Geochemical evidence for the formation of a large miocene “travertine” mound at a sublacustrine spring in a soda lake (Wallerstein castle rock, nördlinger ries, Germany)

Summary “Travertines” (tufa pinnacles) of the Miocene Riescrater basin have been investigated to test whether carbon, oxygen and strontium isotopes can be used for the recognition of fossil subaquatic spring deposits in high-alkalinity settings. The Ries basin “travertines” have so far been interpreted as a product of subaerial to sublacustrine artesian springs discharging calcareous groundwater into a freshwater or slightly saline lake. However, recent studies on microfacies and fabric development propose a formation at Ca²⁺-supplying sublacustrine springs of a soda lake. Geochemical analysis of “travertines” of the castle rock Wallerstein, including “sickle-cell” limestones, thrombolites, non-skeletal stromatolites, and speleothems, now support the latter interpretation. High Sr contents surpassing that of the contemporaneously formed dolomitic algal biocherms of the lake shore point to an aragonitic composition of primary precipitates. the δ¹³C values of diagenetically moderately to weakly altered “travertine” facies types are in the same range of the impact-brecciated Upper Jurassic limestones, thus, are inconsistent with a mixture of soil-derived CO₂ and CO₃ ²⁻ from the Jurassic limestones. In addition, the δ¹⁸O values are too high to support a significant contribution of CO₃ ²⁻ by meteoric waters seeping through marine Jurassic limestones. Instead the δ¹³C and δ¹⁸O values indicate an origin of the CO₃ ²⁻ from a lake water body characterized by evaporation. This is consistent with a sodium-rich lake water as indicated by high sodium contents of aragonitic algal bioherms of the lake shore. The⁸⁷Sr/⁸⁶Sr isotope ratio of the “travertine” mound carbonates are consistent with calculated mixing of spring waters discharging from the crystalline basement and lake water high in dissolved inorganic carbon. This points to an origin of the divalent cations from sublacustrine spring waters. In turn,⁸⁷Sr/⁸⁶Sr isotope ratios of green algal reef carbonates of the lake shore are closer to that of the Upper Jurassic carbonates, due to surface run-off from surrounding limestone uplands.     

Fossilization of the cells of natronophilic endoevaporite cyanobacterium ‘<Emphasis Type="Italic">Euhalothece natronophila</Emphasis>’ in a modelling system

Laboratory simulation of fossilization of cyanobacterial cells in the high-carbonate medium in the presence of calcium was carried out for the haloalkaliphilic natronophilic cyanobacterium ‘Euhalothece natronophila’ Z-M001. This organism was isolated from the Magadi soda lake, where the bioherms consisting of mineralized coccoid cyanobacteria were found in the Quaternary sediments. The structural and chemical heterogeneity of the minerals produced during this process was established, with calcium carbonate and trona being the main products. The differences in the process of cyanobacterial cell carbonatization in soda lakes and marine or freshwater systems were determined. Initial precipitation of calcium carbonate was shown to occur due to a chemical reaction not involving cyanobacteria. At the subsequent stages, amorphous CaCO₃ is sorbed and crystallized on the surface of some of the cells within a cyanobacterial population, resulting in formation of a shell-like mineral layer. The cells embedded in trona in the same system were shown to undergo deformation and destruction. In both cases the mineralized cells were shown to lose their photosynthetic activity.     

Effects of plant harvesting and nutrient enrichment on phytoplankton community structure in a shallow urban lake

The utility of shallow water bodies in urban environments is frequently compromised either by dense beds of submerged plants or cyanobacterial blooms associated with nutrient enrichment. Although submerged plants are often harvested to facilitate recreational uses, this activity may alter the phytoplankton community, which in turn, also may restrict the use of the lake. We tested whether (i) plant harvesting reduced the abundance of flagellate algae and increased the abundance of cyanobacteria, and (ii) whether increasing levels of nutrient enrichment caused shifts in the dominance of heterocytous cyanobacteria, non-heterocytous cyanobacteria and Chlorophyta, in a shallow urban lake in Southern Australia as has been observed for shallow Danish lakes in previous studies. These predictions were tested with large (3000 l), replicated mesocosms in a warm, highly productive, shallow lake densely colonised by the submerged angiosperm, Vallisnaria americana Michaux. The heterokont algae, Chlorophyta, Cyanobacteria and Cryptophyta were the most numerous algal divisions in the lake. The Euglenophyta, although uncommon in early summer, became more abundant towards the end of summer. The Dinophyta and Charophyta were rare. The abundance of the heterokont algae and Euglenophyta was significantly reduced by plant harvesting even after plants had partially re-established 18 weeks after initial harvesting. The decline in the Euglenophyta in response to plant harvesting is consistent with earlier findings, that the relative abundance of flagellate algae tends to be greater in the presence of submerged plants. Contrary to our prediction, we found that the Cyanobacteria did not increase in response to plant harvesting, however the response may be altered under higher nutrient levels. Algal responses to nutrient enrichment in the presence of dense V. americana plants generally followed the patterns observed in shallow Danish lakes despite the large differences in climatic conditions. Both studies found that the abundance of heterocytous cyanobacteria declined at higher levels of nutrient enrichment, whereas non-heterocytous cyanobacteria and chlorophytes increased.     

