Integrated magnetobiochronology of the Early/Middle Eocene transition at Agost (Spain): Implications for defining the Ypresian/Lutetian boundary stratotype

In this paper, we present an integrated study of a 115‐m‐thick section that spans the Ypresian/Lutetian boundary at Agost (Betic Cordillera, SE Spain). Our study includes magnetostratigraphic results and biostratigraphic and palaeoenvironmental data derived from planktic foraminifera, small and larger benthic foraminifera, and calcareous nannofossils. Our results demonstrate that the Agost section is continuous and spans from Zones P9 to P12 (E7 to E10), Zones CP11 to CP14a (NP13 to NP16), Zones SBZ11 to SBZ15, and Chrons C22n to C19r. The first occurrence (FO) of H. nuttalli (base of P10) and the FO of G. nuttalli (base of E8) are found within Chron C20r, at a much younger age (3–5 Myr) than previously considered in standard calibration schemes. Similarly, the boundary between SBZ12 and SBZ13 is located within Chron C21n, also at a younger age than previously considered. On the contrary, the FO of B. inflatus (base of CP12b) is found within Chron C21r, which conforms to the magnetostratigraphically calibrated age of ca. 48 Ma (middle part of C21r) considered in standard calibration schemes. These results corroborate earlier studies and indicate that all the events that have been proposed to mark the Ypresian/Lutetian boundary appear at different stratigraphic intervals and have different ages. Based on our results from Agost and on data from other sections elsewhere, we suggest that the Ypresian/Lutetian boundary might be approximated by the FO of B. inflatus (base of CP12b). The Agost section might be considered as a potential candidate to locate the Global Stratotype Section and Point (GSSP) of the base of the Lutetian Stage, because it includes all the events that might be selected as marker events for the Ypresian/Lutetian boundary and it fulfils most of the geological, biostratigraphic and infrastructure requirements demanded for definition of a GSSP.     

Reassessment of the Early–Middle Eocene biomagnetochronology based on evidence from the Gorrondatxe section (Basque Country, western Pyrenees)

Thirteen Lower–Middle Eocene (Ypresian–Lutetian) successions, including the Gorrondatxe section in the western Pyrenees, show biomagnetostratigraphic correlation schemes that do not agree with the current standard framework. The main discrepancy concerns the position of the boundary between planktonic foraminiferal Zones P9 (=E7, approximately) and P10 (=E8, approximately), which was thought to occur within calcareous nannofossil Subzone CP12a and at the boundary between magnetic polarity Chrons C22n and C21r. However, in the differing correlation scheme the boundary between Zones P9 (=E7) and P10 (=E8) occurs close to the base of Subzone CP13a and to the boundary between Chrons C21n and C20r. An attempt at a new Ypresian–Lutetian boundary biomagnetochronology is made based on data from the Gorrondatxe section, which shows that the boundary between Zones P9 (=E7) and P10 (=E8) is 3.1 Myr younger than hitherto considered. Therefore, the duration of the Early Eocene, most commonly defined according to this planktonic foraminiferal zonal boundary, has generally been underestimated over the last four decades.     

Palaeoenvironmental turnover across the Ypresian–Lutetian transition at the Agost section, Southastern Spain: In search of a marker event to define the Stratotype for the base of the Lutetian Stage

