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1.
Many owl species use the same nesting and food resources, which causes strong interspecific competition and spatio-temporal niche separation. We made use of a recent colonisation of Ural Owls (Strix uralensis) in southern Poland to compare habitat preferences of Tawny Owls (Strix aluco) allopatry and sympatry with Ural Owls. We investigated spatial niche segregation of Ural Owl and the Tawny Owl in sympatry and compared habitat preferences of Tawny Owls breeding in allopatry and sympatry. Tawny Owls breeding in sympatry with Ural Owls occupied forests with higher canopy compactness, sites located closer to forest border and to built-up areas, as well as stands with a higher share of fir and spruce and a lower share of beech as compared to sites occupied by Ural Owls. Allopatric Tawny Owls occupied sites with lower canopy compactness and bred at sites located further from forest borders and in stands with lower share of fir and spruce and a higher share of deciduous as compared to sympatric Tawny Owls. As Ural owls are dominant in relation to Tawny Owls, this indicates that the presence of Ural Owls prevents Tawny Owls from occupying deciduous-dominated and old stands located in forest interior areas, far from buildings and forest edges. The results support habitat displacement between the two species when breeding in sympatry. We also show that protection of large forest patches is crucial for the Ural Owl, a species still rare in central Europe, while small patches are occupied by the abundant Tawny Owl.  相似文献   

2.
We used a 13-year time series of abundance estimates of breeding Northern Saw-whet Owls (Aegolius acadicus), and of small mammals from central Ontario, Canada, to assess the numerical response of the owls to small-mammal prey species. We found that the finite rate of increase of breeding owls was directly related to estimates of red-backed vole (Myodes gapperi) abundance. Thus, it appeared that the owls were nomadic, and made decisions about where to breed based on vole supply. The owls showed a much weaker response to deer mouse (Peromyscus maniculatus) abundance. Across all years, 55% of variation in owl rate of increase could be uniquely attributed to vole abundance, whereas only 3% could be attributed to mouse abundance. Consistent with the model of nomadism, there was only a weak relationship between the proportion of hatch-year owls caught at fall banding stations, and small-mammal abundance. Instead, it appeared that Northern Saw-whet Owls avoided years of widespread reproductive failure through the nomadic strategy of selecting breeding sites based on vole supply.  相似文献   

3.
I examined the diet of breeding White-tailed Kites (Elanus leucurus; Aves; Accipitridae) and Barn Owls (Tyto alba; Aves; Tytonidae) in an agrarian area of southern Brazil by analyzing regurgitated prey remains. The objective was to evaluate how these raptors, which differ markedly in their hunting activity periods (owls are nocturnal and kites diurnal), share their mammalian food component. 2,087 prey consumed by Barn Owls and 1,276 by White-tailed Kites were identified. They presented a high overlap of food-niches (Piankas index was 0.98). Based on the daily activity period of their main small mammal prey, a lower overlap would be expected. The crepuscular/nocturnal Mus musculus was the main prey for the diet of breeding Barn Owls (81%) and White-tailed Kites (63%). This small exotic rodent provided 63% of the small mammal biomass ingested by owls and 44% by kites. Larger native small mammals were also considered important for the diet of kites, mainly because of their biomass contribution. Although these raptors differ markedly in their hunting activity periods, Barn Owls and White-tailed Kites are very similar predators in southern Brazil, overlapping their diets.  相似文献   

4.
To use vocalizations properly for the estimation of owl population size, it is important to identify how environmental factors affect owl calling behaviour. Here, we analyse how intrinsic and extrinsic factors modify the vocal activity of Tawny Owls Strix aluco in two areas of northern Spain. From March 2013 to May 2015, we radiotracked 20 Tawny Owls and also undertook a systematic survey in which we collected data on spontaneous vocal activity (hoot/call) of the tagged owls, plus their mates and neighbours (36 untagged owls). After 223 nights in Valle de Mena and 224 in Duranguesado we obtained a total of 8791 records of vocal activity. The annual proportion of surveys on which an owl called was 6.3% and did not differ between the study areas. Vocal activity of Tawny Owls varied with sex, annual cycle stage and weather. Male owls were significantly more vocal than females year‐round, and vocal activity peaked during the incubation and post‐breeding periods. Wind and rain adversely affected vocal activity of both sexes throughout the year. Tagged owls were detected more often than untagged owls, which we interpret as an observer effect because the systematic survey ensured that short distances to tagged owls increased the probability of detecting vocal activity. In fact, 2.8% of variation in vocal activity was due to detectability differences between tagged and untagged owls. We conclude that if fieldwork is carried out during the optimum period of the year for vocal detection (i.e. incubation, which in our case was around mid‐April), and under good weather conditions (dry and calm nights), censuses based on spontaneous vocal activity would detect only approximately 12% of the true Tawny Owl population.  相似文献   

