Matière organique sédimentaire et paléoenvironnements du Tithonien inférieur quercynois (France)

The Solen 98 well corresponds to the limestones of the Lower Tithonian Cazals Formation (Gigas Zone). The iterative succession of six sedimentary terms expresses a cyclic peritidal dynamic. Limited by two emersion surfaces, each sequence evolves from an upper subtidal lagoon to a tidal flat, upper intertidal or supratidal environment, and ends with open sea depositional bioclastic and oolitic shoals deposits. Hierarchical ascendant classification applied to palynological data define 6 palynofacies types associated with different depositional facies. Type 1, characterizing open marine deposits, shows a diversified and balanced assemblage. The blade-shape woody particles are abundant and the amorphous organic matter is absent. Types 2 and 3 are linked to lagoonal and skeletal shoals deposits. Then microfossil population is dominated by Corculodinium or long-spine Micrhystridium. The Shannon-Weaver and the equitability indices are moderate. Type 4 is associated with the upper tidal flat, lagoonal and skeletal shoal deposits. When microfossils are present, the algal assemblages are more balanced than in type 5. This type, observed in all the palaeo-environments except the open marine, is enriched in elements attributed to the Hyalinsphaeridia complex. The marine component assemblages are balanced. The amorphous organic matter is relatively abundant and the oxydized woody particles absent. Type 6, mainly composed of amorphous organic matter and phytoclasts, is principally associated with the stromatolitic facies of tidal flat deposits. The example of the Solen 98 well, shows that hierarchical ascendant classification method is well suited for identification of palynofacies     

Decouverte d'Ammonites du Lias inferieur et moyendans la zone predorsalienne de la region de Jebha (Rif,Maroc) et ses consequences

Jurassic from rifean “Dorsale” furnished only rare paleontological documents; so it's interesting to point outhere, in the predorsalian zone, associations of ammonites from Domerian (Fuciniceras, Protogrammoceras), from Sinemurian (Charmasseiceras, Epammonites) and from Hettangian (Waehneroceras), this one beeing in this way well characterized for the first time in this zone.     

Nouveaux repères biostratigraphiques à terebratulidés (Brachiopodes) dans le Tithonien inférieur ouest-européen

The discovery of a new species of Terebratulid (Xestosina sera nov. sp.) in the Lower Tithonian allows to specify the biostratigraphical subdivision of the Gigas Zone. This also allows to improve the correlations, when ammonites are lacking, among the Tithonian deposits of Aquitaine, the northern Jura and the eastern border of the Paris Basin. The critical revision of the Xestosina genus gives a nomenclatural setting argued to the index-species of biozones. It constitutes a brachiopods based parallel scale for the Kimmeridgian and Tithonian Stages.     

Sur l’origine bactérienne et fongique de la pigmentation de l’Ammonitico Rosso (Jurassique, région de Vérone, Italie du nord)

Fourteen sections in the Ammonitico Rosso Veronese (Callovian to Tithonian, Trento altipiano) disclose the presence of diverse facies ranging from pelagic to outer platform. In spite of this diversity, red limestones are present at different levels. Many microfacies are similar to those observed in other Paleozoic and Mesozoic red carbonates with an abundance of hematitic bioconstructions. We therefore postulate that the origin of the pigmentation is similar in all the studied cases and due to the activity of iron-oxidizing bacteria. Nevertheless, 2 notable differences are observed: the presence in the Ammonitico Rosso of manganese and the existence of in situ bacterial-fungal mats in the matrix. These “algal” mats can represent up to 20% of the sediment. Their excellent preservation (absence of packing down or crushing) is due to the slow sedimentation rate of the pelagic sediments or of the hardgrounds.     