Marker events to define the stratotype for the base of the Lutetian Stage are poorly defined. To elucidate such markers and characterize palaeoenvironmental turnovers, we conducted an integrated study of the Ypresian–Lutetian (Y–L; early-middle Eocene) transition at the continuous Agost section (southeastern Spain). This 115-m-thick section, which consists of hemipelagic marls intercalated with hemipelagic limestones and turbidity sandstones, spans from planktic foraminiferal Zones P9 to P12 (E7 to E10) and calcareous nannofossil Zones CP11 to CP14a (NP13 to NP16). We report quantitative analyses of planktic and benthic foraminifera and characterization of trace fossil assemblages that are integrated with mineralogical analyses.Relative to benthic forms, planktic foraminifera constitute more than 80% of the foraminiferal assemblage. We found that the most abundant planktic species belong to the genera Acarinina, Morozovella, Subbotina, and Pseudohastigerina. Benthic foraminiferal assemblages are strongly dominated by calcareous taxa, with bolivinids being the most abundant group. Trace fossils showed the succession Nereites–Zoophycos–Cruziana ichnofacies throughout the Agost section. In addition to changes in palaeobathymetry, we deduced that quantity and quality of organic matter flux influenced by turbidity currents are the main factors controlling benthic assemblages. We distinguished several mineralogical boundaries at the Agost section, each associated with lithological facies changes suggesting a change in provenance rather than changes in weathering conditions. We made three observations that indicate an increase in sea water temperatures or a possible hyperthermal event related to the first occurrence (FO) of hantkeninids (i.e., the P9/P10 boundary): 1) a distinct peak in abundance of the benthic foraminifera Aragonia aragonensis; 2) the low-diversity of benthic foraminiferal assemblages; and 3) the occurrence of the planktic foraminifera Clavigerinella eocenica and Clavigerinella jarvisi. Benthic foraminiferal and trace fossil assemblages also suggest an associated relative fall of sea level from upper-middle bathyal to sublittoral depths. These characteristic indicators point to this boundary as a promising feature for defining the Global Stratotype Section and Point (GSSP) for the base of the Lutetian Stage. However, complementary magnetobiostratigraphic studies carried out at the Agost section point to the FO of calcareous nannofossil Blackites inflatus (base of CP12b), which occurred 3–5 Myr before the P9/P10 boundary, as the most suitable primary marker event. Whatever the marker event chosen, all the successive events recognized at the Agost section allow a complete characterization of the Y–L transition, and thus this section may be a suitable candidate to locate the GSSP for the Ypresian/Lutetian boundary.     

Lutetian calcareous nannofossil events in the Agost section (Spain): implications toward a revision of the Middle Eocene biomagnetostratigraphy

This paper presents a detailed calcareous nannofossil biostratigraphy of the entire Lutetian of the Agost section (Betic Cordillera, SE Spain). This investigation integrates and improves on previous study performed through the Ypresian/Lutetian boundary by the authors on this succession. The new revision of the integrated bio‐magnetobiochronology of the Early/Middle Eocene interval revealed highly diversified calcareous nannofossil assemblages, characterizing more than 8 Myr of climatic variability. The studied interval spans from Zone CP11 to Subzone CP14a and from the upper part of Zone NP13 to the base of Zone NP16 of calcareous nannofossil standard zonations. The revision of the calcareous nannofossil content enabled the identification of numerous secondary events which greatly improved the stratigraphic resolution of this time interval. An important re‐organization of the nannoflora was observed during the Y/L transition, when Reticulofenestra and Dictyococcites (Noelaerhabdaceae) became the most important genera in terms of abundance and dispersal, dominating the Middle Eocene nannofossil assemblages and replacing Toweius and Discoaster taxa characteristic of the lower Eocene. Pentaliths and Blackites experience a great expansion and diversification, whereas Discoaster and Chiasmolithus which are well diversified but never abundant during the Lutetian show a slow turnover. A reassessment of the major bio‐events observed in the Noelaerhabdaceae family as well as revision and correlation of these events with the classical Italian sections (Contessa and Bottaccione) are presented. The new results show that biostratigraphic problems related to the Middle Eocene chronology are not limited to the correlation between calcareous nannofossils and planktonic foraminiferans at the Y/L transition but extend to calcareous nannofossil events commonly used for correlating the Bartonian.     

Integrated stratigraphy and astronomical calibration of the Serravallian/Tortonian boundary section at Monte Gibliscemi (Sicily, Italy)

