山东半岛北部沿岸岛屿蝮属一新种(爬行纲,有鳞目,蝰科)

根据对山东半岛沿岸岛屿上的13号蝮蛇标本(7♂♂,6♀♀)外部形态的详细观察,并与同属其他各种比较,均有不同,应为亚洲蝮属1新种.新种的鉴别特征:1)中段背鳞23行;2)腹鳞+尾下鳞187～198,平均191行;3)体棕褐色,背部有两行粗大、周围暗棕色而中心色浅、外侧开放的圆斑,左右圆斑彼此交错或并列;4)眶后有一较宽黑色眉纹,其下缘略呈波状且镶有一条明显的白色细纹.     

Specialization of bone structure in Pachyvaranus crassispondylus Arambourg, 1952, an aquatic squamate from the Late Cretaceous of the southern Tethyan margin

The histological organization of the vertebrae of the Maastrichtian squamate Pachyvaranus crassispondylus Arambourg, 1952 , was compared to that of various extant squamates, in order to further document the causes and functional consequences of the so-called 'pachyostosis', frequently observed in Late Cretaceous squamates. The vertebrae of Pachyvaranus are composed of the same basic bone tissue types as those of extant lizards and snakes. In particular, periosteal cortices are made of a pseudolamellar (or 'parallel-fibred') tissue, with radial vascular canals, Sharpey's fibres and conspicuous cyclic growth marks that are strictly identical to that found in extant varanids. Conversely, the vertebrae of Pachyvaranus are extremely compact, whereas those of extant squamates are very cancellous and lightly built. This difference is due to the absence in Pachyvaranus of a broad internal resorption field that, in extant lizards and snakes, transforms compact cortices into loose spongy formations. This absence of inner bone resorption typically corresponds to an osteosclerotic process. In Pachyvaranus , cortical hyperplasy, or pachyostosis stricto sensu , was restricted to the walls of the neural spine. Extreme vertebral porosity is likely to be a primitive condition in squamates, because all lizards and snakes examined in this study display this feature. Therefore, the high vertebral compactness observed in Pachyvaranus would be a derived condition arising from the loss (or de-differentiation) of a morphogenetic process: the broad internal resorption of the vertebrae. Possible palaeoecological bearings of these results are discussed.     

白垩纪中期克钦琥珀毛蚊科(双翅目: 毛蚊总科)昆虫一新属新种

毛蚊科是双翅目长角亚目的一类昆虫,其种类丰富,广布世界各地.毛蚊化石记录较为丰富,但往往都是保存于湖相沉积地层中,琥珀中的化石较为罕见.本文根据缅甸北部克钦地区产出的白垩纪中期琥珀中保存的昆虫标本建立了 1新属新种——缅甸白垩叉毛蚊(Cretpenthetria burmensis gen.et sp.nov.),归入...     

Hemiclitoris discovered: a fully differentiated erectile structure in female monitor lizards (Varanus spp.) (Reptilia: Varanidae)

In female monitor lizards ( Varanus spp.), the ubiquitous presence of paired evertible and erectile structures in the ventral portion of the tail root, which are miniaturized mirror images of the hemipenes of the males, was discovered. These organs contain all the structural elements (including a forked retractor muscle, supporting ossifications, epidermal flounces etc.) that are also characteristic for the hemipenes of the respective conspecific males. Consequently, it is proposed that the term hemiclitoris be used for this organ. A preliminary survey showed this organ to be present in several other squamate families, suggesting that it is ubiquitous in all female Squamata.

