Pselaphotumulus,a new genus of pselaphine endemic to New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Pselaphitae)

ABSTRACT

Pselaphotumulus Owens and Carlton, gen. nov., is described as the sixth genus in the tribe Pselaphini known from New Zealand. Three new species are described: Pselaphotumulus aorerei, sp. nov., Pselaphotumulus dubius, sp. nov. and Pselaphotumulus unus, sp. nov.. Three species, Pselaphus cavelli (Broun 1893), Pselaphus oviceps (Broun 1917) and Pselaphus urquharti (Broun 1917) are transferred to Pselaphotumulus, nov. combs. Lectotypes from type series in the New Zealand Broun Collection (Natural History Museum, London) are designated for these three species. Habitus photographs, distributional maps and line drawings of diagnostic characters are provided for each species. A key to species is provided. Searches of museum collections have not yielded representatives outside of New Zealand, suggesting that this is the first endemic genus in the tribe Pselaphini described from the New Zealand’s main islands, specifically, the South Island. Pselaphotumulus species exhibit restricted distribution patterns that approximately coincide with the Pacific/Indo Australian Plate boundary.     

Revision of the genus Adorodocia Brenske 1893 (Coleoptera: Scarabaeoidea: Rutelinae: Adoretini) endemic to Madagascar

Summary. The Adoretini of the Malagasy endemic genus Adorodocia Brenske 1893 is revised. Fourteen new species and one new subspecies are described and compared with their most closely related species: A. constricta n. sp., A. cuccodoroi n. sp., A. flava n. sp., A. liliae n. sp., A. marginata n. sp., A. peyrierasi n. sp., A. pseudoconstricta n. sp., A. pseudoflava n. sp., A. pseudostrigata n. sp., A. recta n. sp., A. robusta n. sp., A. sogai n. sp., A. vadoni n. sp., A. viettei n. sp. and A. vittaticollis flavipes n. ssp. The synonymy between Adoretus strigatus Waterhouse 1878, and Pseudadorodocia aenigma Arrow 1901, is confirmed. Thus, based on the results of this study, the genus Adorodocia includes 16 species, and one of them is represented by two subspecies. Diagnostic characters to separate the species in the genus deal mostly with the shape of parameres, color of body and legs, shape of pronotum and female eighth tergite, setation of pronotum and elytra. Key to species, diagnoses and distribution for each species are provided. Endophallus and female genitalia are illustrated for the first time for this genus.     

On the Genus Trypeticus Marseul (Coleoptera: Histeridae: Trypeticinae) in China

This paper describes two new species, Trypeticus fissirostrum n. sp. from Hainan and Trypeticus yunnanensis n. sp. from Yunnan Provinces, China. Three Trypeticus species having been recorded in the territory of China before this study, their total number in this country has now increased to five. A key to the Chinese species is compiled and given in the text. The type specimens are deposited in Institute of Zoology, Chinese Academy of Sciences.     

Preliminary notes on the genus Aeschnosoma Selys 1870 (Odonata: Anisoptera: Corduliidae s. str.)

Three new species of the genus Aeschnosoma are briefly described and illustrated. A. pseudoforcipula n. sp. and A. heliophila n. sp., both from the Brazilian Central Plateau are respectively related to the two Amazonian species A. forcipula Hagen in Selys 1871, and A. auripennis Geijskes 1970. A. louissiriusi n. sp. from Northern Brazil is not closely related to any known species. Based on larval and adult derived characters, the genus Aeschnosoma appears closely related to the Australian genus Pentathemis Karsch 1890, and also to the Madagascan genus Libellulosoma Martin 1907. The clade Aeschnosomata nov. is erected to receive the three genera. Some putative plesiomorphies would place this clade sister group of the remaining Corduliidae s.str.     

A review of the Malagasy endemic genus Cyrtolophus Fairmaire and description of Pseudocyrtolophus n. gen. (Coleoptera: Scarabaeidae: Rutelinae: Adoretini)

Summary

After lectotype designations for both names, the synonymy between Cyrtolophus grandicornis Fairmaire, 1895 and Cyrtolophus viduus Ohaus, 1912, n. syn., is proposed. Pseudocyrtolophus n. gen. is described to accommodate Cyrtolophus limbicollis Ohaus, 1912, its type species, and a second species, Pseudocyrtolophus australis n. sp., which is described and compared to its most related species. The lectotype of Cyrtolophus limbicollis is designated. All species are illustrated. A map showing their distribution in Madagascar is given.     

Revision of the New Zealand genus Novothymbris (Homoptera: Cicadellidae)

The genus Novothymbris Evans is endemic to New Zealand, and is the only representative in this country of the subfamily Ledrinae. The genus and the known species are redescribed; dunensis (Myers) is synonymised with hinemoa (Myers), and hudsonica (Myers) with maorica (Myers). Eleven new species are described; three species‐groups are indicated. A key to species is given.     