Les formations du jurassique supérieur du Bou Rheddou au Nord de Tiaret (Bordure sud-Tellienne,Algérie): âge et milieux de dépôt

In the Bou Rheddou mountain (Southern Tellian border), new biostratigraphic data based on Ammonites, allow precise and modify the age of the Arkell & Busson formations. The Oxfordian «Ammonitico rosso facies begin in the Transversarium zone and go into the Planula zone. This zone is characterized for the first time in this region. Lower Tithonian (Hybonotum zone) and Upper Tithonian (Microcanthum zone) are also recognized. Precisions on depositional environment and paleogeography are given. Facies and low sedimentation show a pelagic shoal (abundant planctonic organisms) slightly subsiding and lately recovered by the Jurassic transgression. During Oxfordian, episodic interruption of the «Ammonitico rosso deposits shows an unstable period with terrigenous deposits (intra-oxfordian tectonic movements). The facies of the Northern-Gondwanian carbonated platform begin earlier in the Bou Rheddou zone (Oxfordian-Kimmeridgian boundary) than in the Southern regions, where the same facies appears only in Upper Kimmeridgian.     

Facies mosaic in the inner areas of a shallow carbonate ramp (Upper Jurassic,Higueruelas Fm,NE Spain)

The internal facies and sedimentary architecture of an Upper Jurassic inner carbonate ramp were reconstructed after the analysis and correlation of 14 logs in a 1 × 2 km outcrop area around the Mezalocha locality (south of Zaragoza, NE Spain). The studied interval is 10–16 m thick and belongs to the upper part of the uppermost Kimmeridgian–lower Tithonian Higueruelas Fm. On the basis of texture and relative proportion of the main skeletal and non-skeletal components, 6 facies and 12 subfacies were differentiated, which record subtidal (backshoal/washover, sheltered lagoon and pond/restricted lagoon) to intertidal subenvironments. The backshoal/washover subenvironment is characterized by peloidal wackestone–packstone and grainstone. The lagoon subenvironment includes oncolitic, stromatoporoid, and oncolitic-stromatoporoid (wackestone and packstone) facies. The intertidal subenvironment is represented by peloidal mudstone and packstone–grainstone with fenestral porosity. Gastropod-oncolitic (wackestone–packstone and grainstone) facies with intercalated marl may reflect local ponds in the intertidal or restricted lagoon subenvironments. Detailed facies mapping allowed us to document 7 sedimentary units within a general shallowing-upward trend, which reflect a mosaic distribution, especially for stromatoporoid and fenestral facies, with facies patches locally more than 500 m in lateral extent. External and internal factors controlled this heterogeneity, including resedimentation, topographic relief and substrate stability, combined with variations in sea-level. This mosaic facies distribution provides useful tools for more precise reconstructions of depositional heterogeneities, and this variability must be taken into account in order to obtain a solid sedimentary framework at the kilometer scale.     

Biostratigraphic distribution of ammonites and calpionellids in the Tithonian of the internal Prerif (Msila area,Morocco)

Because of the relatively thick succession exposed in the Msila area, new bed-by-bed sampling has been carried out in order to investigate late Tithonian ammonites and the correlation between biostratigraphic scales based on ammonites and calpionellids. The outcrop begins with Lower Tithonian beds assigned to the Fallauxi Zone. Ammonites have been found that allow recognition in the same section both Fallauxi and Ponti Zones. The most significant species are: Simoceras admirandum Zittel, which is the marker of the upper subzone of the Fallauxi Zone, i.e. the Admirandum/Biruncinatum Subzone, and several species of the genus Lemencia and Burckhardticeras peroni (Roman), that characterize the Ponti Zone. Ammonites that belong to the Microcanthum Zone appear in bed 18. This zone is usually subdivided into two subzones, of which only the lower one, the Simplisphinctes Subzone, has been recognized. The upper Transitorius Subzone has been recognized on the basis of the occurrence of Moravisphinctes fischeri (Kilian). The last ammonite unit, the Durangites Zone, is easily recognizable because of the occurrence of species of the genera Durangites and Protacanthodiscus. The section ends with the beginning of the Berriasian, which is characterized by a well-exposed, ammonite-rich bed of the Jacobi Zone. Correlation of the ammonite zones with calpionellid zones has been investigated. The Chitinoidella Zone is characterized by calpionellids with microgranular tests that appear in the ammonite Admirandum/Biruncinatum Subzone with species of the Dobeni Subzone, which extends up to the Ponti Zone. Representatives of the Boneti Sub-Zone appear in bed 16, which probably correlates with the base of the ammonite Upper Tithonian Microcanthum Zone. Two horizons have been distinguished within the calpionellid Subzone A3.     