Foraminifera diversity changes and paleoenvironmental analysis: the Lower Cretaceous shallow-water carbonates of San Lorenzello, Campanian Apennines, southern Italy

A high-resolution stratigraphic study, carried out on the carbonate platform strata of the San Lorenzello section (Matese Mountains, southern Italy), Valanginian–Hauterivian in age, has allowed to: recognise lithofacies and their associations; assign the lithofacies associations to specific environments and individuate early meteoric diagenetic modifications, recurring at specific horizons. In this frame the vertical variation of benthic foram diversity has been analysed. On the whole, foraminiferal genera diversity decreases from open to restricted marine environments. Moreover, a climatic control on carbonate sedimentation is suggested by a Milankovitch cyclicity organised in elementary cycles, bundles and superbundles as well as by diagenetic characteristics testifying that humid and arid conditions alternated during the Early Cretaceous times. The orbital cyclicity is also documented by foraminiferal diversity changes, even if some discrepancies between the lithofacies and the diversity locally occur. Therefore, the above diversity changes do not appear to provide sufficient information for the sequence-stratigraphic interpretation of shallow-water carbonates.     

Sequence stratigraphy and paleo-oceanography of an open-marine mixed carbonate/siliciclastic succession (Late Jurassic,Southern Germany)

Summary The Late Jurassic epicontinental sea of South Germany protruded far to the North forming a wide bay which was rimmed by shallow-water platforms (Swiss and French Jura). This wide shelf is characterized by extensive downslope mud accumulations including siliceous sponge buildups. The bioherms are aligned along the more pericontinental parts of this shelf, which graded to the South into the Helvetic Basin of the Tethys Ocean. Five sedimentary cycles of Oxfordian to Middle Kimmeridgian age (ox. 2–ox. 3, ki.1.–ki.1.2, ki.3, ki.1.3–ki.2.1, ki. 2.1–ki. 2.2) were used for interregional correlation. Each cycle is characterized by a vertical suite from marl to pure limestone. The bases of the marls are characterized by abundant open-marine fossils, glauconite and phosphate (fish teeth and pellets) and interpreted as condensed sections. Deepening is indicated by bioherms changing their growth form before demise from large structures into small isolated buildups, which commonly occur within deeper water. Sequence boundaries, are present at the transition from marl into limestone. Two phases (middle ki. 1.3 and ki. 1/2) of debris-flow deposition, one accompanied by the sudden spreadout of biostromes and basinward shift of bioherms, are interpreted as lowstand phases associated with sequence boundaries. The remaining sedimentary cycles described here lack such lowstand deposits. Instead, corresponding positions in the cycles are characterized by omission features. This problem is still unresolved. It may either be due to sediment trapping on the ramp or to a changing origin of the cycles compared. It is proposed here that highstand sediments, having higher contents of fine-grained siliciclastics than lowstand deposits, formed during a humid and warm climate with high rates of continued runoff and sediment transport. Lowstand deposits consist of pure offbank carbonates, because the associated drier climate reduced fluvial input of terrigenous material. Corresponding patterns of climatic change are also seen in platform sediments from the Swiss Jura Range; however, climatic cycles correlate with eustatic sea level fluctuations in only about 50% of the cases. This misfit may partially result from problems with biostratigraphic correlation (boreal— Tethyan). Small-scale sedimentary cycles with an average duration of 66’000 years (Mutabilis chron, ki. 2) to 95’000 years (Planula chron, ox. 3) provide a tool for detailed stratigraphic correlation in biostromes, small scale lenticular bioherms (1–2 m thickness) and large bedded bioherms (many tens of metres thickness). Interruptions of bioherm growth are due to temporal oxygen deficiency related to plankton blooms. The bathymetrically deepest bioherms-small and lenticular in shape—therefore suffered the highest number of ecological break-downs, whereas massive bioherms continuously remained above the critical level.     