5.
The density of great tit Parus major L. and blue tit Parus caeruleus L. was artificially increased by placing nest-box colonies for these species in the vicinity of the nests of breeding tawny owls during 1993–1997. Bird prey composition in the owl nests, the proportion of parents disappearing from the breeding tit populations and the reproductive performance of the widowed parents were analysed. The frequency of predation on tits by tawny owls was greater in areas where tit density had been artificially increased. Owls preyed more on tits during the feeding period of owlets than during the incubation period and more in years when snow covered the ground during the incubation period than when it did not. Mortality due to predation was male biased and more females lost their mates in populations breeding near tawny owl nests. Reproductive performance of the widowed parents was lower and their body weights were lighter at the end of the nestling period than those found in birds rearing youngs with their mates. Predation by owls increased the between-year turnover in the breeding tit population: widowed parents did not return to the nesting site for the next breeding season.  相似文献   

6.
We studied home range and habitat selection of radio-marked adult California spotted owls (Strix occidentalis occidentalis) randomly selected from among the breeding population of owls in the central Sierra Nevada, California from June to October 2006. The most parsimonious home-range estimate for our data was 555 ha (SE = 100 ha). Home-range size was positively correlated with the number of vegetation patches in the home range (habitat heterogeneity). We used resource selection ratios to examine selection of vegetation types by owls within our study area. Owl home ranges contained a high proportion of mature conifer forest, relative to its availability, although the confidence interval for this estimate overlapped one. We also used resource selection functions (RSF) to examine owl foraging habitat selection. Relative probability of selection of foraging habitat was correlated with vegetation classes, patch size, and their interaction. Owls showed highest selection rates for large patches (>10 ha) of pole-sized coniferous forest. Our results suggested that spotted owls in the central Sierra Nevada used habitat that contained a high proportion of mature conifer forest at the home-range scale, but at a finer scale (foraging site selection) owls used other vegetation classes interspersed among mature forest patches, consistent with our hypothesis that spotted owls may use other forest types besides old growth and mature forests when foraging. Our study provides an unbiased estimate of habitat use by spotted owls in the central Sierra Nevada. Our results suggest that forest managers continue to protect remaining mature and old-growth forests in the central Sierra Nevada because owl home ranges contain high proportions of these habitats. However, our results also showed that owls used younger stands as foraging habitat so that landscape heterogeneity, with respect to cover types, may be an important consideration for management but we did not attempt to relate our findings to fitness of owls. Thus management for some level of landscape heterogeneity for the benefit of owls should proceed with caution or under an adaptive management framework. © 2011 The Wildlife Society.  相似文献   

7.
We examined the temporal, spatial, and demographic factors that influenced roadway mortality of barn owls (Tyto alba) along a 248-km stretch of Interstate 84 in southern Idaho using systematic road surveys. Counts of dead animals from surveys can be underestimated because of sampling biases; therefore, we also conducted experiments to assess the effects of search and removal bias on the estimates of roadway mortality of owls. We conducted surveys every 2 weeks over a 2-year period and detected 812 dead barn owls (unadjusted mortality rate of 1.64 owls/km/yr). After adjusting this estimate for search and removal bias, we documented mortality rates of up to 5.99 owls/km/year. Owl mortality was not random in relation to sex, age class, or location along the highway. Females and juveniles, which represent individuals more likely to disperse long distances, were killed more frequently than males and adults. During the nonbreeding season, owls were killed more often near agricultural lands than in shrub-steppe, but this pattern was not apparent during the breeding season. Owls were also killed more often on portions of the roadway closer to the Snake River canyon, perhaps because of the availability of nest and roost sites. Mortality rates differed markedly between the 2 years of study, which could have been related to variability in weather and its subsequent effect on owl productivity. Our data suggest that barn owls in this region may not persist under this level of mortality without significant immigration or management. Thus, roadway management to reduce or prevent owl use of roadways, reduce rodent populations near major roads, alert motorists to the presence of owls, or otherwise reduce the chances that vehicles and owls collide would improve barn owl survival and population persistence. © 2012 The Wildlife Society.  相似文献   

