中国南方二叠纪海绵礁的成礁模式

广泛发育于我国南方碳酸盐岩台地区的二叠纪生物礁,其中绝大多数属于海绵生物礁。从该地区二叠纪海绵生物礁的内部成礁因素分析,即从主要造礁生物－钙质海绵和钙质藻类等的生物学和生态学特征、埋藏和保存特点等方面进行分析,提出了华南二叠纪海绵生物礁主要是由于其主要造礁生物钙质海绵和钙质藻类独特的生物学特征、生态学特征以及它们的共同作用所形成的。此模式与其它地质历史时期生物礁的成礁模式明显不同。     

广西西部二叠纪生物礁中的海绵、水螅动物化石

广西西部田林,凌云、凤山地区二叠纪生物礁的造礁生物－海绵、水螅化石保存完美。文中描述串管海绵7属：Stylothalamia,Lichuanospongia,Tebagathalamia,Imbricatocoelia,Intrasporeocoelia,Rhabdactinia,Lingyunocoelis;纤维海绵3属：Peronidella,Intratubospongia,Cavusonella;水螅1属Radiotrabeculopora以及海托盘类Calathium,其中Lingyunocoelia spherica为新属新种,Lichuanospongia guangxiensis为新种,其它均为中国南方二叠纪生物礁内常见分子。Senowbari-Daryan(1990)提出以Discosiphonella来代替Cystauletes,Ascosymplegma,Lichunospongia3属,但Lichuanospongia的房室形状和中央腔缺失横板（泡沫板）显著不同于Discosiphonella,因此文中仍把Lichuanospongia视作独立属,Lingyuncoelia gen.nov。与Sollasia,Girtyocoelia相似,但房室外壁缺乏任何种类的穿孔,因此明显不同于后两属,作者仍把Radiotrabeculopora视为螅纲内的一属,而没有归于纤维海绵目,因为它最明显的特征是发育分布较规则的放射状支柱（不能视为骨纤构造）,且那些纵向分布的出水管实际上应视为支柱之间的沟道。     

对一些二叠纪-三叠纪房室海绵属种的分类厘订

Ott(1967)归入Cryptocoelia zineli的材料在房室充填物类型方面与Cryptocoelia zitteli的模式标本完全不同,但又不能归入其它属,故建立新属 Vermicoeliao Senowbari-Daryan(1990)归入Cryptocelia siciliana的材料与该属的定义差别很大,应改归入Guangxispongia Wu。Senowbari-Daryan和Rigby(1988)归入Crystothalamia ramosaGlornocystospongia gracilis的材料与该属模式种的水沟系类型不同,应改归入Imbricatocoelia中。Rigby,Fan和Zhang(1989)归入其新种的Glomocystospongia gracilis的材料之一部分(图版19,图5)应改归入Waagen和Wentzel(1887)建立的Carterina之中,即为Carterina gracilis(Rigby,Fan et Zhang)。     

房室海绵化石的微机研究：性状演化

利用微机对房室海绵进行了１）属的清理工作,２）标准化术语的建立与属的标准化术语描述,３）建数据库,４）性状的统计分析。并发现了房室海绵出水管类型、房室排列方式、房室充填物类型、孔和管、体型和房室形态方面的六大演化趋向。     

房室海绵特征与分类

对房室海绵化石的出水管系统类型进行了总结,增添了１１种新类型,提出以出水管系统的类型作为房室海绵科级及亚目级的分类标准,并据此对房室海绵的分类进行了重大改革。     

钙质海绵之古生态

古生代生物礁中钙质海绵(纤维海绵、房室海绵、硬海绵)的生态位在中三叠世以后被生态竞争能力更强的六射珊瑚所占据.在古生代和中三叠世的钙质海绵礁上,0-10m深度内钙质海绵很发育.由于与钙藻共生,典型的造礁钙质海绵生活在透光带以内,并且在其上部更丰富.钙质海绵礁也会生长到破浪带内并受风浪的破坏而形成倒骨岩和骨屑岩.对古生代的钙质海绵礁而言,倒骨岩和骨屑岩形成于0-3m水深范围内,亮晶骨架岩形成于3-10m深度范围内,灰泥骨架岩形成于10-20m的水深,障积岩形成于20-30m的水深,潜障积岩形成于30-40m的水深.钙质海绵的生长形态与水深的关系与六射珊瑚与水深的关系一样:细枝状的钙质海绵生长在最浅的水中(相当于礁生长带的上部),在稍深的水中(相当于礁生长带的中部和下部)各种形态的海绵都会出现,在更深的水中可以出现特别大的、锥状的海绵.     

