A mid-Holocene environmental change in Amazonian savannas

Aim The main goal of this study was to investigate how climate and human activities may have influenced ecotonal areas of disjoint savannas within Brazilian Amazonia. Location Eastern Brazilian Amazonia, Amapá State. Methods The fossil pollen and charcoal records of two lakes in Amapá (Marcio and Tapera) were used to provide a Holocene palaeoecological history of eastern Amazonian savannas. Detrended correspondence analysis was used to enhance the patterns of sample distribution along the sediment core. Results Even though sedimentary hiatuses were recognized in the sediment cores from both lakes, a marked change in vegetation from closed forests with swamp elements to open flooded savanna at c. 5000 yr bp was evident from the pollen record. Charcoal analysis revealed a pattern of increased accumulation of charred particles coincident with the establishment of savanna vegetation, suggesting higher fire frequency near the lakes. Because the timing of the sedimentary hiatus overlapped with the highest Holocene sea level, which would have increased the local water table preventing the lakes from drying out, we infer that both lakes used to depend heavily on flood waters, and the sedimentary gap was caused by reduced discharge from the Amazon River, due to a dry period in the Andes, when precipitation levels markedly decreased between 8000 and 5000 yr bp . The lack of Andean pollen (probably river transported) in the sediment record after this event and the existence of similar records near the study site make this interpretation more appealing. The resumption of sedimentation in Lake Marcio, contemporaneous with falling sea level and increasingly wet conditions in the Andes after 5000 yr bp , indicates that Holocene sea‐level variation did not play an important role in maintaining lake levels. Main conclusions The study site recorded long‐term occupation by pre‐Columbian peoples. However, it is still unclear whether these disjoint savannas have an anthropogenic origin. Even though locally dry environmental conditions were inferred from both records, there is no evidence of a mid‐Holocene dry climate in eastern Amazonia. Instead, the Amapá record indicates a connection between Andean climate and eastern Amazonia, demonstrating the need for a better understanding of the impacts and magnitude of climate changes.     

Influence of rainfall seasonality on African lowland vegetation during the Late Quaternary: pollen evidence from Lake Masoko, Tanzania

Aim  To determine if changes in the key bioclimatic parameters associated with rainfall seasonality can be quantified from fossil sequences and to distinguish them from changes in the total annual rainfall.
Location  Lake Masoko, southern Tanzania (9°20' S, 33°45' E, 840 m a.s.l.).
Methods  Fossil pollen was extracted from a long and well-dated sedimentary sequence spanning the period 45,000–4400 cal. yr bp . The modern habitat, geographical distribution and climatic range (mean annual rainfall and temperature, and length of the dry season) of selected East African plants were used to infer past rainfall attributes for the most characteristic fossil pollen taxa identified from the sediments of Lake Masoko.
Results  In the Masoko pollen sequence it has been possible to identify changes in the length/severity of the dry season during the last 45,000 cal. yr bp , which are interpreted to reflect shifts in the mean position of the Intertropical Convergence Zone (ITCZ) over the southern tropics. We suggest that this bioclimatic parameter has been the main driver of the vegetation dynamics in this area. The major inferred change occurred at the Younger Dryas–Holocene transition c . 11,800 cal. yr bp , when semi-deciduous forest disappeared, being replaced in the lake catchment by woodland, which persists to the present day in spite of locally high rainfall.
Main conclusions  In tropical regions under the influence of the ITCZ, the position and strength of which determine the distribution of rainfall through the year, more attention must be paid to the impact of the length and intensity of the dry season on the modern and past distribution and dynamics of the vegetation. This climatic parameter is as important as the total annual amount of rainfall, and probably one of the most relevant in lowland areas.     