Results are presented of an integrated stratigraphic (calcareous plankton biostratigraphy, cyclostratigraphy and magnetostratigraphy) study of the Serravallian/Tortonian (S/T) boundary section of Monte Gibliscemi (Sicily, Italy). Astronomical calibration of the sedimentary cycles provides absolute ages for calcareous plankton bio-events in the interval between 9.8 and 12.1 Ma. The first occurrence (FO) of Neogloboquadrina acostaensis, usually taken to delimit the S/T boundary, is dated astronomically at 11.781 Ma, pre-dating the migratory arrival of the species at low latitudes in the Atlantic by almost 2 million years. In contrast to delayed low-latitude arrival of N. acostaensis, Paragloborotalia mayeri shows a delayed low-latitude extinction of slightly more than 0.7 million years with respect to the Mediterranean (last occurrence (LO) at 10.49 Ma at Ceara Rise; LO at 11.205 Ma in the Mediterranean). The Discoaster hamatus FO, dated at 10.150 Ma, is clearly delayed with respect to the open ocean. The ages of D. kugleri first and last common occurrence (FCO and LCO), Catinaster coalitus FO, Coccolithus miopelagicus last (regular) occurrence (L(R)O) and the D. hamatus/neohamatus cross-over, however, are in good to excellent agreement with astronomically tuned ages for the same events at Ceara Rise (tropical Atlantic), suggesting that both independently established timescales are consistent with one another. The lack of a reliable magnetostratigraphy hampers a direct comparison with the geomagnetic polarity timescale of Cande and Kent (1995; CK95), but ages of calcareous nannofossil events suggests that CK95 is significantly younger over the studied time interval. Approximate astronomical ages for the polarity reversals were obtained by exporting astronomical ages of selected nannofossil events from Ceara Rise (and the Mediterranean) to eastern equatorial Pacific ODP Leg 138 Site 845, which has a reliable magnetostratigraphy.Our data from the Rio Mazzapiedi–Castellania section reveal that the base of the Tortonian stratotype corresponds almost exactly with the first regular occurrence (FRO) of N. acostaensis s.s. as defined in the present study, dated at 10.554 Ma. An extrapolated age of 11.8 Ma calculated for the top of the Serravallian stratotype indicates that there is a gap between the top of the Serravallian and the base of the Tortonian stratotype, potentially rendering all bio-events in the interval between 11.8 and 10.554 Ma suitable for delimiting the S/T boundary. Despite the tectonic deformation and the lack of a magnetostratigraphy, Gibliscemi remains a candidate to define the S/T boundary by means of the Tortonian global boundary stratotype section and point (GSSP).     

High-resolution intra- and interbasinal correlation of the Danian-Selandian transition (Early Paleocene): The Bjala section (Bulgaria) and the Selandian GSSP at Zumaia (Spain)

The Danian-Selandian (D-S) boundary has been identified for the first time in the Black Sea coast at Bjala (Bulgaria) based on a new integrated bio-, magneto- and cyclostratigraphic study. Several correlation criteria as established for the basal Selandian GSSP from Zumaia (Basque Basin) are evaluated. Noteworthy, is the almost complete lack of calcareous nannoplankton species Braarudosphaera bigelowi in the Bulgarian sections, a sharp decrease of which was indicated as suitable criteria for defining the D-S boundary as it occurred both at Zumaia and in the classical locations of the North Sea basin. Conversely, the second evolutionary radiation of the calcareous nannofossil genus Fasciculithus together with the occurrence of Fasciculithus tympaniformis that define the NP4/NP5 zonal boundary seem to be reliable criteria to approximate the D-S boundary. In detail, however, the best approach is to integrate biostratigraphic data within a magnetostratigraphic and/or cyclostratigraphic framework. Refinements on the placement of chron C27n at Zumaia and robust bed-by-bed correlation between several Basque sections and Bjala indicates that the D-S boundary is located 30 precession cycles (~ 630 ky) above C27n. In addition to the precession-related marl-limestone couplets and 100-ky eccentricity bundles recognized in the studied sections, expression of the stable 405-ky long eccentricity allows direct tuning to the astronomical solutions. A correlation of the land-based sections with previously tuned data from ODP Site1262 from the Southern Atlantic is challenged. Our choice is consistent with original tuning at Zumaia but shifts one 100-ky cycle older previous tuning from Site 1262 along the interval above C27n. Under the preferred tuning scheme the D-S boundary can be given an age of 61.641 ±0.040 Ma on the La04 orbital solution.     

Correlation between the Paleocene/Eocene boundary and the Ilerdian at Campo, Spain

The Ilerdian is a well-established Tethyan marine stage, which corresponds to an important phase in the evolution of larger foraminifera not represented in the type-area of the classical Northwest-European stages. This biostratigraphic restudy of its parastratotype in the Campo Section (northeastern Spain) based on planktic foraminifera, calcareous nannofossils, dinoflagellate cysts and the distribution of the stable isotopes ∂¹³C and ∂¹⁸O is an attempt to correlate the Paleocene/Eocene boundary based on a characteristic carbon isotope excursion (CIE) marking the onset of the Initial Eocene Thermal Maximum (IETM) and the Ilerdian stage. The base of this ∂¹³C excursion has been chosen as the criterion for the recent proposal of the Global Stratotype Section and Point (GSSP) of the base of the Eocene (= base of the Ypresian) in the Dababiya Section (Egypt) to which an age of 54.9 Ma has been attributed. This level is also characterized by a marked extinction among the deep-water benthic foraminifera (Benthic Foraminifera Extinction Event, BFEE), a flood of representatives of the planktic foraminiferal genus Acarinina and the acme of dinoflagellate cysts of the genus Apectodinium. In the Campo Section, detailed biozonations (planktic foraminifera, calcareous nannofossils, dinoflagellate cysts) are recognized in the Lower and Middle Ilerdian. The correlation with the Ypresian stratotype is based on dinoflagellate cysts and calcareous nannofossils. The base of the Ilerdian is poor in planktic microfossils and its precise correlation with the redefined Paleocene/Eocene boundary remains uncertain.     