湖北利川二叠纪生物礁的埋藏学特征及其环境意义*

二叠纪钙质海绵礁的研究已逾百年,但缺乏埋藏学方面的研究。本文选取鄂西利川典型钙质海绵礁剖面,对造架生物的属种组成、保存状态(直立还是倒伏)、个体大小、古石孔藻包壳和泥晶含量进行统计研究。共鉴定出17个属种的钙质海绵和一个属种的水螅;通过统计研究发现,含钙质海绵的灰岩厚98m,其下部4/5厚度所含钙质海绵以倒伏的为主,上部1/5厚度以直立的为主;含倒伏钙质海绵的灰岩中藻包壳(古石孔藻)不发育,钙质海绵以直立为主的灰岩中,藻包壳发育;钙质海绵的个体大小由海绵灰岩下部而上增大不明显;泥晶基质的含量自海绵灰岩的下部而上明显减少。由此推断:下部的钙质海绵灰岩形成时由于没有藻包壳,大多被风浪打倒。上部具有藻包壳的钙质海绵灰岩,尽管形成时的水动力能量更强,但由于有藻包壳,其抗风浪能力得到加强,所以大多直立保存。     

广西、贵州和川东二叠纪生物礁的钙藻化石及其古生态环境

广西,贵州和川东二叠纪礁相岩石和礁后相岩层内获得了红藻Solenoporella,Gymnocodium,Permocalcu-lus;绿藻Anthracoporella,Mizzia以及显微疑难藻类化石Pseudovermiporella,Tubiphytes等,除了广西隆林有中二叠世茅口期藻类化石外,其它均属于晚二叠世乐平世的分子,这些藻类植物一般生活于热带和亚热带,正常盐度的浅海水内,其水深不大于30m,川东,鄂西晚二叠世礁属于浅水海绵礁,而非深水礁。     

贵州紫云猫场剖面二叠纪-三叠纪之交有孔虫的灭绝过程

本研究通过对贵州紫云地区猫场剖面长兴组上部及二叠系-三叠系界线附近地层进行高密度采样和系统切片,获得有孔虫化石标本1 178枚,共鉴定出有孔虫37属65种,包括类有孔虫4属7种。其中35属63种有孔虫消失在生物碎屑灰岩与微生物岩界线之下,包括所有的类有孔虫,其种级和属级灭绝率分别为96.9%和94.6%。这与华南其它微生物岩剖面类似,进一步表明浅水相有孔虫的灭绝发生在长兴期末期。猫场剖面长兴组生物碎屑灰岩中Dagmarita和Cribrogenerina两属非常丰富,其个体数量分别为209枚和126枚,占总数量的28.4%,是该动物群的优势分子,称为Dagmarita-Cribrogenerina组合。猫场组微生物岩中仅发现有孔虫Earlandia和Postcladella两个属,称为Earlandia-Postcladella组合。     

茅口期■类动物的灭绝过程——灭绝选择性与物种个体大小关系的验证

对茅口期类动物群灭绝过程的分析揭示 ,若将类物种按壳体大小分为两类 ,从动物群物种分异度、物种净增速率的变化来看 ,大个体 (壳长 >6mm)物种与小个体 (壳长≤ 6mm)物种的灭绝过程并无明显的区别。根据壳壁等特征 ,茅口期类可分为南京类、希瓦格类、费伯克类和新希瓦格类 4个主要类群 ,各主要类群及其壳体大小不同的物种灭绝过程有明显的差异。在灭绝事件早期 ,各类群及其壳体大小不同的两类物种所受的影响有所不同。在早期的灭绝中 ,南京类、新希瓦格类、希瓦格类中壳长 >6mm和费伯克类中壳长≤ 6mm的物种显著减少 ,而希瓦格类中壳长≤ 6mm以及费伯克类中壳长 >6mm的物种则未受明显的影响。在茅口期晚期 ,由于灭绝压力的增大 ,除南京类外 ,所有类群及其物种无论个体大小均受到重创 ,导致类动物群物种分异度陡然下降研究结果证明 ,在灭绝强度较小时 ,类动物的生物学特征 ,如壳体大小、壳壁特征对物种的存活率有一定影响 ,而在集群灭绝的高峰期间 ,这些特征则不能发挥有效的作用     

