典型社区家庭消费碳排放特征与影响因素——以北京市为例

家庭消费碳排放是中国碳排放总量的重要组成部分,已成为碳排放增长的主要驱动力,从消费角度研究家庭碳排量特征及影响因素对家庭碳减排和低碳社区建设有重要意义。使用碳排放系数法和消费者生活方式法计算北京市5种典型社区家庭消费月均碳排量,通过最优尺度回归和多重比较分析对不同社区家庭碳排放影响因素进行探究。研究发现：北京市5种社区户均碳排放总量及构成差异显著,影响因素不一致。其中：（1）平房类社区家庭直接碳排量732.26 kgCO₂/月高于其他社区,燃煤取暖是平房社区家庭直接碳排放高的主要因素,单位社区、政策性住房社区和商品房社区家庭直接碳排量较低,约50.00 kgCO₂/月。家庭类型显著影响每个社区家庭直接碳排量,家庭积极参与节能环保活动有利于减少家庭直接碳排放;（2）商品房社区家庭间接碳排量最高,达3879.06 kgCO₂/月,平房类社区家庭最低,间接碳排量仅为商品房社区的1/3,间接碳排放是家庭生活消费碳排放的主体。食品和居住消费产生的间接碳排量较高,老龄化社区家庭医疗保健消费碳排量更高;（3）家庭类型和月总收入对所有社区家庭间接碳排量影响显著,但社区环保工作满意度、社区环境满意度、家庭节能环保活动参与度、耐用品使用年限等因素影响程度存在差异,胡同社区和平房类社区中受教育水平高的家庭产生的间接碳排量更高,需积极灌输环保理念。进一步分析了主要影响因素在不同水平下对应的家庭碳排量差异程度与变化规律,有助于社区管理者识别高碳排家庭,为社区低碳管理提供新思路。     

Sustainable livelihood framework-based indicators for assessing climate change vulnerability and adaptation for Himalayan communities

This study evaluated the climate change vulnerability of Himalayan communities, and their potential to adapt to these changes, through assessing their perceived reactions and counter-actions to climate change. The evaluation was conducted through proposing and testing indices for vulnerability (Climate Vulnerability Index – CVI) and adaptation (Current Adaptive Capacity Index – CACI) based on the assumption that a community is an active dynamic entity and has tremendous capability to address the impacts of climate change through an ability to make adjustments based on perceived experiences. Both CVI and CACI include the five forms of capital leading to sustainable livelihood, i.e. human, natural, financial, social and physical capital, and were assessed for each of these forms of capital based on the IPCC framework of vulnerability assessment and its three dimensions (exposure, sensitivity, adaptive capacity). Data for the analysis were collected from randomly selected households located away from district headquarters (ADH) and near district headquarters (NDH). Each dimension was measured based on associated socio-environment-specific indicators for assessing vulnerability and sustainability at community level. The results showed that ADH households had higher human capital and natural capital vulnerability than NDH households. In contrast, NDH households had higher social capital and financial capital vulnerability than ADH households. Overall, ADH households had greater vulnerability than NDH households.These results improve understanding of the environmental and socio-economic changes affecting rural livelihoods and the measures needed to address their specific vulnerabilities by addressing bottlenecks in education and training facilities for skill up-grading, increasing interaction opportunities through local functions and creating opportunities for income generation and effective market and farm linkages. An attempt was made to reduce the gap between bottom-up understanding and top-down policies by suggesting precautionary and ongoing adaptation practices for the communities studied, leading to effective and efficient addressal of vulnerabilities. Vulnerability in the study context was taken to mean externally driven change leading to disturbance in the human environment that could alter internal and external livelihood settings.     

Stream communities across a rural–urban landscape gradient

Rapid urbanization throughout the world is expected to cause extensive loss of biodiversity in the upcoming decades. Disturbances associated with urbanization frequently operate over multiple spatial scales such that local species extirpations have been attributed both to localized habitat degradation and to regional changes in land use. Urbanization also may shape stream communities by restricting species dispersal within and among stream reaches. In this patch-dynamics view, anthropogenic disturbances and isolation jointly reduce stream biodiversity in urbanizing landscapes. We evaluated predictions of stream invertebrate community composition and abundance based on variation in environmental conditions at five distinct spatial scales: stream habitats, reaches, riparian corridors and watersheds and their spatial location within the larger three-river basin. Despite strong associations between biodiversity loss and human density in this study, local stream habitat and stream reach conditions were poor predictors of community patterns. Instead, local community diversity and abundance were more accurately predicted by riparian vegetation and watershed landscape structure. Spatial coordinates associated with instream distances provided better predictions of stream communities than any of the environmental data sets. Together, results suggest that urbanization in the study region was associated with reduced stream invertebrate diversity through the alteration of landscape vegetation structure and patch connectivity. These findings suggest that maintaining and restoring watershed vegetation corridors in urban landscapes will aid efforts to conserve freshwater biodiversity.     