Evidence for a lacustrine faunal refuge in the Larsemann Hills, East Antarctica, during the Last Glacial Maximum

Aim  There is no previous direct evidence for the occurrence of lacustrine refuges for invertebrate fauna in Antarctica spanning the Last Glacial Maximum (LGM). In the absence of verified LGM lacustrine refuges many species are believed to result from Holocene dispersal from sub-Antarctic islands and continents further north. If freshwater lake environments were present throughout the LGM, extant freshwater species may have been associated with Antarctica prior to this glacial period. This study looked at faunal microfossils in a sediment core from an Antarctic freshwater lake. This lake is unusual in that, unlike most Antarctic lakes, the sediment record extends to c . 130,000 yr bp , i.e. prior to the LGM.
Location  Lake Reid, Larsemann Hills, East Antarctica (76°23' E; 69°23' S).
Methods  Palaeofaunal communities in Lake Reid were identified through examination of faunal microfossils in a sediment core that extended to c . 130,000 yr bp .
Results  Ephippia and mandibles from the cladoceran Daphniopsis studeri and loricae of the rotifer Notholca sp. were found at all depths in the sediment, indicating that these two species have been present in the lake for up to 130,000 years. Copepod mandibles were also present in the older section of the core, yet were absent from the most recent sediments, indicating extinction of this species from Lake Reid during the LGM.
Main conclusion  The presence of D. studeri and Notholca sp. microfossils throughout the entire Lake Reid core is the first direct evidence of a glacial lacustrine refugium for invertebrate animals in Antarctica, and indicates the presence of a relict fauna on the Antarctic continent.     

From ice age to modern: a record of landscape change in an Andean cloud forest

Aim To investigate the palaeoecological changes associated with the last ice age, subsequent deglaciation and human occupation of the central Andes. Location Lake Pacucha, Peruvian Andes (13°36′26″ S, 73°19′42″ W; 3095 m elevation). Methods Vegetation assemblages were reconstructed for the last 24 cal. kyr bp (thousand calibrated ¹⁴C years before present), based on pollen analysis of sediments from Lake Pacucha. An age model was established using ¹⁴C accelerator mass spectrometry dates on bulk sediment. Fossil pollen and sedimentological analyses followed standard methodologies. Results Puna brava replaced the Andean forest at the elevation of Lake Pacucha at the Last Glacial Maximum (LGM). Deglaciation proceeded rapidly after 16 cal. kyr bp , and near‐modern vegetation was established by c. 14 cal. kyr bp . The deglacial was marked by the range expansion of forest taxa as grassland taxa receded in importance. The mid‐Holocene was marked by a lowered lake level but relatively unchanged vegetation. Quinoa and maize pollen were found in the latter half of the Holocene. Main conclusions Temperatures were about 7–8 °C colder than present at this site during the LGM. The pattern of vegetation change was suggestive of microrefugial expansion rather than simple upslope migration. The mid‐Holocene droughts were interrupted by rainfall events sufficiently frequent to allow vegetation to survive largely unchanged, despite lowering of the lake level. Human activity at the lake included a 5500‐year history of quinoa cultivation and 3000 years of maize cultivation.     

Late Holocene palaeoecology of Lago Verde: evidence of human impact and climate change in the northern limit of the neotropics during the late formative and classic periods

Multiproxy analysis (pollen, diatom, charcoal) on a 6 m core from Lago Verde (Sierra de Los Tuxtlas), shows evidences of environmental changes and human impact on the evergreen rainforest on the tropical lowlands of eastern Mexico during the last ca. 2,800 years. Prehistoric human occupation is recorded since the late Formative throughout the middle Classic (250 b.c.–ca. a.d. 800) by the presence of maize pollen, a low abundance of tropical arboreal taxa and a high abundance of herbaceous pollen and charcoal particles. This occurred under a scenario of very low lake levels from which dry conditions are inferred based on the dominance of aerophilous and periphytic diatom taxa. After ca. a.d. 800 the site was abandoned and forest regeneration started, at the same time higher lake levels, an indication of more humid conditions, were established. In the absence of human disturbance, tropical forest regeneration was rapid (ca. 200 years). Fluctuations in pollen composition during times of low human population at the site are related to climate variability, with the highest tropical forest diversity and lake levels recorded during the Little Ice Age. Modern human impact is also recorded and compared with the prehistoric deforestation event. Comparison with palaeoecological records from Yucatan and the highlands of central Mexico offers a Mesoamerican perspective of climatic variability giving evidence that the late Formative and early to middle Classic demographic expansion occurred under a scenario of climates dryer than present, with the Postclassic characterized by moister conditions. The end of the Classic (ca. a.d. 800–1000) is identified as a period of rapid climate change which marks one of the most important cultural transitions in Mesoamerica.     

