48%) with respect to the variables that explained age. Moose-oriented hunters using still hunting harvested the youngest male bears. Bears harvested during the first management period (1981–1991) were older and had greater odds of being male than during the subsequent period. It appears that hunters harvesting bears in Sweden are less selective than their North American counterparts, possibly due to differences in the hunting system. When comparing the 4 years immediately prior to the ban on baiting with the 4 years following the ban, we found no differences in average age of harvested bears, sex ratio, or proportion of bears killed with stalking, still hunting, and hunting with dogs, suggesting that the ban on baiting in Sweden had no immediate effect on patterns of brown bear harvest demography and remaining hunting methods. As the demographic and evolutionary side effects of selective harvesting receive growing attention, wildlife managers should be aware that differences in harvest systems between jurisdictions may cause qualitative and quantitative differences in harvest biases. (JOURNAL OF WILDLIFE MANAGEMENT 72(1):79–88; 2008)     

Spatio-temporal variation in male white-tailed deer harvest rates in Pennsylvania: Implications for estimating abundance

The performance of 2 popular methods that use age-at-harvest data to estimate abundance of white-tailed deer is contingent on assumptions about variation in estimates of subadult (1.5 yr old) and adult (≥2.5 yr old) male harvest rates. Auxiliary data (e.g., estimates of survival or harvest rates from radiocollared animals) can be used to relax some assumptions, but unless these population parameters exhibit limited temporal or spatial variation, these auxiliary data may not improve accuracy. Unfortunately maintaining sufficient sample sizes of radiocollared deer for parameter estimation in every wildlife management unit (WMU) is not feasible for most state agencies. We monitored the fates of 397 subadult and 225 adult male white-tailed deer across 4 WMUs from 2002 to 2008 using radio telemetry. We investigated spatial and temporal variation in harvest rates and investigated covariates related to the patterns observed. We found that most variation in harvest rates was explained spatially and that adult harvest rates (0.36–0.69) were more variable among study areas than subadult harvest rates (0.26–0.42). We found that hunter effort during the archery and firearms season best explained variation in harvest rates of adult males among WMUs, whereas hunter effort during only the firearms season best explained harvest rates for subadult males. From a population estimation perspective, it is advantageous that most variation was spatial and explained by a readily obtained covariate (hunter effort). However, harvest rates may vary if hunting regulations or hunter behavior change, requiring additional field studies to obtain accurate estimates of harvest rates. © 2011 The Wildlife Society.     

Hunters and hunting patterns in part of the Kaimanawa Recreational Hunting Area

Abstract

A postal survey of 156 hunters who used the three southeastern-most hunting blocks in the Kaimanawa Recreational Hunting Area (RHA) in 1986 and 1987 achieved a 74% response rate. Comparison of age distributions from this and earlier hunter surveys confirmed an apparent decline in recruitment of young hunters entering the sport during the 1980s. Most hunters had considerable experience (>10 years) and were motivated not only by hunting success but also by the aesthetic value of the outdoor experience. The prime motivation for hunting in the Kaimanawa RHA was the presence of sika deer, and more than two-thirds of the hunters came from outside the Tongariro/Taupo Conservancy. Hunting pressure over current sika deer range in the central North Island may decline if this species continues to disperse naturally or through illegal liberations to other areas. Most (80%) hunters felt that deer densities were acceptable and were satisfied with the present unrestricted hunting system. Fewer than half (41%) the respondents were in favour of active management in the Kaimanawa RHA. Suggestions for active management focused on increasing hunter safety and increasing hunting success rates. Deer densities, as indicated by faecal pellet counts, were inversely related to hunting pressure. Variation in hunting pressure between areas was caused mainly by differences in accessibility. This indicates some potential for manipulation of deer density through options such as increasing access to remote areas or restrictions on hunting pressure.     

Testing Global Positioning System Telemetry to Study Wolf Predation on Deer Fawns

ABSTRACT We conducted a pilot study to test the usefulness of Global Positioning System (GPS) collars for investigating wolf (Canis lupus) predation on white-tailed deer (Odocoileus virginianus) fawns. Using GPS collars with short location-attempt intervals on 5 wolves and 5 deer during summers 2002–2004 in northeastern Minnesota, USA, demonstrated how this approach could provide new insights into wolf hunting behavior of fawns. For example, a wolf traveled ≥1.5–3.0 km and spent 20–22 hours in the immediate vicinity of known fawn kill sites and ≥0.7 km and 8.3 hours at scavenging sites. Wolf travel paths indicated that wolves intentionally traveled into deer summer ranges, traveled ≥0.7–4.2 km in such ranges, and spent <1–22 hours per visit. Each pair of 3 GPS-collared wolf pack members were located together for ≤6% of potential locations. From GPS collar data, we estimated that each deer summer range in a pack territory containing 5 wolves ≥1 year old and hunting individually would be visited by a wolf on average every 3–5 days. This approach holds great potential for investigating summer hunting behavior of wolves in areas where direct observation is impractical or impossible.     

Habitat preferences of ptarmigan hunters in Norway

Hunting for willow ptarmigan (Lagopus lagopus) is a popular recreational activity in Norway, but studies of the hunters are limited. While large game management in Norway is based on sound models and research, management of small game hunters and hunting have not been subject to the same rigour. Compared to the extensive body of knowledge about ptarmigan behaviour and habitat preferences, the knowledge base on ptarmigan hunters is very limited. We surveyed the habitat preferences of 3,056 hunters to identify preferred landscape categories using pictures covering a range of landscape types. We also examined to which extent residence types and forms of hunting were related to habitat preferences. Through factor analysis, we identified three categories of hunting habitat; mountain forest, low alpine and high alpine. Mode of hunting is more important than residence status in terms of preferred habitat. Hunters using dogs preferred mountain forest and low alpine habitats more than hunters without dogs. Hunters without dogs had a higher preference than dog hunters for high alpine habitats. Hunters with mixed modes of hunting have wider habitat preferences. Residence status only affects the perception of the mountain forest habitat. The results may have implications for management as land use and place-based meanings are currently changing in many natural and rural landscapes, and hunting needs to be integrated with other recreational uses.     

