Upper Devonian Sponges from the Holy Cross Mountains, Central Poland

The rich fauna of Late Devonian (Late Frasnian) siliceous sponges from the Holy Cross Mountains, Poland is composed of 15 species and 11 genera. Both astylospongid demosponges (lithistids) and hexactinosan hexactinellids are present. The following new genera and/or species are proposed: D regulara Rigby and Pisera sp. nov., Jazwicella media Rigby and Pisera gen. et sp. nov., Astyloscyphia irregularia Rigby and Pisera gen. et sp. nov., A. turbinata Rigby and Pisera gen. et sp. nov., Astylotuba modica Rigby and Pisera gen. et sp. nov., Paleoregulara cupula Rigby and Pisera gen. et sp. nov., Paleoramospongia bifurcata Rigby and Pisera gen. et sp. nov., Cordiospongia conica Rigby and Pisera gen. et sp. nov., Paleocraticularia elongata Rigby and Pisera gen. et sp. nov., P gigantia Rigby and Pisera gen. et sp. nov., Polonospongiadevonica Rigby and Pisera gen. et sp. nov., P fistulata Rigby and Pisera gen. et sp. nov., Urnospongia modica Rigby and Pisera gen. et sp. nov., and Conicospongia annulata Rigby and Pisera gen. et sp. nov. The investigated fauna contains the youngest astylospongiids known and the oldest well-preserved, and most diversified Palaeozoic hexactinosans. The sponge fauna constituted a significant element of a brachiopod-coral-sponge assemblage that inhabited a deep slope of the local Dyminy Reef structure, during its final phase of growth, in a clearly hemipelagic setting. This fauna is limited to the intrashelf depression within an incipiently drowned carbonate platform.     

Tide-dominated Middle Devonian sequence from the northern part of the Holy Cross Mountains (Central Poland)

Summary The dolomitic Wojciechowice Formation distinctly differs from the remaining, mainly shaly Middle Devonian succession in northern part of the Holy Cross Mountains (Central Poland). The upper Member of the Formation (Crystalline Dolostone Mb.), in greater part dolomitized but also containing limestone beds, is composed of shallowing-upward cyclothems well exposed in Skaly quarry in the Bodzentyn syncline. The lower parts of the cyclothems, interpreted as subtidal facies, contain fossils characteristic for restricted environments. They are grouped into two assemblages. The first, with brachiopods (largeBornhardtina andEmanuella), massive stromatoporoids, and subordinate gastropods and amphiporoids is related to a deeper subtidal environment, while the second (mainly amphiporoids, gastropods, ostracodes and calcareous algae) is shallower subtidal. Towards the top of succession the fossil content radically decreases. The upper parts of cyclothems are composed mainly of different types of laminites. In these parts of the section, interpreted as intertidal/supratidal units, stromatolites, desiccation polygons, intraformational breccias, and common bioturbations are present. The whole succession was deposited in a low-energy environment, only intermittently affected by high-energy events. For their most spectacular example of this, aBornhardtina-coquinite, a tempestitic origin is proposed. The interval with cyclic sedimentation studied correlates with the dolomitized lower “Unit I” of the Stromatoporoid-Coral Kowala Formation from the southern part of the Holy Cross Mountains, which exhibits sabkha-type cyclicity. The differences in development of cyclothems in both regions resemble outer and inner part of an extensive platform, and correspond well with basic trends of the Lower-Middle Devonian transgression in the Holy Cross Mountains. The general succession of formations deposited during this process coincides with transgressive events on Johnson's eustatic curve for the Devonian.     

New Devonian and Carboniferous bryozoans of the Holy Cross Mountains (central Poland)

New bryozoans from the Middle-Upper Devonian (Eifelian-Frasnian) and the Lower Carboniferous (Tournaisian) of the Holy Cross Mountains in central Poland are described: Bigeyella sparsa gen. et sp. nov., B. separata gen. et sp. nov., Eridopora singula sp. nov., Leptotrypa pulchra sp. nov., Kysylschinipora klarae sp. nov., Coelotubulipora rara sp. nov., Alternifenestella genuina sp. nov., Exfenestella polonica sp. nov., and Rectifenestella localis sp. nov. Some paleogeographic and stratigraphic aspects of the Paleozoic deposits of this region are discussed and main bryozoan localities are described.     

