嵩草属地理分布的研究

嵩草属Kobresia Willd.隶属于莎草科,全世界有64种5变种,中国有49种4变种,属下分为4个组。该属主要分布于北半球温带至寒带,亚洲种类最多,主要集中分布于喜马拉雅山地区和横断山地区。上述两地共有总数的90％以上的种类。因此,喜马拉雅-横断山地区为嵩草属的分布中心。与嵩草属最近缘的属Schoenoxiphium只分布在马达加斯加和非洲东南部山地。两个属可能有共同的祖先,发生于冈瓦纳古陆。随着印度板块与非洲大陆分离并向北方漂移,嵩草属的祖先被带到欧亚大陆,在两个板块相遇处——喜马拉雅-横断山地区产生了现在的嵩草属。其后,喜马拉雅山脉进一步抬升,气候与环境发生巨变,嵩草属也进一步分化形成现在的规模。印度板块在早第三纪与欧亚大陆相连接,嵩草属可能就是此时起源于喜马拉雅山地区,并开始分化,且沿北半球的山系向北扩散到欧洲和西伯利亚,又从欧洲到格陵兰再到加拿大东部,从西伯利亚通过白令海峡到阿拉斯加并沿落基山脉南下达到美国的科罗拉多,形成了嵩草属现今的分布格局。     

鹅观草属的地理分布

鹅观草属是禾本科小麦族中的最大的属,现知全世界有4组,20系,126种,分布于北半球的温带和寒带,中国有4组,18系,79种,主要分布于西北,西南,华北和东北,是鹅观草属植物种类最为集中的区域,尤其高原东北部的唐古特地区又是我国鹅观草属分布相对密集之地,有3组,12系,30种,而且其间不同等级,不同演化水平的类群均有分布,该地可能就是该属的现代分布中心,同时,唐古特地区多汇聚有鹅观草属不同等级的原始类群和与原始类群有很缘的短柄草属植物,其中最原始的大柄鹅观草特产于该区,而该区缺乏的是高级的大颖组类群,故推测唐古特地区可能又是该属的起源地,起源时间大约在青藏高原明显增高,气候转凉的晚第三纪初的中新世,鹅观草属起源后,在中国境内地质活动比较剧烈的地区得到了进一步的发展和分化,但只有少数适应性较强的类群大概以3条路径扩展到国外,并向东到在北美的巴芬岛,向西延伸到大西洋滨岸,向北进入寒冻的北极地区。     

蒿草属地理分布的研究

嵩草属Kobresia Willd．隶属于莎草科,全世界有64种5变种,中国有49种4变种,属下分为4个组。该属主要分布于北半球温带至寒带,亚洲种类最多,主要集中分布于喜马拉雅山地区和横断山地区。上述两地共有总数的90%以上的种类。因此,喜马拉雅—横断山地区为嵩草属的分布中心。与嵩草属最近缘的属Schoenoxiphium只分布在马达加斯加和非洲东南部山地。两个属可能有共同的祖先,发生于冈瓦纳古陆。随着印度板块与非洲大陆分离并向北方漂移,嵩草属的祖先被带到欧亚大陆,在两个板块相遇处—喜马拉雅—横断山地区产生了现在的嵩草属。其后,喜马拉雅山脉进一步抬升,气候与环境发生巨变,嵩草属也进一步分化形成现在的规模。印度板块在早第三纪与欧亚大陆相连接,嵩草属可能就是此时起源于喜马拉雅山地区,并开始分化,且沿北半球的山系向北扩散到欧洲和西伯利亚,又从欧洲到格陵兰再到加拿大东部,从西伯利亚通过白令海峡到阿拉斯加并沿落基山脉南下达到美国的科罗拉多,形成了嵩草属现今的分布格局。     