Ein Diplopode (Tausendfüßler i. e. S.) aus den lakustrinen Karbonaten des Nördlinger Rieses (Miozän,Süddeutschland): Morphologie und Integumentstruktur

An eugnath diplopode is described from Miocene lacustrine carbonates of the impact crater Nördlinger Ries. A further systematic classification is impossible, because gonopodes, which are the most important taxonomic characteristics, are not preserved. The primary calcite-impregnated integument has been dolomitized with structural preservation. In fractures, a differentiation of endo-and exocuticle is visible by distinct ultrastructures. The epicuticle is not preserved. Concentric arched structures, which display the orientation of former chitin-protein-microfibres, are preserved by the arrangement of dolomite crystallites. Concerning the diagenetic history of the Ries-lake-carbonates this is an argument for an early, near-surface dolomitization.     

Community succession in small bioherms of algae and sponges in the Lower Permian of New Mexico

Toomey. Donald F. & Cys. John M. 1979 01 15: Community succession in small bioherms of algae and sponges in the Lower Permian of New Mexico. Lethaia , Vol. 12. pp. 65–74. Oslo. ISSN 0024–1164.
Small organic mounds outcrop in the Lower Permian Laborcita Formation of the northern Sacramento Mountains of southcentral New Mexico. These small organic structures, up to I'm in height and 3 m in diameter, are unique because they show a well developed vertical biolic zonation. This zonation consists of an initial pioneer community composed of small digitate or plumose colonies of algae and foraminifers (Stage 1). This is followed by a climax community that is first dominated by laminar colonies of the red alga Archaeolithophyllum (Stage 2). and followed by a terminal community composed of minor Archaeolithophyllum , erect heliosponges and brachiopods, dominated by the form Composita (Stage 3). Evidence suggests that the mounds initially developed in shallow water within an open lagoon but. with continued transgression and rapid deepening of the depositional environment. the biota responded to the changes. The organic buildups were killed off by sudden influx of clastics from a tectonically active nearby landmass.     

Brackish to hypersaline facies in lacustrine carbonates: Purbeck Limestone Group,Upper Jurassic–Lower Cretaceous,Wessex Basin,Dorset, UK

Sedimentary facies and stratigraphic architecture of non-marine carbonates are controlled by a range of environmental parameters, such as climate, hydrology and tectonic setting, but the few published facies models do not account for this variability. Outcrop and petrographic observations from the Mupe Member of the Purbeck Limestone Group (Upper Jurassic–Lower Cretaceous) in Dorset, southern England, are the basis for new depositional models of non-marine microbialites and associated carbonates in an extensional basin. Ten facies are defined, described and grouped into five facies associations. The Mupe Member is characterised by accumulation of in situ microbial mounds developed around tree remains preserved as moulds and silicified wood. Mounds occur within three stratigraphic units, separated by three palaeosoils, characterised by less-porous, bedded, inter-mound packstone–grainstone that commonly onlap mound margins. Mounds are developed mainly in the shallowest areas of the lake, as indicated by their shapes, facies relationships and association with palaeosoils. These microbial mounds are compared to modern (Laguna Bacalar, Mexico and Great Salt Lake, Utah, USA) and ancient (Eocene Green River Formation, Uinta Basin, Utah, USA) analogues to assess their value as palaeoenvironmental indicators. Facies transitions indicate an earlier, brackish-water lake and a later hypersaline lake for the Mupe Member, both within a semi-arid climate setting in an extensional basin. The fact that the microbialites are covered by evaporitic strata, together with sedimentological, palaeontological and stable isotope data, suggest that there was a sharp change from through-flowing brackish-water, to a closed hypersaline, lacustrine system.     