8.
In the temperate zone, food availability and winter weather place serious constraints on European Barn Owl Tyto alba populations. Using data collected over 22 years in a Swiss population, we analysed the influence of early pre‐breeding food conditions and winter severity on between‐year variations in population size and reproductive performance. To estimate pre‐breeding food conditions, we attempted a novel approach based on an index that combines Tawny Owl Strix aluco reproductive parameters and the occurrence of wood mice Apodemus sp. in their diet. Tawny Owls breed earlier in the season than Barn Owls and are strongly dependent on the abundance of wood mice for breeding. This index was strongly positively associated with the number of breeding pairs and early breeding in the Barn Owl. Winter severity, measured by snow cover and low temperatures, had a pronounced negative influence on the size of the breeding population and clutch size. Food conditions early in the breeding season and winter severity differentially affect the Barn Owl life cycle. We were able to use aspects of the ecology and demography of the Tawny Owl as an indicator of the quality of the environment for a related species of similar ecology, in this case the Barn Owl.  相似文献   

9.
Capsule There is a relationship between owl numbers and the availability of the agri-forest patchwork.

Aims To model habitat preferences at three different scales of two predators largely neglected within the framework of Environmental Impact Assessment (EIA) studies.

Methods We studied habitat preferences of Long-eared Owls and Little Owls by comparing habitat composition around 28 and 78 occupied territories respectively with 55 non-occupied territories in Alicante (eastern Spain). Generalized linear models were used to examine patterns of habitat preference at three different spatial scales: nest-site, home range and landscape.

Results At the nest-site scale, Long-eared Owls preferred wooded areas with few paved roads while Little Owls preferred arid plantations. Furthermore, the probability of finding an occupied territory increased with the proximity of another occupied territory in the surroundings. The home range scale models mirror the feeding requirements of the owls. Thus, Long-eared Owls occupied areas with high percentages of forest, arid plantations, edges between these two land uses, short distances between nests, with presence of conspecifics and little human disturbance. Little Owls occupied arid plantations with high availability of linear structures and the proximity of villages. At the landscape scale, Long-eared Owls eluded extensive forests, and Little Owls preferred arid plantations.

Conclusions We suggest a hierarchical process of habitat selection for both owls regarding fitting trophic resources at the broadest scales and adequate sites for breeding and roosting at the smallest scale. EIA studies must consider that protecting small areas around single nests may not be an efficient conservation option compared with preserving clusters of territories for both species.  相似文献   

10.
Monitoring wildlife populations often involves intensive survey efforts to attain reliable estimates of population size. Such efforts can increase disturbance to animals, alter detection, and bias population estimates. Burrowing owls (Athene cunicularia) are declining across western North America, and information on the relative effects of potential survey methods on owl behaviors is needed. We designed a field experiment to compare burrowing owl flight distances, times displaced, and probabilities of being displaced between 4 potential population survey methods (single walking surveyor, single vehicle stop, single vehicle stop with 2 surveyors, and double vehicle stop with 2 surveyors), and an experimental control in the agricultural matrix of Imperial Valley, California. Between 25 April and 1 May 2008, we randomly applied survey methods to 395 adult male owls during daylight hours (0700 hours through 1900 hours). All survey methods increased odds of displacing owls from perches. Survey methods with observers outside the vehicle were 3 times more likely to displace an owl than a single vehicle stop where observers remained inside the vehicle. Owls were displaced farther distances by all survey methods compared to control trials, but distances and time displaced did not differ among survey methods. We recommend that surveys for counting owls during the breeding season in agroecystems like the Imperial Valley where high densities of owls nest primarily along the borders of fields be conducted using single vehicle stops with or without 2 surveyors, depending on conditions for locating owls from roads. © 2011 The Wildlife Society.  相似文献   

