A new enantiornithine bird from Upper Cretaceous non-marine deposits at Villespassans (Hérault,southern France)

A coracoid of an enantiornithine bird from Upper Cretaceous (probably late Campanian) fluvial sediments at Castigno (Villespassans, Hérault, southern France) is described. It differs from all hitherto reported enantiornithine coracoids and is referred a new genus and species, Castignovolucris sebei. This bone is large and robust, indicating a bird that was among the largest known enantiornithines, possibly the size of a Canada Goose (Branta canadensis). The new taxon is an addition to the short list of Late Cretaceous birds from France and confirms that enantiornithines were an important component of European avifaunas until late in the Cretaceous.     

Lindholmemydid turtles (Cryptodira: Testudinoidea) from the Late Cretaceous of Shandong Province,China

A new genus and new species of lindholmemydid turtle (Cryptodira: Testudinoidea), Shandongemys dongwuica n. g. and n. sp. are described on the basis of a partial skeleton with incomplete shell and skull, complete lower jaws and disarticulated limb bones from the Upper Cretaceous Wangshi Group of Zhucheng, Shandong Province, China. Among Lindholmemydidae, the new species is closely related to Mongolemys elegans from the Late Cretaceous of Mongolia. An incomplete shell from the same locality is referred as Lindholmemydidae indet. Glyptops sp. from the Upper Cretaceous Wang Group of Jingangkou, Laiyang, Shandong is revised and assigned to Lindholmemydidae.     

First record of the Late Cretaceous giant bird Gargantuavis philoinos from the Iberian Peninsula

Gargantuavis philoinos is a large terrestrial bird, initially described from several bones (synsacrum, cervical vertebra, femora) discovered in the Late Cretaceous deposits of southern France. The synsacrum described here comes from the late Campanian (Late Cretaceous) site of Laño, in north-western Spain, and is similar in all respects to the other synsacra of Gargantuavis philoinos described from France. This is therefore the first specimen attributed to this species to be reported outside France. This find increases the diversity of the fossil assemblage from Laño and the geographical distribution of this bird, which existed on the Ibero-Armorican island, which included southern France and the Iberian Peninsula, during the Late Cretaceous.     

Euclastes acutirostris,a new species of littoral turtle (Cryptodira,Cheloniidae) from the Palaeocene phosphates of Morocco (Oulad Abdoun Basin,Danian-Thanetian)

A new species of the littoral cheloniid turtle Euclastes, E. acutirostris, is proposed, on the basis of a skull from the Palaeocene Phosphates of Morocco, the first turtle record from the Sidi Chennane area. It is estimated to be Danian-Thanetian in age, possibly younger than the previous Danian Moroccan specimens of Euclastes. It differs from the other species of Euclastes mainly by a more elongated and narrower snout, forming a small hook, the presence of a long and narrow spur-shaped postero-inferior process of the jugal, better delimiting the lateral skull emargination and the medially shorter palate, in relation to American Palaeogene specimens. This study indicates the necessity for a world-wide revision of the “Euclastes group” in order to redefine the taxa. It shows the potential interest of the group in the radiation and dispersion of the faunas of the Tethysian and Atlantic margins during the Cretaceous -Tertiary turnover.     

The tiny horned dinosaur Gobiceratops minutus gen. et sp. nov. (Bagaceratopidae, Neoceratopsia) from the Upper Cretaceous of Mongolia

A new horned dinosaur, Gobiceratops minutus gen. et sp. nov. (Bagaceratopidae, Neoceratopsia), from the Upper Cretaceous Baruungoyot Formation of the Khermin Tsav locality (southern Mongolia) is described based on a 3.5-cm-long skull. The nasal included in the orbital border suggests relationship between the new taxon and Bagaceratops rozdestvenskyi. It is proposed that, unlike other neoceratopsian families, the family Bagaceratopidae is of Paleoasiatic origin.     

Craniology of the Plesiosaur <Emphasis Type="Italic">Abyssosaurus nataliae</Emphasis> Berezin (Sauropterygia,Plesiosauria) from the Lower Cretaceous of the Central Russian Platform

Cranial bones of the plesiosaurus Abyssosaurus nataliae Berezin, 2011 from the Upper Hauterivian (Lower Cretaceous) of Chuvashia are described. It differs from other known Cryptoclididae in the evident pedomorphic characters in the skull structure. The skull is strongly hypertrophic; the visceral part is shortened relative to its large occipital region; the orbits are enlarged, the lower jaw is narrow. The jugal is well developed, positioned vertically, and borders most of the posterior orbital wall. This skull structure is evidence of deepwater mode of life. The species diagnosis is revised.     