Palaeoenvironmental changes across the Permian/Triassic boundary at Shangsi (N. Sichuan,China)

The section at Shangsi in Sichuan contains one of the most detailed and best records of events during the Permian/Triassic (P/T) mass extinction. Continuous deep water deposition is only punctuated by a minor shallowing in the late Changxingian. The micritic mudstones and wackestones of the Changxingian Dalong Formation contain abundant ammonoids and radiolaria and diverse and common benthic taxa (mostly bivalves and brachiopods) in a thoroughly bioturbated sediment. The presence of a well developed tiered burrow profile is just one line of evidence for a fully oxygenated water column in the late Permian. The faunal crisis occurs in the top few decimetres of the Dalong Formation and severely affected all groups (benthos, nekton and plankton). The extinction coincides precisely with the development of anaerobic and dysaerobic facies. The basal Triassic sediments of the Feixanguan Formation are thinnly‐bedded or laminated silicic marls and contain pyrite and several levels of elevated organic carbon concentrations. The fauna is restricted to rare ammonoids and a few bedding planes covered in Claraia. The presence of abundant coccoid cyanobacteria in these sediments may indicate an unusually simple trophic web in the early Triassic seas as these picoautotrophs are normally grazed by zooplankton, they are rarely directly incorporated into seafloor sediment. The recent discovery of black shales in P/T pelagic sediments of Japan indicates that the anoxic event also affected deep ocean waters and further strengthens the link between extinction and anoxia.     

Facies of two important Early Triassic gastropod lagerstätten: implications for diversity patterns in the aftermath of the end-Permian mass extinction

Two important lagerstätten of Early Triassic gastropods, the Sinbad Limestone (Utah, USA) and the Gastropod Oolite (North Italy) yield about 40% of all described Early Triassic species. This great contribution to the global diversity and the exceptional good preservation render high information content, which characterizes fossil lagerstätten. The Smithian Sinbad Limestone contains the most diverse Early Triassic gastropod fauna. At the type locality, it occurs in single, probably storm-induced shell bed within a series of high energy deposits underlain by intertidal microbial mats and subtidal oolite/peloid shoals. The main shell bed contains about 40 invertebrate taxa. Gastropods, scaphopods, and bivalves are most abundant and form an assemblage, which is dominated by small neritaemorphs, the opisthobranch Cylindrobullina convexa and the scaphopod Plagioglypta (annulated tubes). This assemblage lived on shallow, subtidal soft-bottoms based on sedimentological and ecological characteristics. The Dienerian (to Smithian?) Gastropod Oolite Member (North Italy) has extremely abundant, probably salinity-controlled gastropod faunas with low species richness. Almost monospecific assemblages of Pseudomurchisonia kokeni as well as assemblages with about four species are present in the Gastropod Oolite. Modern hydrobiid mudsnail faunas which are adapted to strongly fluctuating salinity in intertidal to shallow subtidal coastal areas form probably a suitable model for the Gastropod Oolite biota. Gastropods from the Werfen- and Moenkopi-Formation lagerstätten are well preserved compared to other Early Triassic deposits. The high contribution to the global diversity of just two sites suggests very incomplete sampling and preservational bias. However, the low richness of the major faunas reflects depauperate Early Triassic faunas and slow recovery from the Permian/Triassic crisis.     