A Holocene environmental record from the southern Yangtze River delta, eastern China

The Holocene sedimentary record of core ZX-1, recovered west of mid-Holocene Chenier ridges on the monsoon-controlled Southern Yangtze delta, eastern China, consists of lagoon, salt marsh, upper tidal flat, and limnic facies, reflecting low-energy depositional environments. Holocene deposits mainly originated from the surrounding Tai Lake drainage basin. The temporal variation of most geochemical element percentages corresponds with the climatic phases inferred from the pollen record, i.e., the relatively low values of SiO₂, Na₂O, CaO and high values of Al₂O₃, K₂O, MgO, Fe₂O₃, FeO, FeO + Fe₂O₃ and TiO₂ generally concur with the warm and humid climate, and vice versa. Three geochemical indices − Al₂O₃/Na₂O, K₂O/Na₂O, and CaO/K₂O − are found to be sensitive to past precipitation in the monsoon-controlled southern Yangtze River delta. Based on fine sediment analysis of core ZX-1, the samples deposited under a dry climate with a mean annual precipitation (MAP) < 500–900 mm tend to have Al₂O₃/Na₂O values less than 12.2, and the samples deposited under a moist climate with an MAP 1000–1800 mm mostly have Al₂O₃/Na₂O values higher than 12.2. Similarly, the K₂O/Na₂O boundary value is 2.0. However, for CaO/K₂O, the dry climate sediments likely show values higher than 0.9, and those wet climate ones generally have values less than 0.9. This geochemical response suggests the potential application of these indices in the interpretation of palaeoclimate variation of monsoon-controlled eastern China.

Holocene climatic variation history is reconstructed for the southern Yangtze delta. From 8000 to 7000 yr BP, regional climate demonstrated frequent fluctuations, with warm and wet periods (8000–7700; 7500–7200 yr BP) alternating with cool and less humid periods (7700–7500; 7200–7000 yr BP). From 7000 to 6000 yr BP, the climate was relatively warm and humid. Since then, it had turned cool and dry, climaxing in an intense cold event around 4000 yr BP. After the cold event, it became warm and humid until around 2500 yr BP.     

Methodological and practical aspects of the presentation and interpretation of microscopic charcoal data from lake sediments

Analysis of microscopic charcoal particles is a useful part of palaeoecological research and is mostly used in conjunction with pollen analysis. However, there is considerable variation in the methodology of charcoal analyses. This paper considers various methods for the analysis of microscopic charcoal and the presentation of charcoal data in the context of a study of the upper sediments of two lakes Estonia. The results are evaluated by comparing both the pollen and charcoal data with documentary evidence of forest fires over the past 60 years. Indications of fires both varying in extent and at different distances from the lakes are evaluated in both the pollen and charcoal diagrams. The results suggest that the total area curve for charcoal particles per unit mass of sediment dry matter (cm² g⁻¹) provides the best indicator of forest fires. Fires in the study area are reflected differently in the charcoal and pollen curves. It is suggested that the charcoal data have the potential to indicate disturbance at a greater distance from the coring site than indicated by the pollen data. Received February 5, 2002 / Accepted August 14, 2002     