Seasonal relation of NTFPs and socio-economic indicators to the household income of the forest-fringe communities of Jaldapara National Park

For India's forest-dependent communities, Non-Timber Forest Products (NTFPs) are a major source of income and survival. NTFPs collection and their varieties differ season to season. However, little attention has been given on the income variation according to the seasons and various socio-economic factors influencing it. The present study examined the household income of the forest dwelling communities from NTFPs and their relation to the time spent for its collection in three different seasons i.e., summer, winter and rainy. We interviewed randomly selected families of the forest-dependent communities living in the forest fringe areas of Jaldapara National Park (JNP), West Bengal state of India located in the Eastern Himalayas using a semi-structured questionnaire. The average annual household income of the communities was greatly affected by NTFPs obtained during the rainy season. The majority of the households supplemented their total annual household income through NTFPs (1–20% of total family income annually). In the rainy season, the majority of the respondents (75%) spent 1–10 days for NTFPs collection, the time spent significantly influenced their household income. Furthermore, we found that socio-economic variables such as age, education, and family size have an effect on total annual household income from NTFPs. The other variable also influenced the family income is time spent, in terms of number of days expended in NTFP collection during summer and winter seasons. The study affirms the contribution of NTFPs in family income is strongly determined by the period spent in the collection, which also varies with season and socio-economic variables of the inhabitants.     

Contemporary Land Change Alters Fish Communities in a San Francisco Bay Watershed,California, U.S.A.

Urbanization is one of the leading threats to freshwater biodiversity, and urban regions continue to expand globally. Here we examined the relationship between recent urbanization and shifts in stream fish communities. We sampled fishes at 32 sites in the Alameda Creek Watershed, near San Francisco, California, in 1993–1994 and again in 2009, and we quantified univariate and multivariate changes in fish communities between the sampling periods. Sampling sites were classified into those downstream of a rapidly urbanizing area (“urbanized sites”), and those found in less impacted areas (“low-impacted sites”). We calculated the change from non-urban to urban land cover between 1993 and 2009 at two scales for each site (the total watershed and a 3km buffer zone immediately upstream of each site). Neither the mean relative abundance of native fish nor nonnative species richness changed significantly between the survey periods. However, we observed significant changes in fish community composition (as measured by Bray-Curtis dissimilarity) and a decrease in native species richness between the sampling periods at urbanized sites, but not at low-impacted sites. Moreover, the relative abundance of one native cyprinid (Lavinia symmetricus) decreased at the urbanized sites but not at low-impacted sites. Increased urbanization was associated with changes in the fish community, and this relationship was strongest at the smaller (3km buffer) scale. Our results suggest that ongoing land change alters fish communities and that contemporary resurveys are an important tool for examining how freshwater taxa are responding to recent environmental change.     

A method for measuring depth-integrated community metabolism in experimental planktonic–benthic ecosystems

We developed incubation chambers of clear acrylic extending the full depth of the water column to measure in situ depth-integrated community metabolism in experimental planktonic–benthic ecosystems. A magnetic stirring apparatus at mid-depth in each chamber prevented stratification and maintained a vertical turnover time that roughly matched turnover in the experimental ecosystems. We used the chambers in light–dark experiments to partition the depth-integrated photosynthesis and respiration occurring in the experimental ecosystems among water column, wall periphyton, and benthic communities. Community metabolism measured with the chambers was positively related to the chlorophyll-a concentrations of the water column and wall periphyton communities. We also found close agreement between the summation of partitioned community metabolism measured in the chambers and an independent measure of total ecosystem metabolism in the experimental ecosystems. Incubations in chambers and in light–dark bottles revealed similar patterns of water column and wall periphyton community metabolism, but magnitudes differed. In addition to application in experimental ecosystems, this incubation chamber technique may provide an efficient and realistic measure of water column and benthic metabolism and nutrient uptake in lakes and estuaries.     