11.
《Ostrich》2013,84(3-4):190-194
We collated the literature available on the breeding activity of the Swainson's Spurfowl Pternistis swainsonii and made use of reliable unpublished reports, nest record cards and field observations within the Springbok Flats, Limpopo Province, South Africa to establish breeding seasons and pairing behaviour. The onset of breeding (egg laying) is closely associated with rainfall, with male gonad development, population density and covey size (pairing behaviour), all correlated with rainfall. Peak breeding activity is from January–April in South Africa, February–May in Zimbabwe and March–June in Botswana. Egg laying has been recorded in all months and sporadic egg laying in the winter months is most likely the result of isolated rainfall. Mean clutch size is 5.2 eggs/hen (n = 140) with an incubation period of 23 days and brood hatching success and chick survival of 69.4% over the southern African sub-region. Current hunting seasons within Limpopo Province are in line with the recommended hunting season for this region and should remain unchanged: 15 June–30 September. The success of this phasianid can be attributed to its extended breeding season, high survival rate of hatchlings and the potential of birds to breed within their first year.  相似文献   

12.
Land conversion for agriculture is an increasing threat to biodiversity conservation, but its ecological effects on African birds is practically unknown. We investigated the impacts of agriculture on the diet and productivity of a small, disjunct population of Mackinder's eagle owls ( Bubo capensis mackinderi  ) in central Kenya. Owl diet was determined by analysis of pellets and other remains and compared to small mammal populations estimated by live trapping in two habitats. Small mammal abundance was low and averaged 7.4 small mammals/ha in farms and 0.5 small mammals/ha in grassland. Owls consumed a wide diversity of prey. The majority were mammals (87%) followed by birds (7%) and insects (5%). The percentage of small mammals in owl diet correlated positively with the relative abundance of small mammals during monthly trapping sessions. Diet composition did not influence owl breeding success. Farming activities affected owl diet composition through crop production. The amount of maize, peas, and carrots growing in farms was correlated with the abundance of Mastomys sp. and Procavia sp. in the owl's diet. Agricultural activities had a large effect on Mackinder's eagle owl diet by increasing the abundance of certain small-mammal prey and attracting owl prey to farms, though farming practices harmful to owls were observed.  相似文献   

13.
In Baranya County (Southern Hungary), tawny owls (Strix aluco) and barn owls (Tyto alba) sequentially use the same nest boxes in a significant number of cases. A total of 460 broods were observed between 1996 and 2003 and, in 12 cases, whole broods of dead tawny owl chicks were registered that had apparently been killed. On investigating the reproductive life characteristics, population sizes, and frequency of killing in these two species, it was concluded that: (1) with growing barn owl population, the number of sequential broods increases but changes in tawny owl population size have no effect on the frequency of sequential broods; (2) the number of killings depends on the number of sequential broods; and (3) with growing barn owl population, the number of killings also grows and this change is unaffected by the size of the tawny owl population. However, no killing occurs as long as 50–60% of the nest boxes are unoccupied. There is no killing either until the percentage of nest boxes occupied by barn owls reaches 40%, although a threshold value like this cannot be shown for the tawny owl. In the cases when a barn owl breeding follows the tawny owl's, the percentage of killing is significantly higher compared to that when barn owls do not breed in the same box. These results indicate that barn owls kill the offspring of tawny owls. By these means, they obtain a breeding place earlier than without killing the chicks of the other species, and this results in higher reproductive success. © 2008 The Linnean Society of London, Biological Journal of the Linnean Society, 2008, 95 , 488–494.  相似文献   

14.
Ecological theory suggests that prey size should increase with predator size, but this trend may be masked by other factors affecting prey selection, such as environmental constraints or specific prey preferences of predator species. Owls are an ideal case study for exploring how predator body size affects prey selection in the presence of other factors due to the ease of analyzing their diets from owl pellets and their widespread distributions, allowing interspecific comparisons between variable habitats. Here, we analyze various dimensions of prey resource selection among owls, including prey size, taxonomy (i.e., whether or not particular taxa are favored regardless of their size), and prey traits (movement type, social structure, activity pattern, and diet). We collected pellets of five sympatric owl species (Athene noctua, Tyto alba, Asio otus, Strix aluco, and Bubo bubo) from 78 sites across the Mediterranean Levant. Prey intake was compared between sites, with various environmental variables and owl species as predictors of abundance. Despite significant environmental impacts on prey intake, some key patterns emerge among owl species studied. Owls select prey by predator body size: Larger owls tend to feed on wider ranges of prey sizes, leading to higher means. In addition, guild members show both specialization and generalism in terms of prey taxa, sometimes in contrast with the expectations of the predator–prey body size hypothesis. Our results suggest that while predator body size is an important factor in prey selection, taxon specialization by predator species also has considerable impact.  相似文献   