A revision of the genus Paronthobium from New Caledonia (Coleoptera: Scarabaeidae: Scarabaeinae: Epilissini)

A revision of the New Caledonian genus Paronthobium Paulian 1984 is presented. Anonthobium Paulian 1984 is synonymized with Paronthobium Paulian 1984 after evaluating the diagnostic characters originally introduced by Paulian to separate them. Upon examination of the type specimens, Onthobium caledonicum Paulian 1935 is separated from O. simplex Fauvel 1903 and restored as valid. Anonthobium moui Paulian 1984 is synonymized with Anonthobium micros Paulian 1984. Fifteen new species are described: P. adio n. sp., P. daghfousi n. sp., P. dierkensi n. sp., P. farino n. sp., P. iac n. sp., P. julieni n. sp., P. memaoya n. sp., P. minutum n. sp., P. nasutum n. sp., P. orientale n. sp., P. peckorum n. sp., P. petchecara n. sp., P. subdentatum n. sp., P. taom n. sp. and P. tchingou n. sp. The genus Paronthobium is now composed of 22 species. Illustrations of parameres and of male protibiae are provided for each species. Distribution maps and a key to species are given.     

Philopotaminae of Madagascar (Trichoptera: Philopotamidae)

The Malagasy fauna includes two genera of Philopotaminae: Wormaldia McLachlan 1865 and an endemic new genus, Ranarijaodes n. gen. The Malagasy and African (sensu stricto) lineages of Wormaldia are distinct. Apomorphic characters (the shape of the phallic apparatus for the African species and the splitting of the median lobe of the tenth tergite for the Malagasy species) confirm their separation. New captures are reported for two of the four previously known species of Wormaldia. Seven new species are described: Wormaldia ambahibeae n. sp., W. faharoa n. sp., W. fahatelo n. sp., W. randriamasimananai n. sp., W. fahefatra n. sp., W. legrandi n. sp. and W. fahadimy n. sp. Ranarijaodes n. gen. is close to Dolophilodes Ulmer 1909. It is described for the three new species Ranarijaodes fiana n. sp., R. angapi n. sp. and R. atsimo n. sp. The 14 Malagasy species of Philopotaminae are endemics of the island. Their geographical distributions and ecological preferences are described and discussed.     

Taxonomic revision and phylogeny of the genus Aneorhachis Kleine 1923 (Coleoptera: Brentidae: Microtrachelizini)

A taxonomic revision of the genus Aneorhachis Kleine 1923 (Coleoptera: Brentidae) and a phylogenetic analysis of species belonging to this genus are carried out. The genus Ipsopisthius Kabakov 2001 is synonymised with Aneorhachis, and two new combinations are proposed: Aneorhachis hirta (Kabakov 2001), n. comb. and A. incerta (Kleine 1935), n. comb. Higonius nitens Goossens 2008 is synonymized with A. incerta. One new species is described from New Guinea, A. papuana n. sp. All the species are redescribed and type-specimens are illustrated; an identification key is proposed. A maximum parsimony analysis is provided on the base of 23 characters from adult morphology. This analysis confirms the monophyly of the genus and seems to show that Aneorhachis originated on continental Asia and then spread out eastward to colonize Pacific islands.     

Three new species of Philocorydoras Suriano, 1986 (Monogenoidea: Dactylogyridae) infecting the gills of callichthyids (Actinopterygii: Callichthyidae) from the Peruvian Amazonia

During a research on gill ectoparasites of callichthyids fishes from the Peruvian Amazonia, the following monogenoideans were found: Philocorydoras peruensis n. sp. from Corydoras splendens (Castelnau); Philocorydoras multiradiatus n. sp. and Philocorydoras jumboi n. sp. from Brochis multiradiatus (Orcés, V.). All new species described herein are mainly differentiated from their congeners based on the morphology of the copulatory complex. In P. peruensis n. sp. the cirrus is “J”-like shaped tube slightly tilted to one side, while in P. jumboi n. sp. is “J”-like shaped tube in a straight position and in P. multiradiatus n. sp. the cirrus is an arced tube with inflated base and distally narrow. Brochis (Orcés, V.) represents a new genus hosting species of Philocorydoras. All new species presented in this work represent the first species of Philocorydoras reported for Peru.

     

Species Diversity of Japanese Clusiidae (Diptera: Acalyptrata) with description of 12 new species

The Japanese Clusiidae are revised and 23 species, including 12 species new to science, are recognized and keyed. Clusiodes angulosus n. sp., Clusiodes discostylus n. sp., Clusiodes tobi n. sp., Clusiodes usikumuri n. sp., Craspedochaeta varicolor n. sp., Heteromeringia crenulata n. sp., Heteromeringia quadrispinosa n. sp., Heteromeringia sexramifera n. sp., Heteromeringia yamata n. sp., Phylloclusia quadrivittata n. sp., Sobarocephala uncinata n. sp., and Tetrameringia borealis n. sp. were described as new. Distribution records were compiled and mapped. Transitional climatic zone between the warm and cool temperate zones is possibly suggested as one of the most richest temperate area in terms of Japanese clusiid species richness. Hendelia beckeri Czerny 1903, Paraclusia japonica Sasakawa 1957, and Clusiodes plumosus Sasakawa 1964 are broadly distributed over all the Japanese temperate forests, but these are not always most abundant species. The clusiid assemblage was heterogeneous in the species composition among various forest types. The femalebiased sample caught by Malaise traps and distinction of clusiid species composition between temperate and subtropical zones are also pointed out.     