Transaction costs of community-based forest management: empirical evidence from Tanzania

Levels of transaction costs in community‐based forest management (CBFM) in four communities adjacent to the Ambangulu mountain forests of the north‐east of Tanzania were assessed through questionnaire responses from 120 households. Costs and benefits of CBFM to the rich, medium and poor groups of forest users were estimated. Costs of CBFM were participation in forest monitoring and time spent in meetings. Benefits included forest products consumed at household level. Transaction costs relative to benefits for CBFM were found to be higher for poorer households compared with medium income and richer households. Higher income groups obtained the most net benefits followed by medium and poorer households. Community involvement in forest management may lower the transaction costs incurred by government, but a large proportion of these costs are borne by poorer members of the community. Transaction costs are critical factors in the success or failure of CBFM and need to be incorporated into policies and legislation related to community‐based natural resource management.     

Combined monitoring of changes in δ13CH4 and archaeal community structure during mesophilic methanization of municipal solid waste

Reconstituted municipal solid waste (MSW) with varying contents of putrescible and cellulosic waste was incubated anaerobically under mesophilic conditions. Standard physicochemical parameters were monitored, together with stable isotopic signatures of produced CH₄ and CO₂. δ¹³C values for CH₄ indicated a change of methanogenic metabolism with time. CH₄ was predominantly produced from H₂/CO₂ at the beginning of the incubations. This period was associated with important shifts in archaeal communities monitored by automated ribosomal intergenic spacer analysis (ARISA) and FISH of oligonucleotidic probes targeting specifically 16S rRNA gene of various methanogenic groups. The onset of the active methane generation phase was characterized by an increase of CH₄δ¹³C, indicating a progressive shift toward an aceticlastic metabolism. When the methane production levelled off, a decrease in the isotopic signature was observed toward values characteristics of hydrogenotrophic metabolism. ARISA profiles were, however, found to be stable from the beginning of the active methane generation phase until the end of the experiment. FISH observation indicated that members of the family Methanosarcinaceae were predominant in the archaeal community during this period, suggesting that these methanogens might exhibit a high metabolic versatility during methanization of waste.     

Urbanization within a Dynamic Environment: Modeling Bronze Age Communities in Upper Mesopotamia

Here agent-based models are employed to explore the behavior of small communities and their associated economic systems within an area of climatically marginal environment in northern Syria and Iraq. The examples, drawn from late prehistoric and Bronze Age societies that developed at the onset of urbanization, show how small communities might respond to various resource stresses and environmental fluctuations. The computer simulations demonstrate that some households gain resources at the expense of others and the community becomes more economically differentiated through time with some households benefiting at the expense of others. The approaches discussed demonstrate that complex societies were capable of a wide range of responses to stresses that could be absorbed or amplified via a number of social, economic, or demographic processes.     

The Burden of Non-Communicable Disease in Transition Communities in an Asian Megacity: Baseline Findings from a Cohort Study in Karachi,Pakistan

Background

The demographic transition in South Asia coupled with unplanned urbanization and lifestyle changes are increasing the burden of non-communicable disease (NCD) where infectious diseases are still highly prevalent. The true magnitude and impact of this double burden of disease, although predicted to be immense, is largely unknown due to the absence of recent, population-based longitudinal data. The present study was designed as a unique ‘Framingham-like’ Pakistan cohort with the objective of measuring the prevalence and risk factors for hypertension, obesity, diabetes, coronary artery disease and hepatitis B and C infection in a multi-ethnic, middle to low income population of Karachi, Pakistan.

Methods

We selected two administrative areas from a private charitable hospital’s catchment population for enrolment of a random selection of cohort households in Karachi, Pakistan. A baseline survey measured the prevalence and risk factors for hypertension, obesity, diabetes, coronary artery disease and hepatitis B and C infection.

Results

Six hundred and sixty-seven households were enrolled between March 2010 and August 2011. A majority of households lived in permanent structures (85%) with access to basic utilities (77%) and sanitation facilities (98%) but limited access to clean drinking water (68%). Households had high ownership of communication technologies in the form of cable television (69%) and mobile phones (83%). Risk factors for NCD, such as tobacco use (45%), overweight (20%), abdominal obesity (53%), hypertension (18%), diabetes (8%) and pre-diabetes (40%) were high. At the same time, infectious diseases such as hepatitis B (24%) and hepatitis C (8%) were prevalent in this population.