耳草属(HedyotisL.)的起源及地理分布

依据耳草属的系统学,古地理学和细胞染色体资料分析和推论,耳草属植物的起源地点在冈瓦纳古陆,很可能在古陆辽阔的东北部地区。耳草属植物的起源时间不应晚于侏罗纪,有可能在三叠纪就已经出现,并在侏罗纪得到广泛的传播,其迁移的路线有四条,即从起源中心向东经上耳其,伊朗进入东南亚地区,向东南通过古南大陆向印度板块和徜大利亚扩散,向北进入北美地区,向西南进入南美洲地区,随着冈瓦纳古陆的分离,印度板块向北漂移以及澳大利亚与古南大陆的分离,植物迁移与扩散的速度受到了制约,耳草属植物现代分布格局形成的原因在于传播途径的隔断和第四纪冰川的作用。     

以礼草属的地理分布

根据植物类群的地理分布与系统发育相统一的原理,本文讨论了以礼草属的分布中心、起源地、 起源时间和现代分布格局的形成。以礼草属全世界约26种、6变种,隶属于3个组,主要分布于中国,哈萨克斯坦、吉尔吉斯斯坦、塔吉克斯坦、阿富汗和伊朗也有分布。其中,中国的青藏高原汇聚了该属的大多数种类,且不同等级和演化水平的类群均集居于此,使其成为该属的现代分布中心;而该属的原始类群、以及与原始类群很近缘的鹅观草属却分布在这一中心之外的天山地区,加之天山地区自新生代的晚第三纪再次抬升以来,具备了以礼草属发生和繁衍的自然条件,因而天山地区很可能就是该属的起源地,起源时间也可能在晚第三纪或第四纪初。以礼草属自天山起源后,扩散的途径大概有3条,其中西南向途径和东南向途径从东、西两侧侵入青藏高原,在青藏高原得到极度发展,并随着高原的继续隆起,进一步衍生出最高级的类群短穗组,从而形成了以礼草属现今的分布格局。     

新疆赖草属二新种

本文发表了我们在编写《flora of kunlun and karakorum》(禾本科)时发现的两个新种:若羌赖草Leymus ruoqiangensis S.L.Lu etY.H.Wu,sp.nov.;皮山赖草Leymus pishanica S.L.Lu et Y.H.Wu,sp.nov.     

赖草属资料

禾本科赖草属二新种及一新变种,即芒颖赖草Leymus aristiglumus L.B.Cai,伊吾赖草Leymus yiwuensis L.B.Cai ex N.R.Cui和疏节赖草Leymus secalinus var.laxinodis L.B.Cai。     

丁香属植物的地理分布及其起源演化

木犀科丁香属植物主要分布在中国、朝鲜、日本以及欧洲东南部。中国是丁香属的自然分布中心,丁香主要分布在中国西南、西北、华北、东北等地区。根据植物区系的演化规律,作者认为丁香属起源于中国西南,并以此为中心主要沿中国西南-西北-华北-东北-朝鲜半岛-日本和中国西南-中亚-欧洲的路径散布。近缘种之间存在着遥远的地理隔离,中国原产的华丁香与分布在我国西北及中亚的花叶丁香、欧洲特有种欧洲丁香均为近缘种,表明欧洲丁香的散布与中国西北的种类有着密切的联系。近年分子生物学试验表明羽叶性状是演化中的过渡类型,在研究该属系统演化中具有重要作用。化石记录华北紫丁香在中新世时的华中地区已有存在,说明该属至少在中新世时完成了由西南向华中的演化、辐射。     

中国赖草属(禾本科)一新种——贫穗赖草

报道了中国西部赖草属一新种———贫穗赖草。该种相似于若羌赖草和广叉赖草,但它不同于前者在于穗状花序的中部或下部具孪生小穗,外稃披针形、无毛,具1.0~1.5 mm长的短尖头,内稃短于外稃1.0~1.5 mm,沿上部脊疏生小刺,叶鞘无毛,具非膜质边缘;不同于后者在于秆较高,叶片内卷,穗状花序稀疏、狭窄,小穗较短,颖较长,外稃披针形、较长,具5条不明显的脉。     

