The timing of breeding and wing-moult of four African Sturnidae

Wing-moult of the Cape Glossy Starling, Red-winged Starling, Pale-winged Starling and Pied Starling was examined primarily from specimens in southern African museums. Breeding data were obtained from nest record cards.
The Cape Glossy Starling breeds from October to March, with the moult period from December to May. There is no evidence of moult-breeding overlap in individual birds. The Red-winged Starling breeds from September to March, while the moult takes place between November and April, overlapping with the second broods. The Pale-winged Starling breeds from October to April and moults between November and May. The Pied Starling moults between November and April, while breeding varies regionally, occurring concurrently with moulting in some areas.     

THE SURVIVAL RATES OF SOME SOUTHERN AFRICAN PASSERINES

Yom-Tov, Y., McCleery, R. & Oatley, T. 1994. The survival rates of some southern African passerines. Ostrich 65:329-332.

Little information is available on survival rates of southern hemisphere passerines, but two recent studies indicate that Australian passerines tend to live longer than their northern hemisphere relatives. The aim of this paper is to provide data on survival of some South African passerines and compare them with Australian data. The survival rates of eleven species (eight genera) of South African passerines were calculated using recapture data provided by the South African Bird Ringing Unit for birds whose retraps occurred more than 12 months after the original ringing. Survival did not depend on either the length of the study or body mass. Mean survival rate was similar to that of 32 species of Australian passerines, which is at least 1,2 greater than that of British passerines.     

THE CHARACTERS OF A NEW RACE OF Alcedo semitorquata Swainson FROM THE LOW COUNTRY OF PORTUGUESE EAST AFRICA

Arkell, G. B. F. 1979. Aspects of the feeding and breeding biology of the Giant Kingfisher. Ostrich 50:176-181.

Giant Kingfishers Megaceryle maxima were studied during 1972–1975 on the Eerste and Blaauwklip Rivers near Stellenbosch (33 53s; 18 50E), South Africa. Nest record cards of the Southern African Ornithological Society were examined for dimensions of the nest, nest lining, clutch-size and brood-size. The feeding rate of nestlings was determined from observations at the nest. Two adults, comprising a pair of breeding birds, and a fledgling wert equipped with breast-mounted miniature radio transmitter packages to monitor the birds activity and home range. Cape River Crabs Potamon perlatus apparently were the main prey of the kingfishers. A correlation was found between the activity patterns of kingfishers and of crabs, which peaked in the mornings and evenings.     

Sociable Weaver biometrics and primary moult

The biometric and primary moult data housed at the South African Bird Ringing Unit (SAFRING) were analysed for the Sociable Weaver Philetairus socius. The average body mass and wing length was 27.9g (SD = 2.2) and 74.1mm (SD = 2.5), respectively. Variation in these parameters is not clearly correlated with region, season or climate, other than a negative correlation of body mass with average annual water deficiency. Body mass of Sociable Weavers near Kimberley showed a longterm decrease of 2.9g, probably due to stabilising selection on mass. Primary moult duration varied from 152 days to 169 days and started between 26 January and 31 December in two populations (socius and South African eremnus respectively). Individual primaries moulted mainly one at a time, each taking 20–28 days to grow fully. Prolonged moult duration in this species is probably an adaptation to reduce energy expenditure, and to grow more durable feathers due to abrasion in entering the nest. The lack of clear patterns of geographical variation in biometrics indicates that the contiguous populations of Sociable Weaver should belong to the nominate species. The biometric and primary moult data housed at the South African Bird Ringing Unit (SAFRING) were analysed for the Sociable Weaver Philetairus socius. The average body mass and wing length was 27.9g (SD = 2.2) and 74.1mm (SD = 2.5), respectively. Variation in these parameters is not clearly correlated with region, season or climate, other than a negative correlation of body mass with average annual water deficiency. Body mass of Sociable Weavers near Kimberley showed a longterm decrease of 2.9g, probably due to stabilising selection on mass. Primary moult duration varied from 152 days to 169 days and started between 26 January and 31 December in two populations (socius and South African eremnus respectively). Individual primaries moulted mainly one at a time, each taking 20–28 days to grow fully. Prolonged moult duration in this species is probably an adaptation to reduce energy expenditure, and to grow more durable feathers due to abrasion in entering the nest. The lack of clear patterns of geographical variation in biometrics indicates that the contiguous populations of Sociable Weaver should belong to the nominate species.     