Filling the North European Early/Middle Eocene (Ypresian/Lutetian) boundary gap: Insights from the Pyrenean continental to deep-marine record

The Early/Middle Eocene (Ypresian/Lutetian) transition is represented by a hiatus in many North European sections, including those in which the classic stratotypes were originally defined. However, the Global Stratotype Section and Point of the Lutetian Stage, which is still pending definition, should be placed at a globally correlatable event included within that unrepresented interval. The Pyrenean Eocene outcrops display sedimentary successions that offer the rare opportunity to analyse the Ypresian/Lutetian boundary interval in almost continuous sections and in very different settings. Seven reference stratigraphic sections were selected on the basis of their quality and correlated by means of biomagnetostratigraphic data. This correlation framework casts light on the sequence of chronostratigraphic events that characterize the Ypresian/Lutetian boundary interval, which may prove useful in defining the main correlation criterion of the base of the Lutetian.All of the Pyrenean sections show a similar sedimentary evolution, despite being up to 350 km apart from each other, containing deposits of different origins (intrabasinal carbonate sediments, siliciclastic sediments sourced from the Iberian plate, and terrigenous sediments sourced from the uplifting Pyrenees) and despite having been accumulated in different sedimentary environments (from continental to deep marine) and in different geodynamic settings (piggy-back basin, foreland basin and cratonic margin). This common evolution can be readily interpreted in terms of a sea-level driven depositional sequence whose lowstand and transgressive systems tracts are included within the Ypresian/Lutetian boundary interval. The Pyrenean Ypresian/Lutetian depositional sequence can reasonably be correlated with depositional sequences from classic North European areas, shedding light on the palaeoenvironmental history which in those areas has not been recorded. Furthermore, these depositional sequences may possibly correlate with others from the Antarctic Ocean and from New Jersey, as well as with oceanic temperature variations, suggesting that they might be the result of climatically-driven glacioeustatic sea-level changes. Should this hypothesis prove correct, it would confirm previous suggestions that the onset of Antarctic glaciations needs to be backshifted to the late Ypresian at least.     

Where and when the earliest coccolithophores?

Gardin, S., Krystyn, L., Richoz, S., Bartolini, A. & Galbrun, B. 2012: Where and when the earliest coccolithophores? Lethaia, Vol. 45, pp. 507–523. New calcareous nannofossil analyses from the Northern Calcareous Alps of Austria are herein used to update and improve the state of knowledge about the oldest occurrence of coccolithophores reported in the literature. Previously reported Norian occurrences of coccoliths were based on an obsolete Triassic chronostratigraphy, in which the Rhaetian stage was subsumed into the Norian (‘Sevatian 2’). The oldest stratigraphical record of coccoliths spp. lies just below the Norian‐Rhaetian boundary and the first coccolith species, Crucirhabdus minutus, is recorded from the base of Rhaetian stage. The latter bio‐event is located just above the First Occurrence of the conodont Misikella posthersteni and the first occurrence of the ammonoid Paracochloceras suessi in the Steinbergkogel section (Austria), Global Stratotype Section and Point (GSSP) candidate for the Norian–Rhaetian boundary. The appearance of the coccolith Crucirhabdus minutus is seen as a robust biochronological datum that will provide useful constraints for Triassic biostratigraphy, palaeoclimatic modelling and phylogenetic reconstructions. The new calcareous nannofossil biochronology of Steinbergkogel that we present herein completes the existing biostratigraphic characterization of the Norian‐Rhaetian transition based on conodonts and ammonoids and strengthens the position of Steinbergkogel as the best GSSP proposed section for the base of the Rhaetian. The record of coccolithophores across the Norian–Rhaetian boundary at Steinbergkogel takes place along with a discernible increase in abundance of Prinsiosphaera triassica, as well as the appearance of Euconusphaera zlambachensis, which are the two most important Rhaetian pelagic carbonate producers. This succession of bio‐events is interpreted as the initiation of the pelagic carbonate production driven by the successful spreading of calcareous nannofossils in the Western Tethys during the Rhaetian. □Austria, Calcareous Alps, coccolithophores, Triassic.     