Conclusion

Our findings highlight the need to monitor risk factors and disease trends through longitudinal research in high-burden transition communities in the context of rapid urbanization and changing lifestyles. They also demonstrate the urgency of public health intervention programs tailored for these transition communities.     

Child nutrition in the transition from nomadic pastoralism to settled lifestyles: individual, household, and community-level factors

The research reported here examines child nutrition in a population that is currently experiencing a transition in subsistence, shifting from nomadic pastoralism to a variety of settled lifestyles. We investigate the range of nutritional consequences of settlement both within and between communities by examining individual, household, and community-level predictors of child nutritional status. Data are drawn from the Rendille Demographic and Health Survey, which contains anthropometric data from 1,088 children ages 6 months to 10 years, as well as socioeconomic data from 640 households drawn from one nomadic and four economically and ecologically diverse settled communities. Comparisons allow us to test the widely held assumption that settlement results in nutritional improvements. The examination of individual and household-level factors highlights several important influences on child nutrition. We find a complex interaction between gender and birthorder, whereby firstborn sons have significantly higher weight-for-height scores than other children, potentially reflecting preference under a system of primogeniture. We also find a complex interaction between defacto female-headed households, where males are absent for over half of the year, and economic status. Young children from poor female-headed households have significantly lower weight-for-height than other children, possibly due to the fact that these households are amongst the poorest in the entire community. However, young children from economically sufficient female-headed households actually fare better than their counterparts in male headed households, suggesting that in households with any discretionary resources, female heads prioritize food acquisition relative to other concerns. Finally, our comparison of child nutritional status across communities, while controlling for individual and household-level variation, does not support the contention that settlement is associated with nutritional improvements. Rather, the effect of community, and its associated changes in subsistence and lifestyle, results in either no nutritional changes, or in the case of young children in the lowland desert community of Korr, diminished nutritional status. Our results underscore the importance of considering variation in sample composition and socioeconomic status when performing community comparisons, and highlight the central role of women in influencing the nutritional welfare of their families.     

Changhsingian brachiopod communities along a marine depth gradient in South China and their ecological significance in the end-Permian mass extinction

Diversity indices (dominance and evenness) and ecological spatial structure (lifestyles and relative abundances) are important features of Changhsingian brachiopod communities prior to the end-Permian mass extinction (EPME) and could predict temporal and spatial extinction patterns during the EPME. In South China, Changhsingian brachiopod communities show higher diversity than other contemporaneous brachiopod communities in the world and have been reported from a variety of sedimentary environments. In this paper, brachiopods from 18 sections in South China were selected to divide communities and compare their ecological structure. Based on the results of network analysis, cluster analysis and quantitative data from the selected sections, we show that Changhsingian brachiopod communities in South China can be categorized into three assemblages along a marine depth gradient: the Neochonetes–Fusichonetes–Paryphella Assemblage from the shallow-water clastic-rock facies, Spinomarginifera–Peltichia–Oldhamina Assemblage from the shallow-water carbonate platform facies and Fusichonetes–Crurithyris Assemblage from the deep-water siliciclastic intracontinental basin facies. Compared with communities from carbonate platform facies, the communities from siliciclastic facies were characterized by high dominance, low evenness and low lifestyle diversity, which might be important biotic factors leading to earlier extinctions. After the extinction began in all environments, the whole earliest Triassic brachiopod community was first dominated by Fusichonetes and then by Crurithyris. These patterns of domination and replacement could be explained by morphological and ecological advantages. The domination of these two genera, which were already adapted to the oxygen and food-limited deep-water habitat, indicates that the cooler deep-water environment might have been a relatively less stressed habitat after the beginning of the EPME. This suggests that global warming might be the main trigger among the previously proposed synergistic environmental stresses, while anoxia might not, at least for the beginning of EPME.     