Recovery from the most profound mass extinction of all time

The end-Permian mass extinction, 251 million years (Myr) ago, was the most devastating ecological event of all time, and it was exacerbated by two earlier events at the beginning and end of the Guadalupian, 270 and 260 Myr ago. Ecosystems were destroyed worldwide, communities were restructured and organisms were left struggling to recover. Disaster taxa, such as Lystrosaurus, insinuated themselves into almost every corner of the sparsely populated landscape in the earliest Triassic, and a quick taxonomic recovery apparently occurred on a global scale. However, close study of ecosystem evolution shows that true ecological recovery was slower. After the end-Guadalupian event, faunas began rebuilding complex trophic structures and refilling guilds, but were hit again by the end-Permian event. Taxonomic diversity at the alpha (community) level did not recover to pre-extinction levels; it reached only a low plateau after each pulse and continued low into the Late Triassic. Our data showed that though there was an initial rise in cosmopolitanism after the extinction pulses, large drops subsequently occurred and, counter-intuitively, a surprisingly low level of cosmopolitanism was sustained through the Early and Middle Triassic.     

Permian and early Triassic isotopic records of carbon and strontium in Australia and a scenario of events about the Permian‐Triassic boundary

The negative shift in δ¹³C values of carbonate carbon at the Permian/Triassic boundary is one of the better documented geochemical signatures of a mass extinction event. The similar negative shift in δ¹³C values in organic carbon from Permian/Triassic boundary marine sediments in Austria and Canada is shown to occur also in marine and non‐marine sediments from Australian sedimentary basins. This negative shift in δ¹³C values is used to calibrate Australian sections lacking diagnostic faunal elements identifying the Permian/Triassic boundary. The minimum in the carbonate ⁸⁷Sr/⁸⁶Sr seawater curve from carbonates across the Guadalupian/Ochoan Stage boundary, mainly from North America, is shown to occur also in brachiopod calcite mainly from the Bowen Basin of eastern Australia, hence providing a second calibration point in the Australian sedimentary record. These two geochemical events support a model of a runaway greenhouse developing about the Permian/Triassic boundary; this is inferred to have contributed to the end‐Permian mass extinction.     

Battenizyga, a new Early Triassic gastropod genus with a discussion of the caenogastropod evolution at the Permian/Triassic boundary

Battenizyga, a new Early Triassic gastropod genus from the Moenkopi Formation of Utah, is described and the speciesAnoptychia eotriassica Batten & Stokes, 1986 is placed in it. The new genus has an axially ribbed planktonic larval shell and a teleoconch with an angulated periphery. This character combination is unknown from the Palaeozoic. Therefore,Battenizyga represents additional evidence that recovery from the end-Permian mass extinction was connected with a faunal turnover. Additionally, the extinction of diverse Palaeozoic groups of the Caenogastropoda in the Permian (e.g., the Pseudozygopleuridae) suggest a turnover. All caenogastropod genera that hold Early Triassic species, have post-Palaeozoic type species and most were not reported from the Palaeozoic. This corroborates the view that there was an intense faunal turnover within the Caenogastropoda.Battenizyga is probably a caenogastropod that is closely related to the superfamily Zygopleuroidea which is abundant in the late Palaeozoic and early Mesozoic.      

Exceptions to the temperature–size rule: no Lilliput Effect in end-Permian ostracods (Crustacea) from Aras Valley (northwest Iran)

The body size of marine ectotherms is often negatively correlated with ambient water temperature, as seen in many clades during the hyperthermal crisis of the end-Permian mass extinction (c. 252 Ma). However, in the case of ostracods, size changes during ancient hyperthermal events are rarely quantified. In this study, we evaluate the body size changes of ostracods in the Aras Valley section (northwest Iran) in response to the drastic warming during the end-Permian mass extinction at three taxonomic levels: class, order, species. At the assemblage level, the warming triggers a complete species turnover in the Aras Valley section, with larger, newly emerging species dominating the immediate post-extinction assemblage for a short time. Individual ostracod species and instars do not show dwarfing or a change in body size as an adaptation to the temperature stress during the end-Permian crisis. This may indicate that the ostracods in the Aras Valley section might have been exceptions to the temperature–size rule (TSR), using an adaptation mechanism that does not involve a decrease in body size. This adaptation might be similar to the accelerated development despite constant instar body sizes that can be observed in some recent experimental studies of ostracod responses to thermal stress.