Précisions biostratigraphiques nouvelles sur le Berriasien des Djebel Nara et Sidi Kralif (Tunisie centrale)

The biostratigraphy of Sidi Kralif formation is precised by means of correlation of calpionellid, ostracod and ammonite faunas in three sections of Nara Mountain (Central Tunisia). The Upper Tithonian (Jacobi zone pars) and the Lower Berriasian (Grandis zone) show 40 m of pelagic sediments above the Nara dolomitic formation. The Middle Berriasian (Occitanica zone) under the same facies points out variations of thickness (60 to 100 m). Only the lower part of the Upper Berriasian (Paramimounum subzone) is present at the upper part with neritic facies (bioclastic limestones, sandstones, dolomites, breccia) and some changes of thickness (10 to 100 m). With the Upper Berriasian begins the thick detritic Meloussi formation.     

Zapaliinae,a new subfamily of Tithonian–Berriasian ataxioceratid ammonites

The recent recognition of the earliest Andean Tithonian ammonite fauna (Picunleufuense Zone) has allowed a balanced classification of the ataxioceratid ammonites, reflecting their phylogenetic relationships. The Picunleufuense Zone fauna, first recognized in the Neuquén Basin at the base of the Vaca Muerta Formation, is represented by numerous records of sparse ammonite faunas from Antarctica, the Austral Basin and the Tarapacá Basin (N. Chile and S. Peru). These Southeastern Pacific faunas include the Tithonian genera Indansites, Choicensisphinctes, Zapalia, Krantziceras (Tithonian–lower Berriasian) and Platydiscus. These five genera are grouped in a new subfamily established in this paper: Zapaliinae subfam. nov. (type genus: Zapalia). The origin of Indansites and Choicensisphinctes (possibly via Indanistes) from Lithacoceras is discussed. Zapalia must have been originated from Indanistes in the upper Proximus Zone. Platydiscus could have been originated from Krantziceras. The Tithonian Southeastern Pacific ataxioceratids (Zapaliinae subfam. nov.) form a group well separated from the Tithonian Indo-Madagascan Virgatosphinctinae.     

Late Ordovician–Early Silurian facies development and environmental changes in the Subpolar Urals

The continuous Upper Ashgill–Sheinwoodian carbonate succession in the most eastern Kozhym River area in the Subpolar Urals comprises the Yaptikshor (Rawtheyan), Kamennaya baba (Hirnantian), Ruchej and Manyuku (Llandovery–?Sheinwoodian) formations. The facies of the deep subtidal Yaptikshor Fm. mark an abrupt sea‐level rise following emergence of the Bad’ya reef (Rawtheyan). Carbonate breccias at the base of the Kamennaya baba Fm. correlate with the beginning of the Hirnantian glaciation and change upwards towards the Ordovician–Silurian boundary with the development of light‐grey massive boundstone/packstone shoal deposits. An abrupt change in facies to the Rhuddanian–Aeronian Ruchej Fm. continental slope environment marked the start of a long‐term sea‐level rise. The uppermost Aeronian–?Sheinwoodian is represented by submarine canyon carbonate conglobreccias of the Manyuku Fm. unconformably underlying the Balban’yu reef. The rapid facies changes at the base of the Hirnantian and at the Ordovician–Silurian boundary were of global eustatic origin. In contrast, the abrupt changes in the Rawtheyan and the formation of the Manyuku Fm. conglobreccias were of local or regional origin associated with tectonics. They were followed by the start of a regional transgression (Yaptikshor Fm.) and a global transgression marked by the initiation of the Balban’yu Reef in the Sheinwoodian.     

A facies model for an Early Aptian carbonate platform (Zamaia, Spain)