DISPERSAL AND SITE FIDELITY OF CAPE GANNETS MORUS CAPENSIS

Klages, N. T. W. 1994. Dispersal and site fidelity of Cape Gannets Morus capensis. Ostrich 65:218-224.

Site fidelity and dispersal of juvenile and sexually mature adults was investigated at the largest southern African gannetry on Bird Island, Algoa Bay, where a minimum of 65000 airs occupy approximatel 2 ha at a mean density of 2,85 nests per m². Based on 512 ring recoveries and live retraps of known-aged birds of this colony, immature and non-breeding adult gannets dispersed widely along the coasts o Africa, reaching Maputo Bay (southern Mozambique) on the eastern side of the continent and the Namibian Skeleton Coast on the western side. Very few birds flew farther. There were conspicuously few ring recoveries at 400–500 km straight-line distance east from Bird Island, corresponding with the Pondoland coast in Transkei. A similar drop was observed in the west for the Tsitsikamma coast and the stretch of coast from Cape St. Blaize to Cape Infanta. In these regions reporting by members of the public is presumed to be low. either because the rocky coast is inaccessible or because of socioeconomic reasons. The geographical spread of the ring recoveries suggested a possible range contraction of the species between the 1950's and the period 1978–1993 but the nature of the data prevented a test of this hypothesis. No evidence was found that gannets from Bird Island breed or roost at other southern African colonies as a matter of habit. On the contrary, all survivors of post-fledgling dispersal eventually returned to their native colony, where they exhibited pronounced fidelity to the site where they hatched from the egg. Based on multiple retraps of 862 known-age individuals, 71.7 % of young adults took over vacant sites in the immediate vicinity of their original nest, 8,1% moved once but were faithful to their new site and in 20,2% of all cases faithfulness to the original site could not be shown. Moreover, in subsequent years edge birds did not vacate their sites to move towards the center, refuting the hypothesis that fringes are physically inferior breeding sites.     

REMARKS ON THE DISTRIBUTION OF BIRDS COLLECTED IN MASHONALAND AND EASTERN SOUTHERN RHODESIA (Part I)

Oatley, T. B. 1983. Twenty-third ringing report for southern Africa. Ostrich. 54:141-149.

Bird ringing activities in southern Africa are summarized for a two-year period from July 1979 to June 1981. A total of 42 374 birds were ringed. Much ringing effort was devoted to marine buds. Palaearctic migrants comprised nearly 12% of the birds ringed, with waders, swallows and warblers being the most frequently ringed families. There were 747 recoveries or recaptures reported during the reviey period, of which 73 were foreign-ringed birds. The remainder involved 103 species of birds, only nine of which contributed 10 or more recoveries to the total. Five of the nine are marine birds and collectively account for 55% of all recoveries and recaptures. A selected list of recoveries and recaptures of southern African-ringed birds is given.     

THE PRIMARY MOULT OF WADERS ON THE ATLANTIC COAST OF MOROCCO

Data from 3659 waders of 23 species live-trapped in the years 1971-73 on the Atlantic coast of Morocco during the period of autumn moult and migration are analysed to estimate duration and timing of primary moult. Common Sandpiper was the only species to moult primaries in its first autumn (unless published ageing criteria are incorrect). Several species showed a low incidence of arrested primary moult and a higher incidence was observed in Ringed, Kentish and Grey Plovers. This is discussed in relation to breeding and migration. Similar rates of primary feather replacement relative to specific moult duration were observed in all species for which information was available. Comparisons between species and with published studies showed that variations in rate of moulting between species and between different geographical populations of the same species were largely due to differences in feather growth rate rather than in the numbers of primaries concurrently in growth. Variations in rate between individuals of the same population were achieved, at least in the first part of moult, by differences in feather dropping rate resulting in differences in the numbers of primaries growing concurrently. The timing and duration of moult in different populations and differences between breeding and non-breeding components were closely related to the requirements of other annual cycle activities, notably breeding and migration. Non-breeding birds summering in Morocco had started moult early. Locally breeding birds had an early start to a fairly slow moult which overlapped with breeding and which in some cases passed through an arrested stage. Birds breeding in cold temperate and arctic regions and wintering in Morocco moulted in a short time soon after arrival. In some cases, notably in Ringed Plovers, birds had commenced moulting on the breeding grounds and arrested moult during migration. Most Redshank and possibly Dunlin migrated in active wing moult. The fastest primary moult was achieved by high arctic breeding birds, Curlew Sandpiper and possibly Little Stint, which stopped to moult in Morocco before moving on to wintering areas further south. This situation is contrasted with that of populations of these two and other species wintering in the southern hemisphere where moult occurs over an extended period during the northern winter.     