High resolution calcareous nannofossil biostratigraphy and paleoecology across the Latest Danian Event (LDE) in central Eastern Desert,Egypt

A detailed study of the calcareous nannofossil assemblages of latest Danian to early Selandian age from Wadi Qena (the Qreiya-1, Qreiya-2, and Araas sections) in the central Eastern Desert and the Gebel Duwi section at the Red Sea Coast has been carried out to understand the local phytoplankton paleocommunities. For the first time in Egypt four calcareous nannofossil zones (NTp6, NTp7, NTp8, NTp9) and five subzones (NTp7A, NTp7B, NTp8A, NTp8B and NTp8C) were distinguished subdividing the standard NP4 and NP5 zones. The calcareous nannofossil zones were correlated with nannofossil zonations from the Tethyan region (e.g. Tunisia and the Zumaia section, the GSSP for the Danian/Selandian boundary). The calcareous nannofossil assemblages are moderately to well preserved. The event beds are characterized by a dark-brown shaley-marl bed rich in organic carbon and coprolites. Three main assemblages are distinguished: a pre-event assemblage, a transitional assemblage and a post-event assemblage. The paleoenvironments vary from cool eutrophic water below and above the event to warm oligotrophic conditions during the event. The result of this study concludes that the “Latest Danian Event” (LDE) may represent an early Paleocene hyperthermal.     

Re‐evaluation of the conodont Iapetognathus and implications for the base of the Ordovician System GSSP

Terfelt, F, Bagnoli, G. & Stouge, S. 2011: Re‐evaluation of the conodont Iapetognathus and implications for the base of the Ordovician System GSSP. Lethaia, Vol. 45, pp. 227–237. In 2000, the International Union of Geological Sciences (IUGS) ratified the decision from the International Working Group on the Cambrian–Ordovician Boundary (COBWG) to place the Global boundary Stratotype Section and Point (GSSP) for the base of the Ordovician System in the Green Point section, Newfoundland, Canada, at a point coinciding with the first appearance of the conodont Iapetognathus fluctivagus. However, a restudy of the conodont successions from Green Point shows that this species is not present at the boundary interval, and as a consequence the section does not fulfil the biostratigraphical requirements of a GSSP. The GSSP horizon as now defined is based on a level part‐way through the range of I. preaengensis– a species with lower first appearance datum (FAD). The true FAD of I. fluctivagus is above the FAD of planktonic graptolites and well above the FAD of I. preaengensis. As a consequence of these problems, a restudy of the GSSP section and the other sections in the Cow Head Group is necessary. A redefinition of the GSSP horizon is suggested. The following four alternative horizons have potential as new horizons for the GSSP level: the FAD of Cordylodus intermedius; the FAD of Cordylodus andresi; the FAD of Eoconodontus notchpeakensis; and the FAD of the agnostoid Lotagnostus americanus. □Boundary, Cambrian, conodont, Global boundary Stratotype Section Point, Iapetognathus, Ordovician.     

Distribution patterns of benthic foraminifera across the Ypresian–Lutetian Gorrondatxe section,Northern Spain: Response to sedimentary disturbance