New Hypogastrura species and new distributional data for Hypogastrura distincta (Axelson 1902) from China (Collembola: Poduromorpha: Hypogastruridae)

Two new species, Hypogastrura analpapillata n. sp., on mushrooms from Yunnan Province, and Hypogastrura microspinata n. sp., on moss from Hunan Province, are described. Hypogastrura analpapillata n. sp. is the sixth species with highly modified mouthparts in the trybomi group. Hypogastrura microspinata n. sp. is close to H. yosii Stach 1964, but can be differentiated from the latter by body granules, tenent hair and lateral teeth on tibiotarsus. A key to Chinese species of Hypogastrura is provided and a list of Chinese Hypogastrura species and their distributional data are given.     

A revision of the Chinese Catops Paykull 1798 of the Catops fuscus species group (Coleoptera: Leiodidae: Cholevinae)

Five species of the Catops fuscus species group are reported from China. Three species are newly described: C. hlisnikovskyi n. sp. from the Beijing municipality and Jiangsu province, C. schuelkei n. sp. from Sichuan and Yunnan provinces, and C. smetanai n. sp. from Sichuan and Yunnan provinces. Female of C. sasajii Nishikawa 2007 is described and the species is reported from Hubei, Sichuan and Yunnan provinces for the first time. Catops nigricans (Spence 1813) is reported for the first time from China (Xinjiang Uygur autonomous region). Important morphological characters are illustrated and the distribution of all species in China is mapped. Preliminar phylogenetic analysis separates (C. hlisnikovskyi n. sp.+C. fuscus fuscus Panzer 1794) as a sister clade to C. nigricans+(C. sasajii+(C. schuelkei n. sp.+C. smetanai n. sp.)), with C. paramericanus Peck & Cook 2002 as outgroup.     

Review of the Fannia jezoensis-group (Diptera: Fanniidae)

The jezoensis species-group is established within the genus Fannia Robineau-Desvoidy consisting of four species, Fannia curvostylata n. sp., Fannia densa Nishida, Fannia jezoensis Nishida and Fannia zhangi Xue. The new species described here is from the Tibetan Plateau. The known species of this group are all described, illustrated and keyed. Geographic distributions are updated. The systematic position and biogeography of the F. jezoensis-group are also discussed.     

Afrotropical and Nearctic genera of Odonata in the French Oligocene: biogeographic and paleoclimatic implications (Insecta: Calopterygidae,Aeshnidae)

New species of the genera Sapho and Epiaeschna are recorded in the Oligocene of Aix- en-Provence, Bouches-du-Rhône, France. Sapho legrandi n. sp. is the third fossil representative of this recent African genus and Epiaeschna pseudoheros n. sp. is the fifth fossil species of this recent North American genus. The fossil species Triaeschna gossi from the Eocene of England, Epacantha magnifica from the Late Oligocene of Kazakhstan, and Mediaeschna matutina from the Oligocene of China, are considered species of Epiaeschna and the three fossil genera Triaeschna Campion 1916, Mediaeschna Zhang 1989, and Epacantha Martynov 1929 are synonymized with Epiaeschna. The closely related genera Umma and Sapho inhabit warm humid forests of Western Africa. Their presence in two Oligocene deposits of France supports the hypothesis of a warm humid palaeoenvironment for Armissan (Aude, France), and Aix-en-Provence.     

Species level taxonomy of the Neotropical hairstreak genus Porthecla (Lepidoptera: Lycaenidae: Theclinae: Eumaeini)

The species level taxonomy of Porthecla is revised. Six previously described species are recognized: P. ravus (Druce 1907); P. barba (Druce 1907); P. dinus (Hewitson 1867); P. porthura (Druce 1907); P. minyia (Hewitson 1867); P. gemma (Druce 1907), and another six new species are proposed: P. forasteira Faynel & Moser n. sp.; P. annette Faynel & Robbins n. sp.; P. johanna Faynel & Robbins n. sp.; P. peruensis Faynel & Moser n. sp.; P. prietoi Faynel & Busby n. sp. and P. willmotti Busby, Faynel & Moser n. sp. For each Porthecla species we present diagnostic characters, images of male and female adults, drawings of male and female genitalia, distribution maps, and notes on habitat and other biological traits. An identification key for males is provided. To stabilize names; lectotypes are designated for Thecla minyia Hewitson 1867, and Thecla gemma Druce 1907. Wing pattern resemblances between Porthecla and other genera are discussed.