基于碳排放约束的北京市社区生态效率评价

在“双碳目标”的背景下,社区作为生态文明建设的基本单元,其碳排放核算及生态效率评估尤为重要。基于家庭消费视角,采用碳排放系数法和消费者生活方式法核算社区居民碳排放,并运用超效率DEA模型评估社区生态效率,最后通过计量分析识别生态效率影响因素。研究结果显示：(1)社区居民人均碳排放的评估结果呈现出商品房社区(528.85kgCO₂/月)>单位社区(475.66kgCO₂/月)>传统社区(368.33kgCO₂/月)的关系;(2)生态效率的评估结果呈现出传统社区(111.04%)>商品房社区(108.78%)>单位社区(105.70%)的关系;(3)社区管理工作参与意愿、家庭光照条件会对社区生态效率产生积极影响;此外,年龄结构(中青年人口占比)对商品房社区的生态效率有消极影响,受教育程度的提升则会提高生态效率。进一步提出针对三类社区特点的转型发展策略,为全国不同类型社区生态转型和绿色低碳发展提供参考范式和决策支持。     

Biotic homogenization of three insect groups due to urbanization

Cities are growing rapidly, thereby expected to cause a large‐scale global biotic homogenization. Evidence for the homogenization hypothesis is mostly derived from plants and birds, whereas arthropods have so far been neglected. Here, I tested the homogenization hypothesis with three insect indicator groups, namely true bugs, leafhoppers, and beetles. In particular, I was interested whether insect species community composition differs between urban and rural areas, whether they are more similar between cities than between rural areas, and whether the found pattern is explained by true species turnover, species diversity gradients and geographic distance, by non‐native or specialist species, respectively. I analyzed insect species communities sampled on birch trees in a total of six Swiss cities and six rural areas nearby. In all indicator groups, urban and rural community composition was significantly dissimilar due to native species turnover. Further, for bug and leafhopper communities, I found evidence for large‐scale homogenization due to urbanization, which was driven by reduced species turnover of specialist species in cities. Species turnover of beetle communities was similar between cities and rural areas. Interestingly, when specialist species of beetles were excluded from the analyses, cities were more dissimilar than rural areas, suggesting biotic differentiation of beetle communities in cities. Non‐native species did not affect species turnover of the insect groups. However, given non‐native arthropod species are increasing rapidly, their homogenizing effect might be detected more often in future. Overall, the results show that urbanization has a negative large‐scale impact on the diversity specialist species of the investigated insect groups. Specific measures in cities targeted at increasing the persistence of specialist species typical for the respective biogeographic region could help to stop the loss of biodiversity.     

快速城市化地区景观格局梯度动态及其城乡融合区特征——以广州市番禺区为例

以快速城市化的广州市番禺区为研究样区,通过解译1990-2008年4期Landsat TM影像得到研究区土地利用/覆被数据.选取景观类型水平和景观水平上的景观指数,采用景观格局与梯度分析相结合的方法,研究快速城市化地区景观格局的梯度变化及其城乡融合区特征.结果表明：研究区景观格局动态变化显著,在城市化缓冲带上具有典型的城乡融合区特征,城市景观逐年增加且向外围扩展,而农业景观破碎化严重.斑块密度、蔓延度、景观多样性等指数在城市化梯度上表现出明显的规律性,景观指数峰值出现在距离城市中心4～6 km的梯度带上.时间序列上的景观格局数据体现了不同梯度带上景观动态的差异,城乡融合区景观格局具有斑块形状复杂、景观多样性和破碎化程度高、景观动态变化剧烈的特点.景观指数的峰值逐年向外推移,城乡融合区有逐渐向外发展的趋势.探讨了城乡融合区景观格局时空演化的一般特征及其驱动因素,对区域土地利用政策和可持续发展规划具有指导意义.     

CO2 emissions and urbanization correlation in China based on threshold analysis

Urbanization and CO₂ emissions trends are driven by worldwide economic development. Studies indicate a direct correlation between urbanization and CO₂ emissions increases with both stimulating and inhibiting factors exhibited in the urbanization process and with periodic and regional characteristics exhibited with CO₂ emission. The relationship between urbanization and CO₂ emissions specifically in China from 1979 to 2013 is researched in this paper The novel points of this paper lies in utilizing a threshold mode to test periodic characteristics and analysis by regions. Results verify the effect of urbanization on CO₂ emissions as: (1) emissions increase when threshold point 0.43 was surpassed; (2) emissions increase as residential income increases; (3) coefficients of urbanization on emissions increase initially and then decrease as a factor of increasing industry percentage in overall GDP; (4) patterns of threshold points vary geographically.     