Zusammenfassung

Die Entdeckung der Hemiclitoris, eines differenzierten erektilen und ausstülpbaren Organs bei Waranweibchen (Reptilia: Varanidae)
In der Schwanzwurzel weiblicher Warane ( Varanus spp.) wurde das obligate Vorhandensein paariger, ausstülpbarer und erektiler Strukturen entdeckt, welche verkleinerte Spiegelbilder der Hemipenes der Männchen darstellen. Diese Organe enthalten sämtliche Strukturelemente (z.B. den gegabelten Rückziehmuskel, Stützknochen, epidermale Paryphasmen), die auch für die Hemipenes der betreffenden artgleichen Männchen charakteristisch sind. Ein erster Erfassungsansatz (mit ausführlicher Artenliste in einer demnächst zu publizierenden, derzeit erstellten Diplomarbeit) zeigt, daß dieses Organ auch bei diversen weiteren Squamatenfamilien nachweisbar ist. Dies legt nahe, daß es tatsächlich obligat bei alien weiblichen Squamata ausgebildet ist.     

Miocene skinks and geckos reveal long-term conservatism of New Zealand's lizard fauna

The New Zealand (NZ) lizard fossil record is currently limited to late Quaternary remains of modern taxa. The St Bathans Fauna (early Miocene, southern South Island) extends this record to 19–16 million years ago (Myr ago). Skull and postcranial elements are similar to extant Oligosoma (Lygosominae) skinks and Hoplodactylus (Diplodactylinae) geckos. There is no evidence of other squamate groups. These fossils, along with coeval sphenodontines, demonstrate a long conservative history for the NZ lepidosaurian fauna, provide new molecular clock calibrations and contradict inferences of a very recent (less than 8 Myr ago) arrival of skinks in NZ.     

The classification of the Lepidosauria

Recent work on modern and fossil genera, and new approaches to classification, have rendered older classifications of lepidosaurs inadequate. This paper discusses the history of lepidosaurian classification from 1860 to the present, and then presents a new hypothesis of lepidosaurian relationships based on phylogenetic analysis. It offers a new definition of the Lepidosauria based on derived character states. Under this definition, squamates, sphenodontids and Gephyrosaurus are lepidosaurs; kuehneosaurs and younginiforms are not. The younginiforms are related to lepidosaurs and may be included within the Lepidosauromorpha. The taxonomic position of kuehneosaurs remains uncertain.     

Hemiclitoris discovered: a fully differentiated erectile structure in female monitor lizards (Varanus spp.) (Reptilia: Varanidae)

In female monitor lizards (Varanus spp.), the ubiquitous presence of paired evertible and erectile structures in the ventral portion of the tail root, which are miniaturized mirror images of the hemipenes of the males, was discovered. These organs contain all the structural elements (including a forked retractor muscle, supporting ossifications, epidermal flounces etc.) that are also characteristic for the hemipenes of the respective conspecific males. Consequently, it is proposed that the term hemiclitoris be used for this organ. A preliminary survey showed this organ to be present in several other squamate families, suggesting that it is ubiquitous in all female Squamata.     

The naso-frontal joint in snakes as revealed by high-resolution X-ray computed tomography of intact and complete skulls

The naso-frontal joint of snakes is described on the basis of high-resolution X-ray computed tomography scans of single individuals of spirit-preserved snake specimens. The suspension of the snout unit from the braincase at the naso-frontal joint shows some broad evolutionary trends among snakes with potential phylogenetic implications, such as sutured or fused medial frontal flanges formed by the medial frontal pillars and the frontal subolfactory processes (in alethinophidians), the restriction of the usually extended dorsoventral contact of the medial nasal flange with the medial frontal flanges to a dorsal or ventral contact (in macrostomatans), and the transfer of the main element of snout suspension from the nasal to the septomaxilla (in colubroids). Some phylogenetic implications of the morphological characters identified in this study are outlined and discussed.     

Amphibians and squamates from the Neogene Siwalik beds of Jammu and Kashmir, India

The Neogene Siwalik deposits of Jammu Province (India) have yielded amphibians and squamates. The collection includes the first amphibians and the first colubroid snakes from the Siwalik Group. Amphibians comprise only anurans: a possible Ranidae and one, or perhaps two, non-ranid frogs Squamates include one lizard,Varanus sp. (Varanidae), whereas snakes are represented by three taxa:Acrochordus dehmi (Acrochordidae), an indeterminate Colubridae, and a snake that is either a Colubridae or an Elapidae.Varanus sp. andA. dehmi have been yielded by the Upper Miocene Ramnagar Member, whereas the anurans and colubroid snakes come from the Upper Pliocene Labli Member. These taxa are indicative of aquatic palaeoenvironment.     