We present a study of benthic foraminiferal assemblages from an Ypresian–Lutetian distal submarine fan system in the lower bathyal Gorrondatxe section (Basque-Cantabrian Basin, northern Spain). The objective of our study is to analyze the benthic foraminiferal distribution patterns and their response to sedimentary disturbance and related factors.Assemblages contain a high percentage of allochthonous taxa, such as asterigerinids and other shallow water taxa, which were transported downslope by turbidity currents.Detailed quantitative analyses, supported by R-mode cluster and Detrended Correspondence Analyses (after removing allochthonous taxa from the foraminiferal counts) allowed us to identify 6 assemblages that are divided into two groups related to the turbidite content in the Gorrondatxe section. Assemblages 1, characteristic of the turbidite-poor intervals with low sedimentary disturbance, include assemblage 1a (with highly diverse common middle–lower bathyal calcareous taxa) assemblage 1b (with common agglutinated taxa, mainly trochamminids), and assemblage 1c (characterized by calcareous taxa that are also common in the turbidite-rich interval).Assemblages 2, characterized by a high dominance, prevail in the turbidite-rich interval, and include assemblage 2a (characterized by the dominance of infaunal bolivinids and epifaunal cibicids), assemblage 2b (typified by moderate to low diversity and dominated by deep-infaunal Globobulimina species), and assemblage 2c (typified by very abundant suspension-feeding astrorhizids). The high abundance of bolivinids and Globobulimina species may be related to an enhanced input of low-quality organic matter transported by turbidity currents to the seafloor, representing different stages of recolonisation after disturbance and different energy regimes. High current activity was probably responsible for the abundance of cibicids, while moderate to low diverse and high dominance assemblages characterize the recolonisation of the substrate after disturbance.We conclude that sedimentary disturbance and other related factors such as current activity, resuspension of sediments at the seafloor, and supply of organic matter (and its quality) played an important role in the distribution of benthic foraminifera in the Gorrondatxe section. The identification of allochthonous taxa emerges as an essential aspect of the study of environments with sedimentary disturbance.     

Effects of diagenesis on the astrochronological approach of defining stratigraphic boundaries in calcareous rhythmites: The Tortonian GSSP

Since the establishment of astrochronology, calcareous rhythmites are frequently used as the basis of high‐resolution chronostratigraphy. In particular for the Neogene, calcareous rhythmites serve as stratotypes and for absolute dating of stratigraphic boundaries (Global Stratotype Sections and Points – GSSPs). However, the exact mechanisms responsible for the formation of the rhythmic intercalation of lithologies in such successions are complex and not easily reconstructed. To a large extent this is the effect of diagenetic modifications of the original sediment. Here, two examples of Neogene calcareous rhythmites are studied; the Monte dei Corvi section and the Monte Gibliscemi section. The first is the GSSP location of the Serravallian–Tortonian boundary, whereas the second is an auxiliary stratotype for the same boundary. During the past years, astrochronological approaches were applied to these successions to considerably increase time resolution compared to the elaborated biostratigraphic database. The present study focuses on micropetrographic, trace element and clay mineralogical methods in order to gain a better understanding of the genesis of the rhythmites. In the Monte Gibliscemi section, sediment parameters that are robust against diagenetic change clearly reflect primary differences, i.e. cyclical environmental changes. In contrast, no clear primary signal is determined for the Monte dei Corvi section on a couplet scale, leaving the origin of the rhythm ambiguous. This impedes the interpretation of the latter and the comparability between the two successions, and also compromises any bed‐by‐bed correlation between the two. The unclear origin of the rhythmites of Monte dei Corvi introduces uncertainty into the applicability of astrochronology to this succession.     

Late Bashkirian and early Moscovian conodonts from the Naqing section,Luodian, Guizhou,South China

Late Bashkirian and early Moscovian conodonts are abundant and diverse at the Naqing section, South China. All conodont genera known to have numerous species in the late Bashkirian–early Moscovian are recorded here, including Declinognathodus, Diplognathodus, Gondolella, Idiognathodus, Idiognathoides, Mesogondolella, Neognathodus, and Neolochriea. At Naqing, many species of these genera provide a succession of conodont chronomorphoclines throughout the Bashkirian–Moscovian boundary interval. They demonstrate that deposition was remarkably continuous through the boundary interval, a major criterion for selecting a Global Stratotype Section and Point (GSSP). This paper describes the current state of knowledge for several of these chronomorphoclines, and also provides an updated range chart of conodonts recovered from the Naqing section and their correlation with other regions.The taxon that best matches the current concept for the base of the Moscovian Stage in its type region is the phylogenetic first occurrence of Diplognathodus ellesmerensis. An ancestral form with most of the characteristics of D. ellesmerensis occurs at Naqing. More specimens are needed to completely document the chronomorphocline, but because D. ellesmerensis is found worldwide — including that close to the base of the type Moscovian — its evolutionary first occurrence would provide an almost ideal GSSP to define the base of the international Moscovian Stage.     