The Cretaceous (Early Aptian, uppermost Bedoulian, Dufrenoyia furcata Zone) Zamaia Formation is a carbonate unit, up to 224 m thick and 1.5 km wide, which formed on a regional coastal sea bordering the continental Iberian craton. A high-resolution, facies-based, stratigraphic framework is presented using facies mapping and vertical-log characterization. The depositional succession consists of a shallow estuarine facies of the Ereza Fm overlain by shallow-water rudist limestones (Zamaia Fm) building relief over positive tectonic blocks and separated by an intraplatform depression. The margins of these shallow-water rudist buildups record low-angle transitional slopes toward the adjacent surrounding basins. Syn-depositional faulting is responsible for differential subsidence and creation of highs and lows, and local emplacement of limestone olistoliths and slope breccias. Two main carbonate phases are separated by an intervening siliciclastic-carbonate estuarine episode. The platform carbonates are composed of repetitive swallowing-upward cycles, commonly ending with a paleokarstic surface. Depositional systems tracts within sequences are recognized on the basis of facies patterns and are interpreted in terms of variations of relative sea level. Both Zamaia carbonate platform phases were terminated by a relative sea-level fall and karstification, immediately followed by a relative sea-level rise. This study refines our understanding of the paleogeography and sea-level history in the Early Cretaceous Aptian of the Basque-Cantabrian Basin. The detailed information on biostratigraphy and lithostratigraphy provides a foundation for regional to global correlations.     

The use of neutral and non-neutral SSRs to analyse the genetic structure of a Tunisian collection of Medicago truncatula lines and to reveal associations with eco-environmental variables

In this study, we investigated the genetic diversity of a collection of 136 Medicago truncatula lines from 10 Tunisian natural populations collected in well-defined locations and in various ecological conditions of soil, salinity and water availability. The genetic diversity was evaluated using a set of 18 microsatellites (SSRs), representing the 8 chromosomes of M. truncatula. A neutrality test showed that 7 SSRs were non-neutral with evidence of balancing selection. The 11 neutral SSRs revealed a geographical pooling with the Tunisian Dorsale axis restricting migration of alleles. The 7 non-neutral alleles demonstrate a correlation with rainfall, altitude and salinity environmental variables suggesting that these SSRs are linked to genes involved in water use efficiency, resistance to salinity or adaptation to altitude, and that there is local adaptation of M. truncatula to these variables. This demonstrates that the choice of so-called neutral markers should be carefully evaluated in population genetic studies. This study illustrates the genetic diversity occurring in natural Tunisian populations of M. truncatula and describes the first collection of this species dedicated to natural variation involved in adaptation to the environment. F. Lazrek and V. Roussel contributed equally to this work.     

Silurian to Mississippian series of the eastern Catalan Pyrenees (Spain), updated by conodonts, foraminifers and algae

Stratigraphic sections and microfossils samples from the upper part of the “Graptolitic Shales-Orthoceras Limestones” and from the Camprodon Formation lying in the area of Camprodon, eastern Pyrenees, Spain, have been studied. Some beds at the top of the “Graptolitic Shales-Orthoceras Limestones” correspond to the Torres Member of the Rueda Formation, and conodont faunas, indicating a Lochkovian age, are described. The Camprodon Fm. is interpreted to be turbidites deposited in deep sea fans, although slope deposits prevail in the eastern sections. One reworked carbonate clast from the Camprodon Fm. provided a valuable early Ludlow conodont fauna, from the Kockelella crassa Zone, reported for the first time in the Iberian Peninsula. Late Devonian conodonts and late Visean (Asbian-Brigantian) foraminifers and algae were also obtained from reworked limestone clasts, and latest Visean or Serpukhovian foraminifers from the sandstone matrix in the Camprodon Fm. The studied microfossils suggest a late Mississippian age for the Camprodon Fm. instead of the previously assigned late Silurian-Lochkovian age. This age must be considered when discussing the distribution of the Culm Facies in the Pyrenees and the significance of the contact between the Camprodon Fm. and the underlying “Graptolitic Shales”.     

Heteromorph ammonites from the Tata Limestone Formation (Aptian–Lower Albian), Hungary

An important new ammonite fauna was collected by Fülöp in 1953 and 1960 from basal lenses between a Tithonian hardground and the overlying Tata Limestone Formation containing many heteromorph ammonites. This assemblage of ammonites has been determined as being of Early Albian ( tardefurcata Zone) age. Among the heteromorphs are species of Tonohamites and Ptychoceras , which are familiar components of Late Aptian or Early Albian ammonite faunas. The genera Ephamulina , Hamites and Protanisoceras are also recorded, the earliest species of which had previously come from sediments of late Early Albian ( mammillatum Zone) age. While Hamites and Protanisoceras are common in Europe, Ephamulina had previously only been known from Madagascar. The discovery of Ephamulina , Hamites and Protanisoceras supports the view that these phylogenetically significant ammonites had radiated close to the Aptian/Albian boundary. Four new species of Hamites ( H. czaszari , H. fazekasensis , H. fueloepi and H. kalvariensis ) and one new species of Tonohamites ( T. boldii ) are described.     