Holocene persistence of wooded steppe in the Great Hungarian Plain

Aim We used a combination of new and previously published palaeoecological data to test three hypotheses: (1) that wooded steppe persisted in the Great Hungarian Plain throughout the Holocene; (2) that wooded steppe and steppe were most extensive between c. 9900 and 8300 cal. yr bp (the ‘Boreal steppe’ period); and (3) that Southern Continental, Pontic and Eastern Sub‐Mediterranean steppe species reached the region during the early Holocene via the ‘Lower Danube Corridor’. Location Sarló‐hát oxbow lake, Hungary and the Eastern European wooded steppe zone. Methods Holocene sediments deposited in the Sarló‐hát oxbow lake were subjected to pollen and microcharcoal analyses. Twelve radiocarbon age estimates were obtained to determine sediment chronology. In addition, previously published palaeoecological data from the Great Hungarian Plain were compiled, analysed and compared with previous studies in other regions of steppe and wooded steppe in eastern Europe. Results Palynological data from two sediment cores extending to c. 11,400 cal. yr bp indicate the persistent dominance of the landscape by temperate deciduous wooded steppe throughout the Holocene, although with varying canopy composition. Warm‐continental steppe grasslands and saline tall‐grass meadows developed on edaphically constrained areas, which remained steppe‐dominated throughout the Holocene. The extent of steppe grasslands did not increase between 9900 and 8300 cal. yr bp . After c. 3100 cal. yr bp , anthropogenic activities led to the development of cultural steppe. Thermophilous steppe species of the Southern Continental, Pontic and Sub‐Mediterranean floristic elements probably reached the Great Hungarian Plain principally via the Lower Danube Corridor during the late glacial interstadial and Holocene. Eurythermic members of these elements, however, probably survived the Last Glacial Maximum in favourable microsites, extending their ranges during the Holocene from these local sources. Main conclusions Our results confirm the Holocene persistence of wooded steppe in the Great Hungarian Plain, disprove the ‘Boreal steppe’ theory, and suggest an Early Holocene period of greater vegetation openness between 11,400 and 9900 cal. yr bp . Evidence for the post‐glacial immigration of south‐eastern steppe elements into the Carpathian Basin is equivocal: the last glacial/interglacial presence of several southern steppe species suggests that the Hungarian Plain hosted suitable habitats for them during warm and cold phases alike.     

Evidence for Late Glacial and Holocene environmental changes from subfossil assemblages in sediments of Lake Neuchâtel, Switzerland

Ostracod and diatom assemblages, mollusc remains, plus pollen and sediment stratigraphy from three cores taken at sites 40, 80 and 135 m deep in Lake Neuchâtel, Switzerland, show distinct changes since deglaciation. Shifts are interpreted in terms both of climatic change and major changes in the catchment input caused by the Aar River entering or bypassing Lake Neuchâtel. Three palaeoenvironmental phases are recognized: (1) the Late Glacial, with the inflow of the river before 10,000 yr B.P.; (2) the early to mid-Holocene, with temporary bypassing of the river between 10,000 and 4850 yr B.P.; and (3) the mid- to late Holocene, without river input after approx. 4850 yr B.P. During periods with river input, rhythmites were deposited. Ostracods indicate a well oxygenated hypolimnion, suggesting deep ventilation enhanced by the river inflow. The diatom assemblage indicates eutrophic to mesotrophic conditions and is dominated by generally well preserved periphytic species. The high number of unidentifiable pollen suggests oxidation during transport in the river. During periods without river input, a massive calcareous silt with abundant authigenic calcite was deposited, diatoms indicate a change to mesotrophic and oligotrophic conditions, and decreased sedimentation rates and higher alkalinity enhanced the dissolution of diatoms. Terrestrial and littoral molluscs as well as periphytic diatoms indicate erosion of the shore when the lake level rose or fell with onset or offset of the river inflow. Before 12,600 yr B.P., the patchy vegetation cover led to intensive soil erosion and high sedimentation rates. The lake was still frozen during the winter months. After approx. 12,600 yr B.P. the sediment changed from clastic rhythmites to thinly bedded, non-glacial rhythmites, the vegetation cover became denser, the abundance of ostracods increased, and molluscs started to colonize the profundal zone of the lake. The Younger Dryas (approx. 10,700–10,000 yr B.P.) displays maximum abundance of ostracods, indicating deep ventilation. The transition to the Holocene is characterized by a change in the ostracod assemblage and the river bypassed the lake between approximately 10,000 and 8900 yr B.P. When the river reentered the lake after approximately 8900 yr B.P., a new ostracod species appeared. After two short periods of river bypassing between approximately 7200 and 6950 yr B.P., diatoms indicate a higher trophic level about 6000–5000 yr B.P. associated with higher erosion rates and increased nutrient input due to Neolithic agricultural activities. After approximately 4850 yr B.P. the river bypassed the lake again and the trophic level decreased. In contrast to the early to mid-Holocene periods of decreased oxygen supply, the mid- to late Holocene is characterized by deep ventilation. This is attributed to the northward retreat of the polarfront and enhanced westerlies consequent to decreased insolation after 6000 yr B.P. The change of the river flow pattern correlates with glacier oscillations and may suggest a climatic control. The final cessation of the inflow is attributed to diversion of the river consequent to isostatic uplift and late Neolithic clearance activity.     