First Record of Soft Tissue Preservation in the Upper Devonian of Poland

Soft tissue preservation is reported from Upper Devonian deposits of the Holy Cross Mountains, central Poland, for the first time. The preserved soft tissues are muscles associated with arthropod cuticle fragments. The muscles are phosphatized with variable states of preservation. Well-preserved specimens display the typical banding of striated muscles. Other muscle fragments are highly degraded and/or recrystallized such that their microstructure is barely visible. The phosphatized muscles and associated cuticle are fragmented, occur in patches and some are scattered on the bedding plane. Due to the state of preservation and the lack of diagnostic features, the cuticle identification is problematic; however, it may have belonged to a phyllocarid crustacean. Taphonomic features of the remains indicate that they do not represent fossilized fecal matter (coprolite) but may represent a regurgitate, but the hypothesis is difficult to test. Most probably they represent the leftover remains after arthropod or fish scavenging. The present study shows that soft tissues, which even earlier were manipulated by scavenger, may be preserved if only special microenvironmental conditions within and around the animal remains are established.     

A new Middle Devonian gasterocomid crinoid from central Turkey and revision of the Gasterocomidae

A juvenile cup, disarticulated arm plates, and columnals of the crinoid Arachnocrinus sarizensis n. sp. are described from the Eastern Taurus Mountains of central Turkey. This is the first Paleozoic crinoid based on a cup and arm plates reported from Turkey. Suggested revision of the Gasterocomidae includes transfer of Arachnocrinus and Ancyrocrinus to the family and rejection of Kopficrinus from the family. The stratigraphic range of Arachnocrinus is extended upward into the Middle Devonian from the Early Devonian. The Devonian of Turkey is of special interest because it includes Laurasian and Gondwanan components on different tectonic blocks. The paleogeographic range of Arachnocrinus is extended from the North American plate onto the Anatolian block, which would have been located on the southern edge of the Paleotethys Sea in the Middle Devonian in a shelf basin off the northern coast of Gondwana at approximately 42° south latitude.     

First <Emphasis Type=Italic>Sycidium</Emphasis> (Charophyta) from the Upper Devonian of Thuringia (Germany)

The well-preserved silicified fragment of a utricular envelope assigned to Sycidium aff. clathratum Peck, discovered in the earliest Famennian “incisus bed” of the Bohlen section near Saalfeld in Thuringia (Central Germany), is the first evidence of charophytes present in the West European Late Devonian. The occurrence of 16 longitudinal ridges is shared with the basal Mississippian S. clathratum. The specimen shows delicate structures of the external layer including ovoid elements at its basal end which are interpreted as casts of rudimentary bract cells and/or bracteoles, rarely observed in the fossil record of charophytes. S. aff. clathratum is among the rare Sycidiaceae that survived the terminal Frasnian Kellwasser Event. This single specimen was probably attached to a piece of wood and thereby transported into fully marine basinal environments that characterize the Famennian of the Bohlen section.     

Middle Devonian dendroid graptolites from the Brilon Reef area (Rheinisches Schiefergebirge, Germany)

Dendroid graptolites from the Middle Devonian (Givetian, middlePolygnathus varcus Biozone) of the Rheinisches Schiefergebirge are described for the first time. The fauna of the Burgberg locality south of the Brilon reef comes from a black shale layer in a succession of calcareous debris flows. The fauna includes five species ofDictyonema andRuedemannograptus described in open nomenclature. The fragmentation of the material is conside-red as an indication of post-mortem transport.      

Kimmeridgian hardground-sequence boundary from the Mesozoic margin of the Holy Cross Mountains (central Poland): implications for the evolution of the northern Tethyan carbonate shelf

The sedimentary succession in central Poland records significant changes in facies at the turn of the Planula and Platynota zones in the Upper Jurassic, expressed by the drowning of the ramp-type platform and development of an extensive isochronous marl horizon. The topmost level of the marl horizon is a regional hardground, which is interpreted as the third-order sequence boundary Kim 1. In some areas, the hardground was eroded and is only preserved as bored and encrusted clasts. The composition of the borings may indicate that colonization and recolonization of the clasts took place in an extremely shallow water environment. The overall low level diversity of clast-encrusting organisms and their occurrence on both sides of clasts indicates frequent overturning and high current activity. However, other extrinsic factors, such as salinity fluctuations, may have been involved. The final redeposition and burial of the clasts were related to subsidence through widespread reactivation of Paleozoic faults. Comparison with Middle Oxfordian–Lower Kimmeridgian sequences of central and southern Poland indicates that the carbonate ramp morphology and paleoenvironmental conditions were periodically subjected to significant modification during phases of extensional tectonics, with the development of fault-controlled intra-platform ridges and basins. These features may have been the NE continuation of parallel swells and basins developed in the southern part of the carbonate platform adjacent to the Tethys.     