Carbonate horizons, paleosols, and lake flooding cycles: Beds I and II of Olduvai Gorge, Tanzania

This study documents the petrology and stable isotope geochemistry of carbonates from six horizons from Beds I and II of Olduvai Gorge, Tanzania. The studied succession, immediately below and above Tuff IF, consists of interbedded waxy and earthy claystones with discrete carbonate horizons and thin sandstones. The succession was deposited in response to repeated flooding and withdrawal of a saline-alkaline lake. The carbonates and their overlying disconformities are important because they help define the surfaces on which hominin activity took place and allow very high-resolution correlation of geographically separated levels of hominin exploitation.The range of different carbonates includes unambiguous land-surface and pedogenic features including calcified rootmat horizons, rhizocretions, and micritic nodules, together with less determinate sparry calcite nodules. Stellate nodules are interpreted as pseudomorphs after sulfate-roses. The carbonate nodules are synsedimentary features, truncated by fluvial and other erosional surfaces. The isotopic composition of the carbonates is variable with δ¹⁸O ranging from −7.0‰ to −4.3‰, and δ¹³C from −8.5‰ to −1.6‰. A covariant increase in δ¹³C and δ¹⁸O repeats in each carbonate horizon and in individual nodules (inner to outer layers): it reflects the evolution of synsedimentary groundwaters. At times of low lake level, the carbonates started to precipitate from meteoric waters with low isotopic values and continued to form as lake levels rose and the waters became increasingly saline. Some of the samples have a last-stage cement of strontium rich dolomite, which supports late-stage flooding by the saline-alkaline lake. Previous studies of carbonate horizons from Olduvai have interpreted carbon isotope values in terms of changes in C₃ and C₄ plants that colonized the land surface. This study demonstrates that in some instances the isotope values from carbonates deposited in these lake marginal settings reflect changes in hydrology rather than vegetation.     

Late Paleozoic reef mounds of the Carnic Alps (Austria/Italy)

The Late Paleozoic (early Kasimovian-late Artinskian) sedimentary sequence of the Carnic Alps (Austria/Italy) is composed of cyclic, shallow-marine, mixed siliciclastic-carbonate sedimentary rocks. It contains different types of skeletal mounds in different stratigraphic levels. The oldest mounds occur at the base of the Auernig Group, within a transgressive sequence of the basal Meledis Formation. These mounds are small and built by auloporid corals. Algal mounds are developed in the Auernig Formation of the Auernig Group, forming biostromes, and Lower Pseudoschwagerina Limestone of the Rattendorf Group forming biostromes and bioherms. The dominant mound-forming organism of these mounds is the dasycladacean alga Anthracoporella spectabilis. In mounds of the Auernig Formation subordinately the ancestral corallinacean alga Archaeolithophyllum missouriense is present, whereas in mounds of the Lower Pseudoschwagerina Limestone a few calcisponges and phylloid algae occur locally at the base and on top of some Anthracoporella mounds. Mounds of the Auernig Formation formed during relative sea level highstands whereas mounds of the Lower Pseudoschwagerina Limestone formed during transgression. The depositional environment was in the shallow marine, low-turbulence photic zone, just below the active wave base and lacking siliciclastic influx. The algal mounds of the Carnic Alps differ significantly from all other algal mounds in composition, structure, zonation and diagenesis; the formation of the mounds cannot be explained by the model proposed by Wilson (1975). The largest mounds occur in the Trogkofel Limestone, they are composed of Tubiphytes/Archaeolithoporella boundstone, which shows some similarities to the “Tubiphytes thickets” of stage 2 of the massive Capitan reef complex of the Guadalupe Mountains of New Mexico/West Texas.     

Early Miocene repetitive vegetation and climatic changes in the lacustrine deposits of the Rubielos de Mora Basin (Teruel, NE Spain)

Pollen analysis of a Lower Miocene succession from the Rubielos de Mora Basin (NE Spain) has been carried out with the aim of reconstructing the flora, vegetation and climatic changes. Previous paleobotanical studies on these sedimentary rocks and adjacent areas interpreted very diverse climates for the Early Miocene: from humid temperate to dry subtropical. In this study, a rich thermophilous pollen spectrum and a diverse subarid flora including Nitraria, Caesalpiniaceae, Ephedra and Acacia, indicative of a dry subtropical climate are identified. On the other hand, mesothermic taxa with high water requirements are also abundant. Therefore, the pollen assemblages evidence the juxtaposition of very contrasted environments: the presence of subdesertic taxa, typical of plants growing in the lowlands and conditioned by a long warm, dry season, together with others with very high water requirements, needing constant water. This can be explained by the presence of the Rubielos de Mora Lake providing local conditions for developing riparian forests. The vegetation was clearly controlled by the water availability under a subtropical and dry-seasonal climate. Pollen changes along the succession, which coincide with sedimentological changes, are related to climatic variations. Alternation in pollen taxa (thermophilous–dry vs. mesothermic–riparian) reflects the influence of the cyclicity of temperature and precipitation on the lake level and vegetation.     