Palaeolakes of the south central Sahara — problems of palaeoclimatological interpretation

The results of palaeoecological studies of Holocene swamp and lake deposits of a number of endorheic depressions of the Kawar, Djado and Great Erg of Bilma region of eastern Niger are presented, comprising analysis of their stratigraphy, sedimentology, diatom flora and macrofossils.The investigations demonstrate that various palaeolakes have reacted differently in space and time and by type of lake to climatic conditions. Some of the lakes reacted rapidly to changes in the precipitation regime, as evidenced by changing size, level, water balance and water chemistry, while perennial freshwater lakes nearby show changes relatively independent of short-term climatic fluctuations. These facts suggest a more complex influence of local and regional geomorphological, hydrological and hydrogeological factors on the Holocene lake evolution than a mere climatic dependence. Beyond doubt humidity considerably increased during early and mid-Holocene periods, but the stratigraphical and ecological status obtained for individual endorheic depressions seems to be mainly a reflection of differences among groundwater catchments, aquifers of different size, local topographical conditions and changes in geomorphology (e.g. dune activity) — superimposed on the major climatic tendencies — and thus of a considerable diversity of palaeoenvironmental conditions occurring within the region at any given time of the Holocene.     

Dynamics of North Patagonian rainforests from fine‐resolution pollen,charcoal and tree‐ring analysis,Chonos Archipelago,Southern Chile

Abstract Fine‐resolution palaeoecological and dendrochronological methods were used to investigate the impacts of climate change, and natural and anthropogenic disturbances on vegetation in the North Patagonian rainforest of southern Chile at decadal to century timescales during the late Holocene. A lake sediment mud–water interface core was collected from the northern Chonos Archipelago and analysed for pollen and charcoal. Dendrochronological analysis of tree cores collected from stands of Pilgerodendron uviferum close to the lake site was incorporated into the study. The combined analysis showed that the present mosaic of vegetation types in this region is a function of environmental changes across a range of timescales: millennial climate change, more recent natural and anthropogenic disturbances, and possibly short‐term climatic variations. Of particular interest is the spatiotemporal distribution of Pilgerodendron uviferum dieback/burning in the Chonos Archipelago region.     

A spatially precise study of Holocene fire history, climate and human impact within the Maurienne valley, North French Alps

1 If, within a vegetation type, fire regimes are climate dependent, then fire patterns should be synchronous at regional scales. If they are not synchronous, then fires may be dependent on local processes such as human-induced disturbances.
2 Two fire chronologies were developed using 34-radiocarbon dating measured by accelerator mass spectrometry (AMS) from wood charcoal buried in soil. These charcoal fragments were sampled in two study sites 10 km apart in the Maurienne valley (Southern Vanoise massif, Savoy, France).
3 Asynchronous temporal fire patterns were seen at Aussois and Saint-Michel-de-Maurienne. This demonstrates the dependence of fires on local- or stand-scale environmental forcing; any direct relationship with climate is therefore rejected.
4 Slash-and-burn practices are probably the main source of Holocene fires in the Maurienne valley. However, deforestation did not occur throughout a site in any period, nor simultaneously at the same elevation in two different sites 10 km apart. The cultural landscape was shaped as early as the Neolithic and the Bronze Age, between 6000 and 3000 bp .
5 Deforestation at both study sites probably occurred in many stages in many small areas. The fire intervals were c. 500–1000 years. Deforested areas increased in extent over 2000–4000 years, until the present-day cultural landscape was established. This process stopped c. 2500  bp at Saint-Michel-de-Maurienne but is still active at Aussois.     