Morphology and wall ultrastructure of some Middle Devonian dispersed megaspores from northern Poland

Specimens of dispersed Middle Devonian megaspores have been isolated from core samples from the Miastko 1 borehole in Western Pomerania. Comprehensive investigations using light, scanning and transmission electron microscopy supplement previous information on morphology and gross structure and provide data on spore wall ultrastructure of four megaspore species. Corystisporites acutispinosus is azonate; the inner layer is laminate, and the lumen is lined by a thick, laterally continuous lamina. The outer layer consists of small, tangentially aligned tabular elements that become wider, more extensive and irregularly arranged toward the outside. Coronispora variabilis is a coronate megaspore; the inner layer appears homogenous and is probably lamellate. The outer layer consists of elongate, cylindrical, branching elements that are overlaid within the proximal part of the body by a lamellate, compact, almost homogenous layer. Grandispora ciliata is pseudosaccate. The inner body is laminate with laminae thickening and becoming less continuous and less tightly packed toward the outside. The outermost region of the inner body and the innermost region of the outer envelope consist of tabular and cylindrical elongate units. The bulk of the outer wall is almost homogenous, and near the surface it is granular. Pomeranisporites subtriangularis is pseudozonate. The inner layer appears homogenous except for the presence of a single innermost lamina. The inner part of the outer layer may represent small tabular and cylindrical elements, and the outer part comprises folded laminae. The megaspores studied share numerous features of morphology and wall ultrastructure with the lycopsids, putative lycopsids, and some enigmatic Devonian plants.     

New Famennian bryozoans (Upper Devonian) from Azerbaijan and Armenia

New bryozoans are described from the Famennian of Southern Transcaucasia: Petalotrypa myunkhbalaensis sp. nov. and Eridotrypella danzikensis sp. nov. (order Trepostomata) and Rectifenestella kadrluiensis sp. nov. and R. famenniensis sp. nov. (order Fenestrata). The paleobiogeographic relations that existed between Nakhichevan (Azerbaijan) and Armenia and other regions of Eurasia and the United States at the end of the Devonian are investigated.     

New finds of vertebrates in the Middle Devonian Brandenberg Group (Sauerland, Northwest Germany)

The arthrodires of the Eifelian Brandenberg Group of Hagen-Ambrock and Lasbeck (Northwest Germany) are described. A new tuberculated holonematid arthrodire (Holonema bruehni n. sp.) is represented by plates of the trunk shield and some plates of the head region. The tuberculation is here regarded as the plesiomorphic character of the Holonematidae compared to an ornamentation of ridges in advanced genera. This agrees with the stratigraphical occurrence of the new form. Additionally, several other plates belonging to at least three different taxa of coccosteid arthrodires have been found. Despite the absence of complete skeletons, the placoderm fauna is more diverse than the fauna of the classical Scottish Eifelian localities.     

New finds of vertebrates in the Middle Devonian Brandenberg Group (Sauerland, Northwest Germany)

Placoderms (Ptyctodontida), acanthodians (incl.Atopacanthus?ambrockensis n. sp.), actinopterygians, dipnoans and crossopterygians — including the first Middle Devonian osteolepids from the Rheinisches Schiefergebirge — are described from the lower Brandenberg Group of Hagen-Ambrock and Lasbeck (northern Sauerland, Northwest Germany). The composition of the fauna is similar to that of contemporaneous Scottish and Baltic Old Red localities, even though typical marine vertebrates are present too (e.g. selachians). This indicates — like the invertebrates — rather a marginal marine deposition.     

New Conodont Species from the Famennian (Upper Devonian) of the Urals

New conodont species of the genus Polygnathus from the Upper Famennian of the Urals and Cis-Urals are described. Po. tsygankoi sp. nov. is distributed from the expansa‑praesulcata conodont zones. This species occupies an intermediate position in the phylogenetic lineage Polygnathus atasuensis–Po. tsygankoi sp. nov.–Po. lobatus sensu Kononova. Po. serriformis sp. nov. is known from the praesulcata Zone. Evolutionary relationships of Po. serriformis sp. nov. are unknown.     

Angustidontidae (Crustacea: Malacostraca) from the Upper Devonian of Ukraine and Belarus

Previously unknown specimens of angustidontid crustaceans from the Frasnian and Famennian of Ukraine and from the Famennian of Belarus are described.     

Cyrtocrinids (Crinoidea) and associated stalked crinoids from the Lower/Middle Oxfordian (Upper Jurassic) shelfal deposits of southern Poland

Five cyrtocrinid crinoid taxa previously unknown from the epicratonic deposits of Poland, as well as associated millericrinids and isocrinids, are described. The studied materials were derived mainly from the Lower and Middle Oxfordian, but crinoids are also from uppermost Callovian and/or lowermost Oxfordian sediments of the Polish Jura Chain (southern Poland). The crinoids, preserved as more or less complete (e.g., basal circlets) cups, include Lonchocrinus dumortieri, Phyllocrinus belbekensis, Remisovicrinus polonicus, Remisovicrinus aff. polonicus, Tetracrinus moniliformis and Sclerocrinus sp. The occurrence of Remisovicrinus polonicus in the late Middle Oxfordian of the southern Poland is confirmed. Moreover, the present study extends the geographic range of all cyrtocrinid species studied and discusses their unusual environmental adaptations.