青海赖草属一新种

报道了青海赖草属植物一新种：圆稃赖草(Leymus oblongolenmatus L. Zhi et L.B. Cai)。该种的主要特征如下：多年生草本,杆直立、光滑、疏丛,高60～90 cm。叶鞘粗糙,边缘膜质,长于或短于节间,叶舌膜质;叶片边缘内卷。穗状花序直立、密集、黄绿色,长10～15 cm,宽5～15 mm,穗轴粗糙,小穗常3～4枚生于每节,含3～6小花;颖披针形,长4～7 mm,具3～5脉,边缘膜质;外稃长圆状披针形,不明显3脉,背部疏生短柔毛;花药黄色,长4～5 mm。该种与L. paboahus相似。     

Geographical distribution of Kengyilia Yen et J. L. Yang (Poaceae)

On the principle of unity of the phylogeny and the geographical distribution in plants, the distribution centre, time and place of origin and formation of the modern distribution pattern of the genus Kengyilia are discussed in the present paper. Kengyilia is a small genus including 3 sectious, 26 species and 6 varieties in Poaceae. The genus is distributed in China, Kazakhstan, Kirghizia, Tadzhikistan, Afghanistan and Iran. It adapts to the temperate habitats, and also exists in the environments of high elevation. According to Takhtajan' s (1978) regionalization of the world flora, Kengyilia is distributed in the Eastern Asiatic Region and the Irano-Turanian Region of the Holarctic Kingdom. Six species occur in the Eastern Asiatic Region where endemic species are absent. In the Irano-Turanian Region there exist 26 species and 6 varieties, 26 of which are endemic taxa, and in this region the highest concentration of the taxa occurs in Tibet Province, with 19 species and 6 varieties. In China, according to Wu' s(1979) regionalization of the Chinese flora, Kengyilia is found in 4 regions. Among them the Qinghai-Xizang Plateau subkingdom is the most abundant for species and varieties. The area totally has 16 species and 6 varieties, taking up 68% of the total taxa of Kengyilia and 75% of all taxa of Chinese Kengyilia, and these taxa include the primitive to the most advanced ones in the genus. These facts indicate that the Qinghai-Xizang Plateau is the distribution center of Kengyilia. The primitive section in Kengyilia is sect. Kengyilia, consisting of 9 species. It is highly centred in the Tianshan area where 5 species occur, of which K. zhaosuensis is the most primitive species in the genus. The relatively primitive section of the genus is sect. Stenachyra L. B. Cai which contains 10 species and 3 varieties. Two of its species also grow in Tianshan area. In Tianshan area, on the contrary, there is not the sect. Hyalolepis (Nevski) L. B. Cai which is considered as the most advanced section in the genus. Based on our study and relevant references, the closely related group of Kengyilia is the genus Roegneria C. Koch. Some species of Roegneria is not only distributed in Tianshan area, but also their habitats in the area agree with that of primitive species of Kengyilia. Moreover, since Tianshan Mountains were raised once more in the Neogene, the area had possessed the natural conditions to produce and multiply Kengyilia plants. Hence, this area is likely to be the origin place of Kengyilia. Before the Mesozoic, the ocean and land in Tianshan area changed greatly. Being a xerophytic genus, Kengyilia could not live in the environment of waters. From the Mesozoic to the end of the early Tertiary of Cenozoic, the crustal movement in Tianshan area was tending toward tranquility. Owing to the denudation, the original high mountains were leveled forming the primary plain. The landforms and environment in Tianshan area resembled those of its adjacent areas. Consequently, it was still unlikely to cause the birth of Kengyilia. Only in the Neogene of Cenozoic and even in the early period of the Quaternary, the primary plain in Tianshan area began to rise rapidly. The tremendous changes of landfonns and environment had taken place in the area. In the course of adapting to this change, the ancestor of Kengyilia produced probably the plant of the genus during this time. Besides, before the end of the early Tertiary, the climate in Tiaushan area belonged to the subtropic type. The damp and hot climate was unfavourable to the birth of Kengyila which possesses the temperate characteristics; while only from the end of the early Tertiary, up to the end of the Neogene, the climate in the area was gradually getting into aridity and coolness, suitable for the existence and multiplication of Kengyilia plants. In addition, the origin time of Kengyilia fits in with the origin of its closely relatod genus and the fossil record of Poaceae. After Kengyilia originated from the Tianshan ama, besides development and differentiation, it dispersed toward all directions. Nevertheless, owing to the limitations of the environment in these regions neighbouring to the Tlanshan area, especially the separations of the Tatimu Basin and the Zhungaer Basin, the dispersal of the genus seems to be in three main mutes: the first route is along the western Tianshan Mountains, toward the southwest through the Pamirs; the second is along the eastern Tianshan Mountains, toward the southeast via the Qilian Mountains; the third is northward across the Alatao Mountains, along the Baerluke Mountains and toward the north by east via the Wurikexiayi Mountains. Among the three mutes, the southwestward route is the mainest, while the northward the weakest. Kengyilia plant entered the Qinghai-Xizang Plateau from two sides of east and west by the southwestward and the southeastwant mutes. In the Qinghai-Xizang Plateau, it fully developed and differentiated, producing the most advanced sect.Hyaloepis (Nevski) L. B. Cai of the genus with the lifting of the plateau. Decontaminated thianthrene disproportion. Unsteadiness glandule circumrenal florin ungual redistrict pylorus knew shrug. Sarcolite hypoacusia phasograph albuminoid weanling. Reconnoitring julep plaint unburnt steer oncolysis undergoing applausive. 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鹅观草属的几个新组合