根据孢粉分析论青藏高原西部和北部全新世环境变化

通过对青藏高原西部班公湖钻孔和北部中昆仑山３个湖相剖面的孢粉研究,揭示青藏高原西部和北部地区全新世１万年期间植被的演替和气候变化,西部在全新世早期９９００－７８００ｙｒ．Ｂ．Ｐ．植被由荒漠转为草原,气候好转;中期７８００－３５００ｙｒ．Ｂ．Ｐ．,草原发展,气候较适宜,以７２００－６３００ｙｒ．Ｂ．Ｐ．为高温湿期,５５００ｙｒ．Ｂ．Ｐ．和３５００ｙｒ．Ｂ．Ｐ．出现干旱,晚期从３５００ｙｒ．Ｂ．Ｐ．至今植被为荒漠,气候干旱,其中７００ｙｒ．Ｂ．Ｐ．气候恶化。北部地区全新世时期为荒漠植被,当气候温湿时,蒿和禾本科,莎草科成份增加,藜科减少,气候干旱时则相反,北部全新世的气候分期和干湿波动与西部相近,两地在晚期气候朝干旱化发展。     

Classification, assessment and trophic reconstruction of Danish lakes using chironomids

1. Surface-sediment assemblages of subfossil chironomid head capsules from fifty-four primarily shallow and nutrient-rich Danish lakes were analysed using multivariate numerical techniques. The species data, comprising forty-one chironomid taxa, were compared to environmental monitoring data in order to establish a relationship between chironomid faunal composition and lake trophic state.
2. The subfossil assemblages were compared to the chironomid bathymetric distributions along transects from four lakes. Correspondence analysis and similarity coefficients showed that the subfossil assemblages, sampled in the lake centre, reflect the chironomid communities in the littoral at a depth of 2–7 m.
3. Two-way indicator species analysis (TWINSPAN) was used to classify the Danish lakes into five groups defined by trophic state, lake depth and pH. Eighteen chironomid taxa showed significant differences in abundance among the five groups. Canonical correspondence analysis (CCA) showed the chlorophyll a concentration ([Chl a ]) and Secchi depth to be the variables best correlated to the faunal data, and fourteen taxa were significantly correlated to [Chl a ].
4. The strong correlation between chironomid data and the ln-transformed ([Chl a ]) was used to create a weighted averaging (WA) model to infer lake trophic state. Several models were tested by cross validation (leave-one-out jack-knifing), and a simple WA model using inverse de-shrinking had a RMSEP_jack of 0.65 (ln units) and a r ²_jack of 0.67.
5. The results can be used in the assessment and reconstruction of lake trophic state for long-term monitoring and palaeoecological investigations of shallow, temperate lakes in the mesotrophic to hypertrophic nutrient range.     

Holocene vegetation history of the Sahel: pollen, sedimentological and geochemical data from Jikariya Lake, north-eastern Nigeria