15.
The effects of competition and risk of predation on secondary cavity breeders were examined between the 2008 and 2009 breeding seasons using an experimental design manipulating two nest entrance sizes (large entrances allowed Barn Owls (Tyto alba) to enter, while the small entrances excluded them). During the 2009 breeding season, the entrance sizes of nest boxes were exchanged, so that if during one year a nest box in a particular location had a small entrance, the second year it had a large entrance and vice versa. Barn Owls and Eurasian Kestrels (Falco tinnunculus) occupied 67.3 and 17.3%, respectively, of large entrance nest boxes. Significantly more Jackdaws (Corvus monedula), House Sparrows (Passer domesticus) and Scops Owls (Otus scops) bred in nest boxes with small than with large entrances. After nest box entrance sizes were exchanged, Barn Owls and smaller species did not breed in the same nest boxes with the new entrance size. Jackdaws probably did not breed in large entrance nest boxes due both to exploitation competition (Barn Owls and Eurasian Kestrels occupied the majority of large entrance nest boxes), and may also have avoided empty nest boxes because of the risk of interference competition; whereas smaller species may have also avoided large entrance nest boxes due to risk of predation.  相似文献   

16.
Highway programmes typically focus on reducing vehicle collisions with large mammals because of economic or safety reasons, while overlooking the millions of birds that die annually from traffic. We studied wildlife–vehicle collisions along an interstate highway in southern Idaho, USA, with among the highest reported rates of American Barn Owl Tyto furcata road mortality. Carcass data from systematic and ad hoc surveys conducted in 2004–2006 and 2013–2015 were used to explore the extent to which spatial, road geometric and biotic factors explained Barn Owl–vehicle collisions. Barn Owls outnumbered all other identified vertebrate species of roadkill and represented > 25% of individuals and 73.6% of road‐killed birds. At a 1‐km highway segment scale, the number of dead Barn Owls decreased with increasing numbers of human structures, cumulative length of secondary roads near the highway and width of the highway median. The number of dead Barn Owls increased with higher commercial average annual daily traffic (CAADT), small mammal abundance index and grass rather than shrubs in the roadside verge. The small mammal abundance index was also greater in roadsides with grass vs. mixed shrubs, suggesting that Barn Owls may be attracted to grassy portions of the highway with more abundant small mammals for hunting prey. When assessed at a 3‐km highway segment scale, the number of dead Barn Owls again increased, with higher CAADT as well as with greater numbers of dairy farms. At a 5‐km scale, the number of dead Barn Owls increased with a greater percentage of cropland near the highway. Although human conversion of the environment from natural shrub‐steppe to irrigated agriculture in this region of Idaho has probably enhanced habitat for Barns Owls, it simultaneously has increased risk for owl–vehicle collisions where an interstate highway traverses the altered landscape. We review some approaches for highway mitigation and suggest that reducing wildlife–vehicle collisions involving Barn Owls may contribute to the persistence of this species.  相似文献   

17.
P. HANSEN 《Bioacoustics.》2013,22(2):147-154
ABSTRACT

The focus of this study was to determine whether individual vocal identification of Scops Owls Otus scops was possible and if there was a stability of the hoot-calls over a short time period in the same individuals. Spontaneous vocalizations of 13 owls were recorded in 2004 in Southern Tuscany, Italy. Visual analysis of spectrograms and quantitative multivariate analysis of six vocal features showed marked individual differences. In some owls a repertoire of two different hoot types was found. In 2005, 10 Scops owls were recorded three times in the same breeding season (2 hours and 10 days after the first session). Statistical analysis of data showed that 60% of owls did not change call features over time. However a slight but significant variability between successive vocal performances of the same owl was found in 40% of cases. This variability may decrease the recognition power by acoustic analysis. To overcome this obstacle I suggest a multi step qualitative/quantitative approach. A Difference Index (DI) was calculated to set a threshold between the slight intra-individual and the very high inter-individual variability. This method allowed the recognition of calls of each owl recorded over time in 2005.  相似文献   

18.
R. K. Brooke 《Ostrich》2013,84(3):164-169
Crawford, R. J. M., Cooper, J. &; Shelton, P. A. 1982. Distribution, population size, breeding and conservation of the Kelp Gull in southern Africa. Ostrich 53:164:177.