氮、磷对热带浅水湖泊惠州西湖蓝藻的控制

湖泊富营养化常导致蓝藻生物量的增加,水质恶化.于2011年2月至12月对热带浅水湖泊惠州西湖六个湖区的蓝藻群落结构进行研究,以了解其时空变化特征及主要影响因素.结果表明,以沉水植物为优势的元妙观湖区与南南湖蓝藻无明显的优势种;平湖蓝藻优势种为银灰平裂藻(Merismopedia glauca)和湖丝藻(Limnothrix sp.),南丰湖、北丰湖和北南湖的主要优势种均为银灰平裂藻(Merismopedia glauca).平湖、南丰湖、北丰湖和北南湖蓝藻丰度及生物量存在显著的季节变化.相关分析显示惠州西湖夏季蓝藻生物量受氮、磷盐控制.冬季温度的影响,蓝藻生物量与氮、磷的相关性不显著.     

Lower Ordovician sponge bioherms in the Makkol Formation, Taebaeksan Basin, mideast Korea

Summary Isolated sponge bioherms are documented from the Lower Ordovician Makkol Formation of the Taebaek Group in the Taebaeksan Basin, mideast Korea. They are formed by an association of a lithistid spongeArchaeoscyphia, a receptaculidCalathium and stromatolitic algae, and share many features with the Lower Ordovician buildups known elsewhere. These bioherms were established in an incised bottom and reached up to about 1 m in height. As the bioherms grew upward, they were more severely affected by intense wave action and frequent storms, which eventually perished the bioherms. The occurrence ofArchaeoscyphia-Calathium association suggests a close biogeographic link between Korea and North China, supporting the paleogeographic model that the Taebaeksan Basin was connected through contiguous shallow waters to North China in the early Paleozoic.     

Late Holocene palaeoecology of Lago Verde: evidence of human impact and climate change in the northern limit of the neotropics during the late formative and classic periods

Multiproxy analysis (pollen, diatom, charcoal) on a 6 m core from Lago Verde (Sierra de Los Tuxtlas), shows evidences of environmental changes and human impact on the evergreen rainforest on the tropical lowlands of eastern Mexico during the last ca. 2,800 years. Prehistoric human occupation is recorded since the late Formative throughout the middle Classic (250 b.c.–ca. a.d. 800) by the presence of maize pollen, a low abundance of tropical arboreal taxa and a high abundance of herbaceous pollen and charcoal particles. This occurred under a scenario of very low lake levels from which dry conditions are inferred based on the dominance of aerophilous and periphytic diatom taxa. After ca. a.d. 800 the site was abandoned and forest regeneration started, at the same time higher lake levels, an indication of more humid conditions, were established. In the absence of human disturbance, tropical forest regeneration was rapid (ca. 200 years). Fluctuations in pollen composition during times of low human population at the site are related to climate variability, with the highest tropical forest diversity and lake levels recorded during the Little Ice Age. Modern human impact is also recorded and compared with the prehistoric deforestation event. Comparison with palaeoecological records from Yucatan and the highlands of central Mexico offers a Mesoamerican perspective of climatic variability giving evidence that the late Formative and early to middle Classic demographic expansion occurred under a scenario of climates dryer than present, with the Postclassic characterized by moister conditions. The end of the Classic (ca. a.d. 800–1000) is identified as a period of rapid climate change which marks one of the most important cultural transitions in Mesoamerica.     

Allelopathy among Cyanoprokaryota and Other Algae Originating from Lake Neusiedlersee (Austria)

During the late eighties, the eutrophic shallow lake Neusiedlersee located at the austrian – hungarian border showed a massive development of cyanobacteria, which coincided with a strong decline of chlorococcales. Besides a change in nutrient conditions, allelopathy may have contributed to the dominance of cyanobacteria, too. To get a first insight into potential allelopathic effects, 13 cyanobacteria strains were tested originating for the most part from lake Neusiedlersee. Plate diffusion assays showed allelopathic activity of 6 strains indicating chemical interactions between cyanobacteria and other algae.     