Willow grouse bag size is more sensitive to variation in hunter effort than to variation in willow grouse density

Annual variation in harvested animals (hereafter bag size) is often used as an index of population abundance when investigating population dynamics. Few studies have evaluated how well bag size tracks population change despite its widespread use. Two recent studies on grouse harvest statistics have reached contrasting conclusions. Th ere is limited information about the functional response of hunters in relation to varying game densities, and effort is seldom recorded. We investigated how much of the variation in bag size (total number of harvested grouse km^?2) is explained by variation in willow grouse Lagopus lagopus density (adult and young grouse km^?2) and hunting eff ort (total number of hunting days km^?2). We also evaluated catch per unit effort (CPUE) as an index of grouse abundance, and estimated the response in harvest rate (total bag size in relation to total grouse density) to varying hunting effort. We used data from the 88 management areas on state land in Jämtland county, Sweden (1996–2007), where hunting days and bag size are recorded in detail. Willow grouse density was estimated in four of these management areas in August using line transects and distance sampling. The hunting effort and total grouse density explained most of the variation in bag size (R²= 0.89). Bag size was twice as sensitive to changes in hunting effort compared to changes in grouse density. More than a ten times change in the grouse population density was required to one unit change in bag size. The use of CPUE did not provide a better alternative index of grouse density, and variation in density only explained 23% of the variation in CPUE. Harvest rate showed a strong relationship with hunting effort, and we suggest that an upper limit in hunting effort can be used to reduce the risk of high harvest rates. Hunters became more efficient at low densities and controlling hunting effort is most important when there are indications of population lows and/or poor breeding. CPUE may be less sensitive to changes in game abundance than previously assumed, and bag size as a proxy for population density would then depend on the ability of hunters to adjust their effort according to population change. We speculate that this ability will depend on whether or not hunters have long‐term experience of a hunting area where they can return to hunt throughout the hunting season. We propose that recording hunting eff ort should be encouraged and possible correlations with game abundance and other factors such as weather should be investigated for game species.     

Habitat and weather influence body condition in white-tailed deer,Wisconsin, USA

In temperate and northern ecosystems where there are pronounced seasonal patterns in weather and available energy, there are corresponding patterns of body condition among white-tailed deer (Odocoileus virginianus). Body condition of white-tailed deer can affect survival and reproduction, which has large repercussions for state-level natural resource agencies that allocate hunting permits. In this study, we investigated how variation in winter weather, spring phenology, habitat composition, and browse quantity affected white-tailed deer body condition across a large spatial scale. Several body condition indicators (e.g., carcass mass, heart fat, antler size) were measured by hunters for 795 deer during September–December 2016–2018 in Wisconsin, USA. Winter severity in the previous year was an unreliable predictor of fall body condition of deer when winters were considered mild or moderate. The timing of spring green-up had a consistent effect on the body condition of all age and sex classes of deer. Earlier spring green-up resulted in heavier fawns and larger antlers among adult males. Region and spring green-up interacted to affect the heart fat of adult females. Earlier springs resulted in adult females in northern and central Wisconsin having a higher probability of heavy heart fat, whereas spring green-up had no effect on adult female heart fat in southern Wisconsin. Effects of habitat differed by age and sex class of deer, and by the body condition metric being evaluated, indicating that there are important physiological differences among age and sex classes of deer that are affected by the environment. Our study demonstrates that the hunting public can contribute large-scale, cost-effective, and quality data to deer monitoring and research projects. It is important that natural resource agencies be able to identify and recruit highly engaged members of the hunting public to ensure project success. The timing of spring green-up can have lasting effects on deer health that can be consistently observed the following fall, which is in contrast to the effects of winter severity that did not appear to persist when previous winters were mild or moderate. We encourage managers in northern or temperate regions to consider measures of spring green-up timing in conjunction with traditional winter severity when making deer population management decisions, such as antlerless tag allocation.     

The economic value of publicly accessible deer hunting land

Access to lands for hunting is widely recognized as critical to sustaining and developing participation in hunting. Deer hunters comprise a substantial portion of recreational hunters in the United States, and numerous stakeholders benefit from the deer management services provided by hunters on public lands. We estimated the economic value of publicly accessible deer hunting land in Michigan using economic models of hunting destinations for hunters in the archery season and for hunters in the firearm season. We found that the amounts of state-owned hunting land, federally owned hunting land, and publicly accessible, privately owned Commercial Forest Act land are significant (P < 0.001) and positive determinants of hunting destinations. The annual economic value of Michigan's publicly accessible hunting land to archery and firearm deer hunters, beyond hunter expenditures, was estimated to be over $80 million. Access to state-owned land accounted for approximately $50 million in annual economic value to Michigan deer hunters. The average per-acre value of publicly accessible hunting land is greatest in the Southern Lower Peninsula of Michigan, where public access is scarce and the majority of hunters reside. Understanding the economic values hunters derive from publicly accessible hunting lands will enable managers and policy makers to make better decisions when considering policy options that involve reducing or increasing public access. © 2011 The Wildlife Society.