Moult of the giant petrels Macronectes halli and M. giganteus at South Georgia

Moult scores were collected from colour-ringed individuals of known reproductive status of the two species of giant petrel, Macronectes halli and M. giganteus , at Bird Island, South Georgia between 1978–81.
Both species showed a substantial overlap between breeding and wing-moult, unlike most other Southern Ocean seabirds. Males started moult before females and both sexes of M. giganteus moulted at an earlier stage of the breeding cycle than M. halli , which breeds six weeks earlier than its congener.
Changes in moult rate during the breeding season are documented for both species, with Id. halli showing a rapid increase as the chick nears fledging. Male M. giganteus show a notably different pattern to the other three species-sex groups, starting moult much earlier (at egg-laying), with greater individual synchrony and usually suspending primary moult throughout the main chick growth period, whereas only two male M. halli and no females of either species suspended moult. Differences in pattern, timing and rate of moult are interpreted in terms of availability of food resources and the competing energy demands of other activities, especially chick-rearing.
Completion of primary moult could not be observed in the field but was estimated using data frcsm non-breeding birds and failed breeders; the latter started a rapid moult almost immediately they failed. In both sexes of both species moult is probably concluded at least by early winter.
The general pattern of moult in giant petrels at Bird Island is contrasted with that of other populations and species of Southern Ocean seabirds. It is suggested that the unusually extensive overlap between breeding and moult in giant petrels is a consequence of the very abundant and easily available summer food supplies (especially carrion) and the much diminished winter resources, favouring a completion of moult by the beginning of the winter.     

Notes on the moult of red-backed shrikes ( Lanius collurio ) in their non-breeding range

Moult data from 302 museum skins and 11 trapped birds from sub-Saharan Africa show the course of flight feather moult. Most birds seem to start flight-feather moult soon after arrival in their southern African non-breeding ranges. About 75% of the birds had started before mid-December, i.e., during the main arrival time of the species. The mode of moult scores 1 and 2 was reached on 7 December; the last birds with a score of zero occurred in the first days of January. The mode of moult scores 5 and 6 was reached on 27 February. Thus, the time elapsed between the days when 50% of the population had reached the first and last stages of recorded moult was about 82 days; nine days later 75% had reached this last stage before moult was completed. Thus, individual moult may be estimated to cover about 80–90 days. The main moulting period is between mid-November and mid-March, thus covering about four months. No temporal difference was detected between males and females. A tendency for an advancement of adults compared to young birds was not statistically significant. According to the progress of the moult, sexing of young birds in the field is possible for 50% of the birds towards the end of January and for most birds before mid-February.     

NOTES ON THE PEARL-BREASTED SWALLOW HIRUNDO DIMIDIATA IN THE SOUTH-WESTERN CAPE

Earlé R. A. &; Herholdt, J. J. 1988. Breeding and moult of the Anteating Chat Myrmecocichla formicivora. Ostrich 59: 155–161.

The general breeding biology and moult of the Anteating Chat Myrmecocichla formicivora was studied in open grassveld over a two-year period. During the winter (July), groups were significantly smaller than during summer (December) (1,81 ± 0,50 versus 2,85 ± 1,35 birds per group). There was a large turnover of individuals in the study area but the total population stayed the same. The breeding season in the study area lasted from September to February but analysis of nest record cards from a larger area gave a breeding season of August-April. Two types of nests were used: 90,6% were burrows in sand banks or other excavations, but 9,4%were in the mud pellet nests of Greater Striped Swallows Hirundo cucullata (n = 53). Consecutive breeding attempts were never made in the same burrow. Clutches consisted of three, four or five eggs ([Xbar] = 3,73 ± 0,67). Incubation lasted 14–14,5-15 days. The nestling period lasted 15–18 days. Fledgling/egg breeding success was 41,8% with 48,2% of all eggs not reaching the hatching stage. Juveniles showed an unequal sex ratio of 0,57 ♂ ♂: 1,0 ♀ ♀ but adults had a nearly equal ratio (0,9 ♂ ♂: 1,0 ♀ ♀). There was a significant positive correlation between the primary moult score and the week of the seven months in which moult was recorded. Juveniles underwent a complete body moult and partial primary moult 3–4 months after fledging.     