Comparative biology and the importance of cladistic classification: a case study from the sensory biology of squamate reptiles

Evolutionary taxonomy has all but succumbed to cladistic methodology, but it continues to exert considerable influence in the realm of higher classification. Some systematists accept cladistic methods in phylogeny inference, but allow paraphyly in formal classifications. Most important, however, many traditional classifications based on paraphyletic groups (e.g. 'Reptilia') remain in force, deeply entrenched in the literature. Cladists have argued that such paraphyletic classifications can mislead comparative biologists into false evolutionary generalizations, but this assertion has rarely, if ever, been supported by example. This paper provides a case study, illustrating in detail the influence of a traditional paraphyletic classification of squamate reptiles on the historical development of ideas regarding the evolution of sensory modes (chemoreception vs. vision) in the group. The paraphyletic classification is shown to have led to false generalizations and incorrect conclusions stemming directly from the fact that the classification did not reflect accurately the phylogeny of Squamata, particularly the cladistic relationships of Gekkota. This study provides direct evidence that evolutionary generalization must be rooted in the branching pattern of phylogeny and not the potentially arbitrary categorical ranks of traditional taxonomies. It further supports recent calls for a truly phylogenctic taxonomy that has as its philosophical core the concept of descent.     

A new species of Thecadactylus from Sint Maarten, Lesser Antilles (Reptilia, Squamata, Gekkonidae)

We describe a new species of Thecadactylusfrom the Caribbean island of Sint Maarten. The new species differs from all other species in the genus by having a distinct dorsal pattern of numerous irregular but sharply deliminated black spots and blotches on an otherwise almost patternless background.     

A new mosasauroid (Squamata) from the Late Cretaceous (Turonian) of Morocco

Tethysaurus nopcsai gen. et sp. nov. is described on the basis of both cranial and postcranial material from the Late Cretaceous (Early Turonian) of the Goulmima region, southern Morocco. This new mosasauroid is mainly characterized by a parietal table ending posteriorly in two pointed pegs; jugal with a large ascending ramus; splenial with a large and notched dorsomedial process; surangular exposed medially ventral to the coronoid; large paracotylar and parazygosphenal foramina on vertebrae. A phylogenetic analysis shows that Tethysaurus is the sister-group of Mosasauridae. It fills the gap between the aigialosaurids (mainly Cenomanian) and the mosasaurids (known from the Middle-Late Turonian to the Latest Maastrichtian). To cite this article: N. Bardet et al., C. R. Palevol 2 (2003).     

中国东北马斯特里赫特阶的Wulagasaurus dongi(鸭嘴龙亚科)的重新评估与系统发育分析(英文)

通过对黑龙江省乌拉嘎地区晚白垩世渔亮子组地层发现的董氏乌拉嘎龙(Wulagasaurus dongi)的原有材料和新材料进行对比研究和重新评估,可以认定以前归入乌拉嘎龙的部分骨骼(包括不完整的脑颅骨、上颌骨和肩胛骨)实际上应该属于赖氏龙类,并相应地修订了其鉴定特征。乌拉嘎龙具有同北美地区发现的短冠龙和慈母龙十分相似的骨学特征:长而楔形的前顶点位于颧骨前支的一半背腹高度处;颧骨后支呈扇形;乌喙骨的腹突相对较长且纤细;肱骨三角胸嵴较不发育,适度地向前外方扩展;髂骨髋臼上突的腹缘呈不对称的U形,缺乏一条强烈的嵴连接其后部区域与髋臼后突的背缘。系统发育分析结果显示乌拉嘎龙、短冠龙和慈母龙共同构成了鸭嘴龙亚科的一个基干支系。该支系起源于桑托期的亚洲,可能在中坎帕期之前发生分异并通过白令陆桥向北美地区迁徙辐射。