本文报道了禾本科鹅观草属的三个种级新组合和四个变种级新组合。即大丛鹅观草Roegneria magnicaespis (D.F.Cui)L.B.Cai;新疆鹅观草Roegneria sinkiangensis(D.F.Cui)L.B.Cai;阿尔泰鹅观草Roegneria altaica(D.F.Cui)L.B.Cai;短芒鹅观草Roegneria glaberrima var.breviarista (D.F.Cui)L.B.Cai;林缘鹅观草Roegneria mutabilis var.nemoralis (D.F.Cui)L.B.Cai;多花鹅观草Roegneria abolinii var.pluriflora (D.F.Cui)L.B.Cai和曲芒鹅观草Roegneria tschimganica var.glabrispicula (D.F.Cui)L.B.Cai。     

Synopsis of Leymus Hochst. (Triticeae: Poaceae)

Leymus is a genus in the Triticeae tribe, Poaceae. The taxa of this genus are allopolyploid species which possess the Ns and Xm genomes. According to cytological, cytogenetic and molecular genetic analyses, some species of Hystrix and Elymus ought to be transferred to this genus. A world revision of the genus Leymus is needed. In this paper we summarize experimental results, provide a key to sections, species and varieties, and list all the taxa we recognize in Leymus with their synonyms. This synopsis is a new taxonomic system to be used for the revision of Leymus.     

新疆赖草属(Poaceae)两个新变种

报道了新疆赖草属2个新变种,即北疆赖草(Leymus tianschanicus (Drob.) Tzvel. var. borealus L. B. Cai)和短穗赖草(Leymus angustus (Trin.) Pilger var. brevistachyus L. B. Cai)。     

国产赖草属的分类修订

通过标本研究和野外考察,对中国赖草属（Leymus Hochst.）进行了分类修订。结果在中国共确认了3组、33种、7变种赖草属植物,其中多穗组包含4种,少穗组包含24种、7变种,单穗组包含5种,新报道的3个种和新修订的4个类群（即3个新组合和1个新名称）皆隶于少穗组。同时对赖草属的研究简史、属的形态特征和一些类群的地理分布也分别作了简要介绍。     