Migrant European Swallows Hirundo rustica in southern Africa: a southern perspective

Oatley, T.B. 2000. Migrant European swallows Hirundo rustica in southern Africa: a southern perspective. Ostrich 71 (1 & 2): 205–209.

A total of 1 457 recoveries and controls of European Swallows Hirundo rustica in the South African Bird Ringing Unit data bank were analysed to provide an update of the apparent origins and dispersion of migrant European Swallows in southern Africa, to investigate local movements within southern Africa, and to yield an estimate of the age structure of the visiting migrants. Although birds from the former USSR and from Britain and Ireland predominate, over 200 recoveries from 20 other European countries have been recorded. More than half of the swallows migrating to southern Africa are from the central third of the species' breeding range. The dispersion of recoveries of swallows ringed in Britain and Ireland is widespread, with 55% recorded from the Eastern Cape, Free State and KwaZulu-Natal. Controls of southern African-ringed swallows at roosts and recoveries elsewhere in the subcontinent suggest that the daily range of migrant swallows is encompassed within a circle of 100 km diameter, but wide-ranging intra-seasonal movements within South Africa have also been recorded. Conservative estimates of the age of European Swallows at recovery indicate that as many as 14% of migrant swallows ringed in southern Africa may Survive to more than three years of age. It is suggested that the experience of migrants that survive to return to Africa one or more times may permit some flexibility in choice of final destination and that mixing of populations in roosts may result in birds arriving at new destinations.     

THE MIGRATION SYSTEM OF THE CURLEW SANDPIPER CALIDRIS FERRUGINEA IN AFRICA

Elliot, C: C. H., Waltner, M., Underhill. L. G., Pringle, J. S. & Dick, W. J. A. 1976. The migration system of the Curlew Sandpiper Calidris ferruginea in Africa. Ostrich 47:191-213. Data on ringing and recoveries of Curlew Sandpiper, mainly from the Cape, South Africa are presented. Possible migration routes to the breeding grounds are considered in the light of these and other recoveries from the rest of Africa. Retraps show that the species exhibits ortstreue and some evidence is presented which suggests that some birds may travel together and stay in the south in the same flock during one and subsequent migrations. Sex ratio statistics show an excess of females. Adults complete a full primary moult in the Cape between September and February, taking about 140 days but there is a lot of individual variation. Data from Mauritania show primary moult starting faster, a month earlier than in the Cape, and arrested moult in a few adults. The difference may be because Mauritanian birds move on further south while the Cape is the end point of the migration. Kenyan moult records from the Rift Valley follow the Cape pattern except that some birds arrest moult and finish later. Juvenile moult is shown to be different from that of adults, involving only a moult of the outer primaries and taking place during the overwintering period, April to August. All juveniles in the Cape are thought to overwinter and the modified moult to be an adaptation to this behaviour. The weight of adults but not juveniles increases markedly in the six weeks before migration. Fat and protein analyses suggest that the increase is entirely due to deposition of migratory fat. Kenyan birds have lower mean weights and deposit fat about two weeks later than those at the Cape. The nearer the non-breeding quarters are to the breeding grounds, the earlier moult starts and the later fat deposition takes place.     

Bucking the trend: the African Black Oystercatcher as a recent conservation success story

The African Black Oystercatcher Haematopus moquini is a charismatic, southern African near-endemic, wader species, that is often seen as a flagship species for coastal bird conservation, as it was recently down-listed regionally to Least Concern on the IUCN Red List of Threatened Species. To celebrate this rare conservation success story, BirdLife South Africa named it the 2018 Bird of the Year and ran a year-long programme in collaboration with the Nature’s Valley Trust highlighting aspects of the species’ biology, current threats, and conservation success. We used data collected by the Southern African Bird Atlas Project (SABAP1 and SABAP2) to examine changes in the species’ range and relative abundance, both in the records between the two projects, as well as trends within the SABAP2 sampling period (2008–2017). This case study enabled us to assess whether such metrics can accurately reflect abundance and distributional changes in a species. We found increases in the reported range and the reporting rates between the two Atlas projects, and that the SABAP2 reporting rate was stable. Regionally, across four coastal categories, the reporting rate was lowest in KwaZulu-Natal, though this region also showed an increase in the probability of reporting during the SABAP2 period. While corroborating the recent change in the species’ conservation status, we also provide good evidence that the long-term SABAP data can be used successfully to assess population trends and range changes over time.     