Zonocypris digitalis (Ostracoda, Crustacea), nouvelle espèce du Fuvélien (Campanien continental) de Provence (sud-est France)

Many specimens of non-marine Ostracodes, especially the new species Zonocypris digitalis were recovered from continental deposits of fuvelian (Campanian) age at Auriol (Provence, south-east France). First species of that genus described in the Cretaceous of southern Europe, Z. digitalis is associated with Charophytes and mainly Gastropodes remains, thus providing confirmation of the datation proposed and age of the studied outcrops. Z. digitalis may be integrated within the current biostratigraphic scale which is accepted as valid for the non-marine Upper Cretaceous of the above mentioned area.     

Ostracodes limniques de la Formation d'Irbzer, Crétacé terminal du Moyen-Atlas, Maroc : taxonomie, biostratigraphie, paléoécologie, paléobiogéographie

An association of limnic ostracodes has been discovered in the upper part of the Irbzer Formation, of Maastrichtian age, in the Moroccan Middle-Atlas. A new species is described, Gomphocythere achloujensis nov. sp. Paracandona occitanica, which is very common in terminal Cretaceous non-marine deposits of southern Europe (France, Spain), is found for the first time on the south Tethyan margin.     

Structure of fossil dinosaur eggshell from the Aix Basin,France

At least four types of dinosaur eggshell are distinguished among samples of fossil eggshell from Late Cretaceous deposits of southern France. Recognition is based on shell microstructure, porosity, and shell thickness. Estimated values of water vapour conductance for these dinosaur eggshells are much greather than predicted for avian eggs, suggesting that the eggs were incubated under conditions of high humidity, such as would occur underground.     

Did the historical range of the European bison (<Emphasis Type="Italic">Bison bonasus</Emphasis> L.) extend further south?—a new finding from the Yenikapı Metro and Marmaray excavation,Turkey

The origin of the European bison (Bison bonasus, Linnaeus, 1758) has been widely discussed and investigated in recent years. The species had a wide historic geographic distribution throughout the European continent during the middle and late Holocene, ranging from France in the west to the Caucasus in the east. However, archaeological evidence is needed to resolve the southern extent of the European bison distribution. We discovered one bison skull fragment during archaeological excavations in 2008 in the area of Yenikap? Metro and Marmaray (Turkey). Radiocarbon dating indicated the skull was deposited during the Byzantine period (seventh to eighth century AD). Mitochondrial genome analyses provided clear evidence that the skull was from a European bison. This is the first unambiguous evidence of the presence of this species in southeastern Europe during Byzantine times, which validates the historical written records of a potentially wider range of the European bison in historical times.     

New Crocodyliforms from Southwestern Europe and Definition of a Diverse Clade of European Late Cretaceous Basal Eusuchians

The late Campanian-early Maastrichtian site of Lo Hueco (Cuenca, Spain) has provided a set of well-preserved crocodyliform skull and lower jaw remains, which are described here and assigned to a new basal eusuchian taxon, Lohuecosuchus megadontos gen. et sp. nov. The reevaluation of a complete skull from the synchronous site of Fox-Amphoux (Department of Var, France) allows us to define a second species of this new genus. Phylogenetic analysis places Lohuecosuchus in a clade exclusively composed by European Late Cretaceous taxa. This new clade, defined here as Allodaposuchidae, is recognized as the sister group of Hylaeochampsidae, also comprised of European Cretaceous forms. Allodaposuchidae and Hylaeochampsidae are grouped in a clade identified as the sister group of Crocodylia, the only crocodyliform lineage that reaches our days. Allodaposuchidae shows a vicariant distribution pattern in the European Late Cretaceous archipelago, with several Ibero-Armorican forms more closely related to each other than with to Romanian Allodaposuchus precedens.     

a new elasmosaurid plesiosaur from the Lower Jurassic of southern France

Plesiosaurus tournemirensis Sciau, Crochet and Mattei, based on a nearly complete skeleton with skull from the Upper Toarcian (Lower Jurassic) of Tournemire (Aveyron Department, southern France), is here redescribed and reinterpreted. Comparisons with other plesiosaurs indicate that it belongs to a new genus, Occitanosaurus . O. tournemirensis is characterized mainly by its spatulate premaxillae with short facial process, very high postorbital broadly contacting posterior ramus of the maxilla, trapezoidal jugal excluded from orbital margin, orbit diagonally oriented, temporal fenestra with a sigmoidal anterior margin, 43 cervical vertebrae, powerful interclavicle-clavicle complex and coracoids with a pointed protuberance on lateral border and expanded posterolateral cornua. Cranial and cervical vertebra features show that this new genus is undoubtedly a representative of the Elasmosauridae. A preliminary cladistic analysis of long-necked plesiosaurs reveals that, within Elasmosauridae, Occitanosaurus is a close relative of Microcleidus and Muraenosaurus .     