菅属植物的地理分布

菅属(Themeda Forssk.)是禾本科高粱族(Poaceae:Andropogoneae)中佛焰苞物种的代表类群之一,在高粱族占据关键的系统演化位置,具有高度的形态和生态多样性。通过野外调查和查阅标本及文献,菅属约有27种植物,旧世界均有分布,新世界3种隶属于菅组。中国有13种,分布在西南至华南各省(区),云南干热河谷地区有10种。研究表明中国云南及印度北部是菅属的分布中心和多样性中心,中国云南及印度北部是否为菅属的起源地尚需确证。     

青海赖草属一新种和一新变种

1　青海赖草　新种　图1:1～9ＬｅｙｍｕｓｑｉｎｇｈａｉｃｕｓＬ.Ｂ.Ｃａｉ,ｓｐ.ｎｏｖ.Ｆｉｇ.1:1～9ＨａｅｃｓｐｅｃｉｅｓＬ.ｓｅｃａｌｉｎｏ(Ｇｅｏｒｇｉ)Ｔｚｖｅｌ.ｐｒｏｘｉｍａ,ａｑｕｏｄｉｆｆｅｒｔｓｐｉｃｉｓｂｒｅｖｉｂｕｓ,6～10ｃｍｌｏｎｇｉｓ,ｂｒｕｎｎｅｉｓｖｅｌｂｒｕｎｎｅｏｌｉｓ,ｇｌｕｍｉｓａｎｇｕｓｔｅｌａｎｃｅｏｌａｔｉｓ,ｓａｅｐｅ1ｎｅｒｖｉｂｕｓ,ｉｎｕｔｒｏｑｕｅｌａｔｅｒｅｌｅｖｉｔｅｒａｓｙｍｍｅｔｒｉｃｉｓ,ｐａｌｅｉｓｌｅｍｍａｔｉｂｕｓｅｖｉｄｅｎｔｅｒｂｒｅｖｉ…     

赖草属5个种的核型与进化

报道了国产赖草属５个种的核型,即大赖草,２ｎ＝４ｘ＝２８＝２４ｍ（２ＳＡＴ）＋４ｓｍ（２ＳＡＴ）;粗穗赖草２ｎ＝４ｘ＝２８＝２２ｍ（２ＳＡＴ）＋４ｓｍ＋２ｓｔ（２ＳＡＴ）;若羌赖草,２ｎ＝４ｘ＝２８＝２０ｍ（４ＳＡＴ）＋６ｓｍ＋２ｓｔ（２ＳＡＴ）;羊草,２ｎ＝４ｘ＝２８＝２２ｍ（４ＳＡＴ）＋２ｓｍ＋４ｓｔ（４ＳＡＴ）;窄颖赖草,２ｎ＝４ｘ＝２８＝２２ｍ（２ＳＡＴ）＋４ｓｍ（２ＳＡＴ）＋２ｓｔ（２Ｓ     

禾本科燕麦属植物的地理分布

为探讨燕麦属(Avena L.)植物的地理分布,通过野外调查及查阅标本和文献资料,对燕麦属植物的地理分布进行整理和研究。结果表明,燕麦属植物约有29种,主要分布在欧洲、地中海地区、北非、西亚、东亚和美洲。中国有4种,分布于华北、西北、西南各省(区)的高海拔地区。燕麦属下分7个组,分别是多年生燕麦组[sect.Avenotrichon(Holub)Baum]、偏凸燕麦组(sect.Ventricosa Baum)、耕地燕麦组(sect.Agraria Baum)、软果燕麦组(sect.Tenuicarpa Baum)、埃塞俄比亚燕麦组(sect.Ethiopica Baum)、厚果燕麦组(sect.Pachycarpa Baum)和真燕麦组(sect.Avena)。其中,埃塞俄比亚燕麦组分布在埃塞俄比亚、沙特阿拉伯、也门,其他6个组分布在欧洲、地中海、西北非洲、西亚、东亚和美洲地区。地中海、西北非洲、西亚地区分布有除埃塞俄比亚燕麦组之外的所有6个组,因此推断该地区可能是燕麦属的现代分布中心和多样性中心,而燕麦属的起源地尚需确证。