Planktonic foraminiferal succession in late Cretaceous to early Palaeocene strata in Meghalaya, India

A roughly 10.5-m-thick succession within the Langpar Formation of the Um Sohryngkew River section, Meghalaya, India, constrained by the last occurrence of Globotruncanita stuarti and the first occurrence of Parasubbotina pseudobulloides , spans the K/T (Cretaceous–Tertiary) transition. The unit is divisible into three parts with the lower consisting of shaly limestone, weakly calcareous shale and silty shale with coal streaks. The middle part is dominated by calcareous shale with mud flakes, coprolites, burrows and pyrite nodules, followed by alternating limestone and marlite at the top. Planktonic foraminifera are rare to frequent within the unit. Based on the distribution of zonal indices, seven successive planktonic foraminiferal zones are recognized from across the K/T boundary. From base to top, these are CF4, CF3, CF2 and CF1 in the upper Maastrichtian part and Zone P0, Zone Pα and Subzone P1a in the lower Danian part. The biozones indicate that the section is biostratigraphically continuous across the K/T boundary. A similar foraminiferal succession and K/T transition is observed in the Langpar of the Cherrapunji-Mahadeo road section at a distance of over 5km. These K/T outcrops from Meghalaya provide the first record of a continuous K/T sequence in the Indian subcontinent with respect to planktonic foraminifera.     

A proposed new global stratotype for Aeronian Stage of the Silurian System: Hlásná Třebaň section,Czech Republic

The current Global Stratotype Section and Point (GSSP ) for the Aeronian Stage (Llandovery Series, Silurian System), on Trefawr track in the Llandovery area of Wales, is an inadequate marker for precise, global, correlation. The International Subcommission on Silurian Stratigraphy has, therefore, undertaken the selection of a new GSSP for this level. The lowest occurrence of the graptolite Demirastrites triangulatus , 1.38 m above the base of the black‐shale succession of the ?elkovice Formation at the Hlásná T?ebaň section in Central Bohemia, is proposed to mark the base of the Aeronian Stage. The section, which fulfils all formal requirements for stratotype of a chronostratigraphical unit, should be considered as a candidate for the new GSSP . An abundant, well‐preserved, diverse graptolite fauna occurs through the section along with common chitinozoans, which indicate that the Spinachitina maennili Biozone spans the boundary interval. The section comprises the lower–middle Aeronian (D. triangulatus–Lituigraptus convolutus graptolite biozones) along with underlying Rhuddanian (Akidograptus ascensus–Coronograptus cyphus biozones) and Hirnantian strata. Several graptolite genera of primary importance in global correlation (Demirastrites , Petalolithus , Rastrites and Campograptus ) first appear in the lower part of the triangulatus Biozone. The structurally simple section is somewhat condensed, but there is a uniform succession of black shale without any evidence of disconformity in the broad boundary interval. The C_org isotope record exhibits a minor positive excursion just above the base of the triangulatus Biozone, whereas TOC and N isotope and elemental geochemical records provide evidence for uninterrupted sedimentation under stable, anoxic conditions.     

Late Cretaceous-Eocene nannofossil and magnetostratigraphic correlations near Gubbio,Italy

Using a modified sample preparation technique, we have been able to establish a detailed lower Campanian to upper Eocene nannofossil stratigraphy in the Bottaccione and Contessa Highway sections near Gubbio. Appearance and extinction levels of virtually all the commonly used calcareous nannofossil zonal markers have been recognized and can now be closely correlated with the planktonic foraminifera zonation and the magnetic reversal stratigraphy previously established in these sections. Comparisons with the nannofossil calibrations of the oceanic magnetic anomaly sequence in Deep Sea Drilling Project (DSDP) sites suggest that magmetic Subchrons C17N and C25N are missing in the Bottaccione section. The observed variability of the relative stratigraphic position of most plankton events is confirmed to less than one magnetic subchron. Absolute abundance, paleobiogeographic restriction, and differential preservation render some of the traditionally used biostratigraphic events less reliable than others.     