A revision of xinjiangchelyid turtles from the Late Jurassic of Sichuan Basin,China

The type specimens of the Jurassic turtles from the Sichuan Basin (China) previously referred to plesiochelyid Plesiochelys are revised, which confirm that they belong to the family Xinjiangchelyidae. The study of a large number of additional shell material shows, as the dominant group, the Xinjiangchelyidae were greatly diversified in the Late Jurassic of the Sichuan Basin. By the absence of mesoplastron and other shell characters, Chengyuchelys baenoides is moved to Xinjiangchelyidae and considered as a valid taxon. Of the xinjiangchelyids from the Late Jurassic of Sichuan Basin, four genera are recognized: Chengyuchelys, Tienfuchelys, Yanduchelys and cf. Protoxinjiangchelys. The phylogenetic analysis results in that Chengyuchelys includes C. baenoides, C. latimarginalis and C. radiplicatus, and Tienfuchelys consists of T. tzuyangensis, T. chungkingensis and T. zigongensis. All xinjiangchelyids from the Sichuan Basin are more primitive than Xinjiangchelys in the plastron sutured to the carapace.     

First radiation of Cretaceous planktonic foraminifera with radially elongate chambers at Angles (Southeastern France) and biostratigraphic implications

The study of planktonic foraminifera from the Lower Cretaceous succession at Angles (Southeastern France) directly correlated with ammonites, confirms that the origin of morphotypes with radially elongated chambers occur earlier than previously recorded. In particular, the species Hedbergella semielongata and Hedbergella roblesae bearing subclavate to clavate chambers first appear in the upper Hauterivian, just predating the onset of the oceanic anoxic Faraoni event. Based on these observations, a new zonation is proposed.     

First Jurassic occurence of Enoploclytia M’Coy, 1849 (Crustacea: Decapoda: Erymidae)

Among the Erymidae Van Straelen, 1925, known as early as the Late Permian (Changhsingian) and widespread in the Jurassic, the genus Enoploclytia M’Coy, 1849 seems to have a late appearance because of its lack before the Cretaceous. Until now, the oldest representative was E. augustobonae Devillez, Charbonnier, Hy?ný and Leroy, 2016 from the Early Cretaceous (Barremian) of the eastern Paris basin (France). However, a new fossil collected in Normandy (France), showing the typical carapace groove pattern of Enoploclytia, attests the presence of the genus in the Late Jurassic (Oxfordian).     

Predation on bellerophontiform molluscs in the Palaeozoic

Shell repair assumed to result from failed predation is documented in 66 specimens of Ordovician-Carboniferous bellerophontiform tergomyan and gastropod molluscs to examine the relationship between the distribution and appearance of injuries, shell morphology and the internal anatomy of the molluscs, as well as the attack strategies of the presumed predators. Furthermore, the distribution of repaired injuries from failed attacks along the apertural margin as a reflection of the nature of the margin and emarginations is investigated. Bellerophontiform molluscs are ideal for this study because of their distinctive isostrophic morphology and the possibility to directly compare broad and narrow conchs with either deep or shallow medial emarginations. The results show that taxa with a deep medial emargination in the form of a slit have significantly more medial injuries than lateral ones. Near-equal frequencies of lateral and medial injuries in specimens with a shallow emargination (slit or sinus) suggest random distribution. Shell form (narrow or broad) does not exert overall control on the distribution of injuries except, perhaps, in some broad explanate shells with an insignificant medial emargination. While this suggests that it is the type of medial emargination that governs distribution of injuries in these forms, it is not clear if this is a result of passive selection due to structural geometry or preferential targeting by predators (i.e. site-specific mode of attack). Predation strategies on bellerophontiform molluscs thus seem to be dependent on the morphological features of the shells rather than their interpretation as tergomyan or gastropod.     

A Ceratopsian Dinosaur from the Lower Cretaceous of Western North America,and the Biogeography of Neoceratopsia