Time for a change: dynamic urban ecology

Contemporary cities are expanding rapidly in a spatially complex, non-linear manner. However, this form of expansion is rarely taken into account in the way that urbanization is classically assessed in ecological studies. An explicit consideration of the temporal dynamics, although frequently missing, is crucial in order to understand the effects of urbanization on biodiversity and ecosystem functioning in rapidly urbanizing landscapes. In particular, a temporal perspective highlights the importance of land-use legacies and transient dynamics in the response of biodiversity to environmental change. Here, we outline the essential elements of an emerging framework for urban ecology that incorporates the characteristics of contemporary urbanization and thus empowers ecologists to understand and intervene in the planning and management of cities.     

城市景观格局视角下空气微生物研究进展

城市化进程将原有自然生态系统改造为以不透水面为主的人工景观,这种变化影响了空气微生物群落的生存环境及其时空异质性。空气微生物受城市化的影响程度与其生态功能的发挥关系密切,微生物群落特征的改变会在一定程度上影响当地生态环境质量并给人群健康带来潜在威胁。城市化进程和空气微生物群落动态分属两个时空尺度差异巨大的生态过程,二者的联合分析已成为目前生态安全与环境健康领域的研究热点。从空气微生物的来源及其组成特征、空气微生物群落的时空异质性及其影响因素以及空气微生物的生态环境效应三个方面系统梳理和总结了近年来探索城市化和空气微生物群落动态之间关系的研究,从宏观视角探讨了当前空气微生物研究的不足,并引入社会-经济-自然复合生态系统、景观格局与过程等相关理论和方法来分析城市化对空气微生物群落特征的影响,旨在明确城市景观格局作用下的空气微生物群落对人群健康的潜在威胁程度,为快速城市化过程中的人居环境的改善提供参考。     

Exploring the community phylogenetic structure along the slope aspect of subalpine meadows in the eastern Qinghai–Tibetan Plateau,China

Exploring the community assembly has been important for explaining the maintenance mechanisms of biodiversity and species coexistence, in that it is a central issue in community ecology. Here, we examined patterns of the community phylogenetic structure of the subalpine meadow plant community along the slope gradient in the Qinghai–Tibetan Plateau of China. We surveyed all species and constructed the phylogenetic tree of the plant community based on data from the Angiosperm Phylogeny Group III. We selected the net relative index (NRI) and evaluated the community phylogenetic structure along the five slope plants communities. We found that the phylogenetic structure varied from phylogenetic clustering to phylogenetic overdispersion with the slope aspect from north to south. In the north slope, the community phylogenetically cluster indicated that the limiting similarity played a leading role in the community assembly and the maintenance of biodiversity. Community phylogenetic overdispersion in the east, southeast, and south slopes indicated that habitat filtration was the driving force for community assembly. The NRI index of the northeast slope was close to zero, implying random dispersion. But it may be driven by the neutral process or limiting similarity, in that the community assembly process was the result of a combination of several ecological factors and thus required further study.     

陕西子午岭生态因素对植物群落的影响

为探讨生境对植被格局分布的影响,对黄土高原马栏林区60块样地进行植被学调查的基础上,采用17个环境指标刻画植物群落的空间位置、地形和土壤特征;利用双向指示种分析(TWINSPAN)划分了该区不同演替阶段的植物群落类型;利用前向选择法(forward selection)及Monte Carlo检验对不同演替阶段植物群落物种组成影响显著(p＜0.05)的环境因子进行筛选;利用去势典范对应分析(DCCA)排序方法分析不同演替阶段植物群落分布格局与环境的关系;利用偏典范对应分析(partial CCA)定量分离环境、空间及其交互作用对植被格局总体变异的影响.结果表明:(1)马栏林区的植物群落可划分为13个类型,分别属于4个不同的演替阶段;(2)对演替初期阶段群落影响显著的环境因子是土壤含盐量和碱解氮,对演替过渡阶段群落影响显著的环境因子是海拔和腐殖质厚度,对演替亚顶级阶段群落影响显著的因子是海拔、坡向、枯落物厚度、腐殖质厚度和pH值,而对演替顶级阶段群落分异影响显著的因子是海拔、坡向、pH值和速效磷;(3)不同演替阶段群落的生态学特性和分布规律与环境空间的生态梯度格局吻合较好;(4)随着演替的进行,环境因子单独对植物群落的影响越来越大,而样地位置单独作用和样地位置与环境因子的交互作用之和越来越小.