从塔北隆起奥陶纪钙藻化石探讨奥陶纪的古环境

塔里木盆地塔北隆起轮南46井、英买1、2井奥陶纪石灰岩内含有大量的钙藻化石、蓝细菌以及疑难微体化石。这些钻井岩芯均在深达5000-6200m的地下深部取得。钙藻主要为绿藻类的Dasyporella,Ver‘miporella,Moniliporella以及?Plexa;红藻类的管孔藻类Solenoporaceans;钙化蓝细菌则有Girvanella,Botomaella,?Subtifloria等;疑难微体化石有Bevocastria,Nuia,Rothpletzella。这些钙藻生活于热带或亚热带正常盐度的浅海水内,其水深不到20m。世界各地的奥陶纪Vermiporella均位于古赤道的两侧,这表明它们是在气候炎热、温暖海水中生活的一类海洋藻类。Girvanella以藻灰结核和内碎屑最为常见,某些球粒可能代表Girvanella破碎后形成的单管或小棒。塔里木盆地钙藻植物群相似于哈萨克斯坦、波罗的海周围地区以及北美同时代植物群,这表明这些钙藻和蓝细菌化石具有遍布于全球的性质。塔北隆起早、中奥陶世沉积属于典型的碳酸盐岩台地相沉积。到晚奥陶世时,碳酸盐岩沉积被浅水陆棚沉积所取代,以陆源碎屑岩为主,夹少量的碳酸盐岩。     

Facies features and paleoenvironmental reconstruction of the Early to Middle Devonian syn-rift volcano-sedimentary succession (Padeha Formation) in the Eastern-Alborz Mountains,NE Iran

The Padeha Formation in the Eastern-Alborz Mountains (northeast Iran) is an Early to Middle Devonian syn-rift succession. Siliciclastic rocks (conglomerates, sandstones, mudstones, and red paleosols) to non-marine carbonates (yellowish calcretes, dolocretes, stromatolites, and sandy bivalve packstone) associated with volcaniclastic rocks (basalts or andesites and tuffs) are present in this succession. Facies analysis led to the recognition of three facies associations that are deposited in three terrestrial environments (alluvial fan, distal fan, and palustrine/lacustrine). Its sedimentary fill pattern and association with mafic volcaniclastic rocks indicate that this formation is related to the initial phase of a rift basin (intracratonic rift). Paleosols of this formation are good indicators for recognition of paleoenvironmental factors (climatic changes from semi-arid to sub-humid with annual fluctuation, small and short gross-like vegetation cover, subaerial exposure, very shallow lake, volcanic source rocks, and early meteoric diagenesis).     

<Emphasis Type="Italic">Alona alsafadii</Emphasis> n. sp. from Yemen,a primitive,groundwater-dwelling member of the <Emphasis Type="Italic">A. karua</Emphasis>-group

The effect of algae on the production of musty-smelling compounds by actinomycetes was studied. Streptomyces spp., causing intensive musty odor, were isolated from hypertrophic Lake Kasumigaura and cultured in association with algae from the same lake. Isolate E and I effectively utilized the cyanobacteria, Microcystis aeruginosa and Anabaena spiroides, and the diatom, Synedra acus, as a carbon source and produced a musty-smelling 2-methylisoborneol in the shaken sediment cultures. High populations of algae and actinomycetes, and aerobic condition in the sediment seem responsible for the occurrence of musty odor in Lake Kasumigaura.     

The role of pingos in the development of the Dzhangyskol lake–pingo complex,central Altai Mountains,southern Siberia

Dzhangyskol is a small lake of glacial origin in the central part of the Altai Mountains in southern Siberia. Pollen stratigraphies and chronologies of two cores record the vegetational development of the area from the Late Glacial treeless landscape to the forest and steppe of today. The modern lake is a remnant of a much larger ice-dammed lake, which was reduced in size and then temporarily drained after diversion of the inflowing mountain meltwater stream, which had low δ¹⁸O values. The dry lake floor allowed development of permafrost and small pingos (frozen mounds of lake sediments). With the onset of greater climatic humidity in the mid-Holocene, the input of local water with higher δ¹⁸O caused a rise in lake level, drowning the earlier pingos. Growth of a broad fen on the margin of the lake led to formation of a modern pingo complex.