DISTRIBUTION,MIGRATION AND TIMING OF MOULT IN THE SOUTH AFRICAN CLIFF SWALLOW

Earle, R. A. 1987. Distribution, migration and timing of moult in the South African Cliff Swallow. Ostrich 58:118-121. The South African Cliff Swallow Hirundo spilodera breeds in South Africa mainly between 25 and 31S and 24 and 31E. In some years with exceptionally high rainfall the breeding range is more extensive. Man has probably had a pronounced influence on the present-day distribution of this species. Seven winter recovery/collecting localities are known from the lower Congo basin in Zaire. Possible migrating birds were observed in Zambia and Malawi in the east, and on the Namibian coast in the west. The few sight records suggest a direct migrational route over Botswana. Moult takes place between March and September primarily in the winter quarters, but about 2% of the birds handled during March and April started moult before migrating.     

6.—DATA OF BIRDS' EGGS IN MY COLLECTION

Van Zyl, A.J, Jenkins, A.R. &; Allan, D.G. 1994. Evidence for seasonal movements by Rock Kestrels Falco tinnunculus and Lanner Falcons F. biarmicus in South Africa. Ostrich 65:111-121.

Patterns of seasonal movement by Rock Kestrels Falco tinnunculus and Lanner Falcons F. biarmicus in South Africa were determined using data from the Southern African Bird Atlas Project, road transect counts, ringing recoveries and other sources. Both species were found to be partial, facultative migrants, and Lanner Falcon movements involved mostly immature birds. Rock Kestrels were found to concentrate in the southwest of the country in the breeding season, and to move north and east in the non-breeding season. Lanner Falcons concentrated in the east of the country during the breeding season, and moved into the west in the non-breeding season. The movement by both species into the arid northwest of the country is probably associated with dramatic increases in prey abundance, in turn associated with summer rah events. Lanner Falcon movement into the southern Cape apparently coincides with cereal farming practices, affecting the availability of prey (granivorous birds and rodents). Rock Kestrel movement into the east of the aunt is probably to exploit increases in insect abundance mated with summer rain. The pattern of Rock Kestrel movements suggests that it does not compete for food with the Lesser Kestrel F. naumanni. a Palearctic migrant to the east of the country.     

GRASSLAND BIRDS NESTING IN HAYLANDS OF SOUTHERN SASKATCHEWAN: LANDSCAPE INFLUENCES AND CONSERVATION PRIORITIES

Abstract: To determine the benefits to grassland birds of converting cropland to hayland in southern Saskatchewan, Canada, we quantified the relative nest abundance and success of grassland nesting birds in haylands and the influence landscape variables have on these parameters. We found nests of 26 species of grassland nesting birds, primarily waterfowl and vesper sparrow (Pooecetes gramineus). With the exception of the northern pintail (Anas acuta), few nesting attempts were recorded for species of high priority in the Prairie Pothole Bird Conservation Region. Mayfield nest success for all waterfowl (20 and 13% in 1999 and 2000, respectively) was high relative to previously reported nest success estimates in other habitat types—especially spring-seeded cropland—and was near levels thought to be required to sustain populations (15–20%). Vesper sparrow nest success (39 and 33% in 1999 and 2000, respectively) also was high relative to that reported in other studies. Haying destroyed few nests as wet weather delayed operations in 1999 and 2000. More nests may be destroyed by haying in other years as approximately 25% of nests in this study were still active on the long-term average haying date for southern Saskatchewan. Among models we developed to explain waterfowl relative nest abundance, amount of cropland in the surrounding landscape and field area were the most informative. Evidence that a specific set of landscape variables was important to models of waterfowl nest success was equivocal. Landscape variables did not explain variation in vesper sparrow relative nest abundance or nest success. Within our study area, conversion of cropland to hayland appears to provide significant benefits to a variety of grassland species, including some species of high conservation priority (e.g., northern pintail). Grassland species of conservation concern nested less frequently in hayland than in native grassland.     

OCCURRENCE AND BEHAVIOR OF THE SOUTHERN RIGHT WHALE DOLPHIN LISSODELPHIS PERONII OFF NAMIBIA

The southern right whale dolphin Lissodelphis peronii is a seldom seen cetacean of southern latitudes. Off southern Africa, observations have been made predominantly in the major upwelling area off Lüderitz, Namibia, but the species has now also been seen south of the Orange River in South African waters. Sightings for the years 1980–1990 are documented and they reveal that L. peronii occurs off southern Africa both inshore and offshore and throughout the year. An opportunity for extended observation in January 1989 afforded the opportunity to record some interesting behavior patterns.     