Calcareous nannofossil biostratigraphy of the M. del Casino section (northern Apennines,Italy) and paleoceanographic conditions at times of Late Miocene sapropel formation

A detailed quantitative calcareous nannofossil analysis has been performed on 138 samples from the astronomically dated Monte del Casino section with the aim to identify and precisely date the most important calcareous nannofossil events across the Tortonian/Messinian boundary in the Mediterranean, and to unravel paleoceanographic conditions at times of sapropel formation during the Late Miocene. From the biostratigraphic perspective, the genus Amaurolithus provides three successive first occurrences (FOs): A. primus, A. cf. amplificus and A. delicatus, dated at 7.446, 7.434 and 7.226 Ma, respectively. Other bioevents include the base and top of the `small reticulofenestrids' Acme, dated at 7.644 and 6.697 Ma, and the FO, FCO and LO of R. rotaria, dated at 7.405, 7.226 and 6.771 Ma. These events appear to be useful in improving biostratigraphic resolution in the Tortonian–Messinian boundary interval, at least for the Mediterranean. Quantitative analysis revealed changes in the calcareous nannofossil assemblage associated with the sapropels. The observed fluctuations suggest a single mechanism for sapropel formation in the Mediterranean during the late Neogene. Sapropels are characterized by a decrease in the total number of coccoliths, interpreted mainly as a reduction in calcareous nannofossil production due to increased siliceous plankton production during spring blooms; and an increase in reworked specimens, interpreted to reflect enhanced continental input via river run-off. An increase in abundance of the genus Rhabdosphaera can be explained by opportunistic behavior at the end of the spring bloom when nutrient levels start to become impoverished. As far as sea surface water temperature indicators are concerned, warm water D. pentaradiatus shows positive fluctuations in sapropels while cooler water D. intercalaris and C. pelagicus show negative fluctuations.     

Sea-level dynamics and palaeoecological factors affecting trace fossil distribution in Eocene turbiditic deposits (Gorrondatxe section,N Spain)

Ichnological analysis of the upper Ypresian–lower Lutetian interval at the Gorrondatxe section (W Pyrenees, N Spain), reveals a relationship between sea-level dynamics and the eco-sedimentary factors influencing trace fossil assemblages. The 600 m thick section of deep-sea turbiditic deposits contains 41 ichnospecies belonging to 28 ichnogenera, which are typical of the Nereites ichnofacies, and mostly of the Paleodictyon ichnosubfacies, suggesting deposition in a basin plain to fan-fringe setting. The trace fossil diversity and abundance fluctuate, irrespective of turbidite frequency. These ichnological features are strongly affected by trophic level changes related partly to sea-level dynamics according to the sequence stratigraphic interpretations for the studied section. Temperature, oxygenation and substrate changes are also considered as relevant factors. Increased ichnodiversity, particularly among graphoglyptids, coincides with moderate oligotrophy and stable ecological conditions. Eutrophisation, lowered oxygenation and drop of temperature, typical of low sea level, can reduce ichnodiversity.     

Paleogene terrigenous (flysch) sequences in Etoloakarnania region (W. Greece). Plankton stratigraphy and paleoenvironmental implications

Based on calcareous nannofossils and planktonic foraminifera1 biostratigraphic data from flysch sequences, we give evidence for the paleoenvironmental evolution of Gavrovo and Ionian foreland basins (External Hellenides, Etoloakarnania region). Our data suggest that the onset of clastic sedimentation in both foreland basins in the study area is chronostratigraphically placed at Late Eocene (from 36.2-34.4 Ma; nannofossil biozones NP19-20, planktonic foraminifera biozones P16-17). During the earliest Oligocene (NP21-22 nannofossil biozones/34.4-32.45 Ma), both basins represent restricted accumulation of sediments, mainly composed of clays and silts. The presence of thick flysch deposits, accumulated during Early Oligocene (33.4-30 Ma, nannofossil zone NP23), indicates an increasing rate of sediment supply. The flysch sequences in the Ionian basin are associated with a distal depositional environment, while in the same time the sedimentation in the external part of Gavrovo basin is related to a more proximal environment that is gradually deepening. On the contrary, the internal part of Gavrovo basin is characterised by deep-water facies, deposited in the Early Oligocene. At the end of Early Oligocene and the onset of Late Oligocene (nannofossil zone NP24/30-27.2 Ma, planktonic foraminifera zone P21), the deposition of coarse grained sediments in both basins indicates a shift to shallower depositional environment. The accumulation of fine-grained sediments during Late Oligocene (27.2-23.2 Ma, NP25 nannofossil biozone) in the Ionian basin marks the youngest flysch sediments in the Etoloakarnania region and specifies the time of the Gavrovo nappe emplacement on the Ionian zone. Moreover the emplacement of Pindos nappe on the Gavrovo zone is estimated between 30-27.2 Ma (NP24 biozone) as supported by the nannofossil analysis of samples in front of Pindos thrust.