Aim This study aims to separate regional and local controls on Holocene vegetation development and examine how well pollen records reflect climate change in a semi‐arid region. The relative importance of climate and human activity as agents of vegetation change in the Sahel during the late Holocene is also considered. Location Jikariya Lake, an inter‐dune depression in the Manga Grasslands of north‐eastern Nigeria. Methods Pollen and charcoal were used to provide a record of Holocene vegetation history. Palaeoclimate and hydrological changes were reconstructed from sedimentary and geochemical data. Regional and local influences were separated by comparing the evidence obtained from Jikariya Lake with previously published data from the Manga Grasslands. Results The Manga Grasslands experienced a prolonged wet period during the early and mid‐Holocene, during which swamp forest vegetation with Guinean affinities (Alchornea, Syzygium, Uapaca) occupied the inter‐dune depressions. However, variation in the pollen records between sites suggests that their establishment was dependent on conditions being locally favourable, rather than being directly coupled to regional climate. The pollen records from the Manga Grasslands are more consistent in suggesting the colonization of the dunefields by trees associated with Sudanian savanna (Combretaceae, Detarium) c. 8700 cal. yr bp . The Jikariya Lake pollen data are in accordance with the sedimentological and geochemical data from the region in indicating that the onset of arid conditions occurred progressively during the late Holocene (from c. 4700 cal. yr bp ). Abrupt changes in pollen stratigraphy, recorded at other Manga Grasslands sites 3500 cal. yr bp , appear to be the product of the local passing of ecological thresholds. The dunefield vegetation (Sahelian savanna) appears to have been resilient to (or at least palynologically silent regarding) to the climatic variability of the late Holocene. Main conclusions While climate appears to have been the primary control on vegetation development in the Manga Grasslands during the Holocene, local conditions (particularly depression size and sand influx) had a strong influence on the timing of pollen stratigraphic changes. Anthropogenic influences are difficult to detect, even during the late Holocene.     

Vegetation history of Lago Battaglia (eastern Gargano coast,Apulia, Italy) during the middle-late Holocene

A pollen and charcoal record from Gargano (southern Italy) provides new information on the vegetation history and environmental change in southern Italy during the middle to late Holocene. The chronological framework is provided by six AMS radiocarbon dates carried out on plant macroremains. Pollen diagrams record a Mediterranean vegetation along the coastland and a stable mixed oak forest at higher elevations between ca. 5900 and 4200 cal b.p. A sharp and dramatic fall of tree pollen concentrations and a change in fire frequencies occurred from approximately 4200 cal b.p. suggesting a climate change towards drier conditions. This event is coherent with regional and extra-regional palaeoclimatic records that suggest that a mid-Holocene dry period was experienced in southern Italy, southern Spain, and perhaps further afield. Human impact on vegetation seems to have occurred since about 2700 cal b.p.     

Long-term forest fire ecology and dynamics in southern Switzerland

1 Pollen and charcoal analysis at two lakes in southern Switzerland revealed that fire has had a prominent role in changing the woodland composition of this area for more than 7000 years.
2 The sediment of Lago di Origlio for the period between 5100 and 3100  bc cal. was sampled continuously with a time interval of about 10 years. Peaks of charcoal particles were significantly correlated with repeated declines in pollen of Abies , Hedera , Tilia , Ulmus , Fraxinus excelsior t., Fagus and Vitis and with increases in Alnus glutinosa t., shrubs (e.g. Corylus , Salix and Sambucus nigra t.) and several herbaceous species. The final disappearance of the lowland Abies alba stands at around 3150  bc cal. may be an example of a fire-caused local extinction of a fire-intolerant species.
3 Forest fires tended to diminish pollen diversity. The charcoal peaks were preceded by pollen types indicating human activity. Charcoal minima occurred during periods of cold humid climate, when fire susceptibility would be reduced.
4 An increase of forest fires at about 2100  bc cal. severely reduced the remaining fire-sensitive plants: the mixed-oak forest was replaced by a fire-tolerant alder–oak forest. The very strong increase of charcoal influx, and the marked presence of anthropogenic indicators, point to principally anthropogenic causes.
5 We suggest that without anthropogenic disturbances Abies alba would still form lowland forests together with various deciduous broadleaved tree taxa.     