Nest temperature fluctuations in a cavity nester,the southern ground-hornbill

Southern ground-hornbills Bucorvus leadbeateri inhabit savanna and bushveld regions of South Africa. They nest in the austral summer, which coincides with the wet season and hottest daytime temperatures in the region. They are secondary cavity nesters and typically nest in large cavities in trees, cliffs and earth banks, but readily use artificial nest boxes. Southern ground-hornbills are listed as Endangered in South Africa, with reintroductions into suitable areas highlighted as a viable conservation intervention for the species. Nest microclimate, and the possible implications this may have for the breeding biology of southern ground-hornbills, have never been investigated. We used temperature dataloggers to record nest cavity temperature and ambient temperature for one artificial and 11 natural southern ground-hornbill tree cavity nests combined, spanning two breeding seasons. Mean hourly nest temperature, as well as mean minimum and mean maximum nest temperature, differed significantly between southern ground-hornbill nests in both breeding seasons. Mean nest temperature also differed significantly from mean ambient temperature for both seasons. Natural nest cavities provided a buffer against the ambient temperature fluctuations. The artificial nest provided little insulation against temperature extremes, being warmer and cooler than the maximum and minimum local ambient temperatures, respectively. Nest cavity temperature was not found to have an influence on the breeding success of the southern ground-hornbill groups investigated in this study. These results have potentially important implications for southern ground-hornbill conservation and artificial nest design, as they suggest that the birds can tolerate greater nest cavity temperature extremes than previously thought.     

NOTES ON THE BIOLOGY AND ETHOLOGY OF THE NATAL ROBIN COSSYPHA NATALENSIS

A preference for bushy evergreen undergrowth determines both the local and the general distribution of Cossypha natalensis. The lack of such undergrowth makes equatorial rain-forests unsuitable. Tropical and subtropical evergreen forests provide excellent conditions from this point of view, and they are thus the main habitat type from which C. natalensis has secondarily extended into tropical and subtropical deciduous forests with evergreen undergrowth.
Adult birds remain in or close to their territories throughout the year. First-year birds undertake longer movements during the winter. Breeding territories are relatively small, from 2,500 up to (exceptionally) 10,000 square yards (2,100-8,400 sq. m).
Breeding begins in October. Both sexes build the nest, but only the female incubates. In southern Africa there is one brood a year, and the care of the offspring is protracted. The juvenile moult begins in the third and ends in the sixth month; adults have only one (post-breeding) moult a year.
The begging behaviour of the young is somewhat different from that of most other turdine birds, and is considered to be more primitive. The ontogeny of the adult displays is described as observed in young birds kept in captivity.
The development of vocal behaviour is described. Special attention is given to the different functional forms of the song, and to two of its special features, imitation and a tonal quality termed vox humana.     

BIRDS OF THE UNDERSTORY OF LAKE-SHORE FORESTS ON THE ENTEBBE PENINSULA, UGANDA

Monthly mist-netting of low-flying forest birds was conducted in three lake-shore forests in southern Uganda from September 1970 to July 1972. It was found that the edge of the forest formed a distinct ecological barrier that was rarely crossed by species on either side of it. Catches were concentrated in the hours of daylight, in two peaks, a morning peak at 09.00 hrs and an afternoon peak at 16.00 hrs (sun-time) for all birds combined, although there were slight species differences. Most bird species were found to be sedentary in a fixed home-range, and covered lateral distances of up to c. 300 m. No movement between the different forests was recorded.
The most common birds such as bulbuls were often caught at heights between 0·6 and 1·8 m above the ground, but the upper limit of their vertical range was not determined. It is suggested that the bi-modality of flight activity may be intrinsic, even though it may be correlated with such extrinsic factors as light and radiation. On a month to month basis, the numbers of birds caught reflected breeding and moult activity. This was best shown by the Pygmy Kingfisher, taken in higher numbers towards the end of the first rainy season when a majority of the birds caught were young and many birds were moulting the remiges. On the other hand higher numbers of Olive Sunbird were found during the second rainy season, again at a time corresponding to a general moulting of remiges. The bulbuls showed fluctuations in numbers caught that were matched by an equally fluctuating pattern of moult. The role of food as a possible limiting factor in bird breeding is discussed.