The Kelp Gull Lams dominicanus in Africa occurs coastally between Luanda, Angola and Delagoa Bay, Moçmbique. It breeds between Cape Cross Lagoon, South West Africa/Namibia and Riet River, eastern Cape, South Africa. Censuses of nests and breeding birds at all known southern African breeding localities in the period 1976–1981 indicated that 11 199 pairs bred at 52 localities; 79.5% of this population occurred in South Africa, 57,1% in the Saldanha Bay to Dassen Island region, southwestern Cape. Of the breeding pairs 83% occurred on offshore islands and rocks. Colony size at islands is related to their surface area andMayalso be influenced by food availability and the level of human disturbance. The species breeds in a wide variety of habitats ranging from cliffs and rock stacks to wooden platforms, lowlying vegetation among sand dunes and estuarine sandbars. Any available material is used in the construction of nests, whichMaybe as dense as 4/m2 Clutch size is 2–3 eggs. In 1978 breeding took place earlier in South Africa than in South West Africa/Namibia. 92% of the population breeds m sites which are legally protected. Kelp Gulls have decreased or increased in numbers at some breeding localities but there is no clear overall trend. Any increases in colony size near urban areasMayresult in added airstrike hazards.  相似文献   

19.
Owl predation on snowshoe hares: consequences of antipredator behaviour   总被引:4,自引:0,他引:4  
We show evidence of differential predation on snowshoe hares (Lepus americanus) by great horned owls (Bubo virginianus) and ask whether predation mortality is related to antipredator behaviour in prey. We predicted higher predation on (1) young and inexperienced hares, (2) hares in open habitats lacking cover to protect from owl predation, and (3) hares in above average condition assuming that rich food patches are under highest risk of predation. Information on killed hares was obtained at nest sites of owls and by monitoring hares using radio-telemetry. The availability of age classes within the hare population was established from live-trapping and field data on reproduction and survival. Great horned owls preferred juvenile over adult hares. Juveniles were more vulnerable to owl predation before rather than after dispersal, suggesting that displacement or increased mobility were not causes for this increased mortality. Owls killed ratio-collared hares more often in open than in closed forest types, and they avoided or had less hunting success in habitats with dense shrub cover. Also, owls took hares in above average condition, although it is unclear whether samples from early spring are representative for other seasons. In conclusion, these results are consistent with the hypothesis that variation in antipredator behaviours of snowshoe hares leads to differential predation by great horned owls.  相似文献   

20.
ABSTRACT Competition with barred owls (Strix varia varia) is an important factor contributing to the continued decline of threatened northern spotted owl (Strix occidentalis caurina) populations in the Pacific Northwest, USA, but basic information on habitat selection and space use patterns of barred owls is lacking for much of the region. We investigated space use and habitat selection by tracking radiotagged barred owls in the Eastern Cascade Range of Washington, USA, from 2004 to 2006. We surveyed for barred owls across the 309-km2 study area and confirmed presence of barred owl pairs at 21 sites. We collected movement data on 14 barred owls from 12 sites. Mean annual 95% fixed-kernel home-range size was 194 ha for females (n = 4, SD = 70) and 288 ha for males (n = 5, SD = 114). Home ranges were located more frequently than expected in areas with low topographic position, gentle slopes, large overstory tree-crown diameter, high normalized difference vegetation index (NDVI), overstory tree canopy closure >72%, and a moderate amount of solar insolation. Within home ranges, areas that had large tree-crown diameters, low topographic positions, and gentle slopes were used more frequently than expected. The resource selection function we developed for barred owls in our study area indicated that barred owls used areas with the combination of low values for topographic position and slope and higher values for NDVI, solar insolation, and an interaction term for canopy closure and tree-crown diameter. In comparison to published information on northern spotted owls, barred owls used areas with similar canopy closure and tree size classes, but barred owl home ranges were much smaller and more concentrated on gentler slopes in valley bottoms. This information may contribute to the development of management practices that maintain forest characteristics appropriate for spotted owl habitat and prey in areas where spotted owls are least likely to be excluded by territorial barred owls in the Eastern Cascades of Washington.  相似文献   

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