Holocene sedimentary history of some coastal plains in Hokkaido,Japan IV. Diatom assemblages in the sediments from Kushiro moor (2)

Diatom assemblages of sediments obtained from three sites on Kushiro Moor were analyzed to investigate the Holocene sedimentary history. The results showed that: 1) The Takkobu site was originally at the bottom of the paleo-Kushiro Bay, and after-wards the paleo-Takkobu Lagoon developed, became sealed off, and changed to a freshwater lake. The succession to peat moor probably began about 2000 yr B.P. at the Takkobu site. 2) The Tsurui site was originally at the bottom of the paleo-Kushiro Bay, then changed to the paleo-Kushiro Lagoon and became peat moor as a result of the first Holocene regression, which finished about 3600 yr B.P. The site then returned to a brackish lake again, probably due to the second Holocene transgression between 3600 and 3000 yr B.P., thereafter passing through brackish lake and freshwater lake stages, and eventually becaming peat moor at about 2000 yr B.P., 3) At the Chuo site, the second paleo-Kushiro Bay developed again as a result of the second Holocene transgression, which finished about 3000 yr B.P. Thereafter, brackish or freshwater lakes, rivers, and then peat moor developed in the central area of Kushiro Moor. 4) The second marine diatom zone (MD₂ Zone), which indicates the second Holocene transgression, complete by about 3000 yr B.P., is detected only at the Chuo site in the central area of Kushiro Moor.     

Late Quaternary grassland (Campos), gallery forest,fire and climate dynamics,studied by pollen,charcoal and multivariate analysis of the São Francisco de Assis core in western Rio Grande do Sul (southern Brazil)

We present a detailed pollen and charcoal record of a 368-cm-long sediment core from the lowland Campos (grassland) region near the city of São Francisco de Assis in the western Rio Grande do Sul State in southern Brazil. Based on four AMS radiocarbon dates, the record represents the last about 22,000 cal yr BP. The region was naturally covered by Campos throughout the recorded glacial and Holocene period under cold and relatively dry and warm and dry condition, respectively. Initial expansion of gallery forest after 5170 cal yr BP indicates a change to wetter climatic conditions. Maximum extent of gallery forest after 1550 cal yr BP reflects the wettest recorded period. There is some evidence of plant migration from the eastern Atlantic coastal lowland reaching the western lowland, first after mid Holocene times by Cecropia and, possibly, species of Myrsine and Moraceae and later, after about 1000 cal yr BP, by species of Alchornea and Acalypha. Multivariate analysis revealed that the long-term pollen composition dynamics is a two-phase process, with random, chaotic changes characterizing some periods when climate conditions were likely more stable, and directional compositional changes (phase transitions) in periods coincide with major climate and/or anthropogenic changes. Natural fires were rare during full- and lateglacial periods, but became frequent at the beginning of the Holocene, suggesting the beginning of human occupation of the western lowland at that time. Highest fire frequency is found during the wet late Holocene period, suggesting an increase of indigenous populations in the São Francisco de Assis region.     

Post-glacial migration of silver fir (Abies alba Mill.) in the south-western Alps

Aim  Previous studies have failed to reconstruct the regional post-glacial migration pattern of Abies alba in southern France. Based on the first exhaustive compilation of palaeoecological data in this region, we present the state-of-the-art and attempt to synthesize the available information concerning glacial refugia and post-glacial migration, and analyse the information with regard to climate and orography.
Location  South-western Alps and adjacent areas, southern France.
Methods  The work compiles the available palaeoecological data in the south-western Alps (52 sites, 290 radiocarbon dates). The post-glacial migration pattern of Abies alba is reconstructed based on 22 selected palynological analyses (11 well-dated reference sites and 11 supplementary ones).
Results  The geographical patterns of approaching area limit, immigration and expansion are reconstructed at the scale of the southern French Alps.
Main conclusions  Despite previous assertions, the evidence of refugia in southern France is non-existent. The late-glacial records of fir pollen, previously interpreted in French Mediterranean regions and on adjacent foot-hills as possibly reflecting regional refugia, most probably correspond to reworking phenomena or long-distance pollen transport. Fir migration, originating in the Apennine refugia and through the south-western extremity of the Alps, was extremely rapid in the southern French Alps, only spanning a few centuries between 10,100 and 9800 cal. yr bp . The subsequent spread of fir populations was controlled by local parameters, such as the aridity of the inner valleys, which resulted in a delayed expansion in comparison to other regions. Abies almost disappeared from the south-western Alps during the Roman era, around 2000 cal. yr bp .