The fossil record for neoceratopsian (horned) dinosaurs in the Lower Cretaceous of North America primarily comprises isolated teeth and postcrania of limited taxonomic resolution, hampering previous efforts to reconstruct the early evolution of this group in North America. An associated cranium and lower jaw from the Cloverly Formation (?middle–late Albian, between 104 and 109 million years old) of southern Montana is designated as the holotype for Aquilops americanus gen. et sp. nov. Aquilops americanus is distinguished by several autapomorphies, including a strongly hooked rostral bone with a midline boss and an elongate and sharply pointed antorbital fossa. The skull in the only known specimen is comparatively small, measuring 84 mm between the tips of the rostral and jugal. The taxon is interpreted as a basal neoceratopsian closely related to Early Cretaceous Asian taxa, such as Liaoceratops and Auroraceratops. Biogeographically, A. americanus probably originated via a dispersal from Asia into North America; the exact route of this dispersal is ambiguous, although a Beringian rather than European route seems more likely in light of the absence of ceratopsians in the Early Cretaceous of Europe. Other amniote clades show similar biogeographic patterns, supporting an intercontinental migratory event between Asia and North America during the late Early Cretaceous. The temporal and geographic distribution of Upper Cretaceous neoceratopsians (leptoceratopsids and ceratopsoids) suggests at least intermittent connections between North America and Asia through the early Late Cretaceous, likely followed by an interval of isolation and finally reconnection during the latest Cretaceous.     

A large iguanodont from the Upper Barremian of the Paris Basin

The remains of an ornithopod dinosaur from the Upper Barremian (Lower Cretaceous) of Auxerre (Burgundy, France) are described. They consist of several vertebrae and a fragmentary scapula. Despite obvious morphological and dimensional affinities with the species Iguanodon bernissartensis, the state of the specimen does not allow any identification more precise than Iguanodontia indet. Reappraisal of the French Upper Barremian record of medium and large-sized ornithopods reveals that no specimen can be definitely determined to the specific level.     

Two sympatric new species of woodlizards (Hoplocercinae, Enyalioides) from Cordillera Azul National Park in northeastern Peru

We report the discovery of two sympatric new species of Enyalioides from a montane rainforest of the Río Huallaga basin in northeastern Peru. Among other characters, the first new species is distinguishable from other Enyalioides by the combination of the following characters: strongly keeled ventral scales, more than 37 longitudinal rows of dorsals in a transverse line between the dorsolateral crests at midbody, low vertebral crest on the neck with vertebrals on neck similar in size to those between hind limbs, projecting scales on body or limbs absent, 96 mm maximum SVL in both sexes, and caudals increasing in size posteriorly within each autotomic segment. The second new species differs from other species of Enyalioides in having strongly keeled ventral scales, scales posterior to the superciliaries forming a longitudinal row of strongly projecting scales across the lateral edge of the skull roof in adults of both sexes, 31 or fewer longitudinal rows of strongly keeled dorsals in a transverse line between the dorsolateral crests at midbody, vertebrals on neck more than five times the size of vertebrals between hind limbs in adult males, projecting scales on body or limbs absent, and caudals increasing in size posteriorly within each autotomic segment. We also present an updated molecular phylogenetic tree of hoplocercines including new samples of Enyalioides rudolfarndti, Enyalioides rubrigularis, both species described in this paper, as well as an updated identification key for species of Hoplocercinae.     

A New Notosuchian from the Late Cretaceous of Brazil and the Phylogeny of Advanced Notosuchians

A new notosuchian crocodyliform from the Late Cretaceous Bauru Group found in the southeastern State of São Paulo (Brazil) is described here. The new taxon, Caipirasuchus stenognathus, is referred as a new species of the recently erected genus Caipirasuchus within the clade Sphagesauridae based on a phylogenetic analysis of basal mesoeucrocodylians. Caipirasuchus stenognathus is represented by an almost complete skull and lower jaw that has autapomorphic characters that distinguish it from other species of Sphagesauridae. These autapomorphies include: maxilla forming part of the orbital margin (absence of lacrimal-jugal contact), nasal with smooth depressions on the posterior region close to the contact with the maxilla and lacrimal, postorbital with posterior palpebral facet that extends posteriorly underneath the ear-flap groove, and a distinct anterior process of the medial flange of the retroarticular process. Additionally, the new taxon lacks autapomorphic features described in other sphagesaurids. The phylogenetic analysis results in a monophyletic genus Caipirasuchus, that is the sister group of a clade fomed by Sphagesaurus huenei, Caryonosuchus pricei, and Armadillosuchus arrudai. Sphagesaurids also include a basal clade formed by Adamantinasuchus navae and Yacarerani boliviensis. Other notosuchian taxa, such as Mariliasuchus amarali, Labidiosuchus amicum, Notosuchus terrestris, and Morrinhosuchus luziae are successive sister taxa of Sphagesauridae, forming a clade of advanced notosuchians that are restricted to the Late Cretaceous of South America. These results contrast with most previous phylogenetic hypotheses of the group that depicted some members of Sphagesauridae as more closely related to baurusuchids, or found Asian (e.g., Chimaerasuchus) or African (Malawisuchus, Pakasuchus) forms nested within advanced notosuchians that are, according to our analysis, endemic of the Late Cretaceous of South America.