Tithonian ammonoid biostratigraphy in eastern Himalayan Tibet

The Tithonian and Lower Berriasian sediments in the eastern Himalayas of Tibet contain an extensive sequence of ammonoid fauna. New collections in situ through the Lanongla, Pure, Gucuo and Jiapeila sections have facilitated a major revision of the ammonoid assemblages. Probably due to depositional facies segregation, the Belemnopsis galoi-bearing beds can be regarded as the oldest Tithonian sediments in which the basal Tithonian ammonoid Kossmatia is not present. The Lower Tithonian includes the Virgatosphinctes-Aulacosphinctoides and Uhligites-Aulacosphinctes assemblages; the Upper Tithonian includes the Haplophylloceras pingue, Blanfordiceras wallichi and Haplophylloceras strigile-Corongoceras-Himalayites assemblages. The Spiticeras assemblage is suggested to be from the Lowermost Berriasian. The new ammonoid assemblages at well-defined levels in Himalayan Tibet provide some crucial links for correlation with other regions of the SW Pacific domain where these ammonoid genera have been widely distributed.     

Presence de Graptolites de l'Arenig moyen a la basede la Formation de Postolonnec (Massif Armoricain); Consequences stratigraphiques et paleogeographiques

The lowest part of the Postolonnec Formation (western Brittany) can be assigned to the middle Arenig Didymograptus nitidus Zone on graptolite evidence. Preliminary information on the Chitinozoa agrees with this age assignment, which defines more precisely the age of the underlying Grès armoricain Formation and also demands reconsideration of the problems associated with early Ordovician palaeogeography and sedimentology in the Massif armoricain.     

Ammonite habitats in the Early Tithonian of Western Tethys

Cccca, F. 1992 07 15: Ammonite habitats in the Early Tithonian of Western Tethys.
Habitats of Early Tithonian ammonites of Western Tethys are inferred from analysis of faunal spectra, functional morphology. facies characteristics and palaeogeography. Phylloceratina and Lytoceratina are included in a pelagic group characterized by taxa which could reach bathyal depths. This group is very abundant and dominant in some localities. A neritic nektobenthic group, relatively abundant in the Cmbria-Marche Apennines only. is composed of taxa characterired by species bearing simple sutures (some Simoceratidae and Pseudolissoceras ). probably living at shallower depth. The habitat of Haplploceras and some Oppeliidae. is interpreted to be close to the shelf margin/continental slope. Perisphinctids. Aspidoceratidae and some other Oppeliidae are provisionally included in an 'indeterminate' group including deep and shallow taxa. Quantitative studies of assemblage composition and comparison of faunal spectra successions from different pelagic regions allow the characterization of different pelagic environments. Bathymetric hypotheses by means of ammonites are discussed. Z Ammonoidea. habitats. facies, faunal spectra. Jurassic, Tithonian. Tethys .     

Le passage Lias–Dogger de la Dorsale de Tunisie septentrionale: Nouveaux apports biostratigraphiques. Âge Toarcien supérieur de la distension téthysienne

In the Jebel Bou Kornine of Hammam Lif (northern Tunisia), the transition between Lias and Dogger is located in a carbonate formation that is approximately 150 m thick (Kef El Orma Formation). Based on ammonite faunas, its age can be determined as Upper Toarcian (Aalensis Zone) up to Lower Bajocian (Propinquans Zone). Four successive conglomeratic units are recognized in this formation. The two earlier ones are situated in the Uppermost Toarcian (Aalensis Zone, Lugdunensis Subzone), while the later two are in the Opalinum Zone of the Lower Aalenian (Opalinum and Comptum Subzones). These deposits, which associate carbonate-bearing conglomerates with laminated/slumped calcarenites, result from a gravitational flow on a palaeoslope induced by extensive palaeofaults. They testify to a tectonic instability during the Upper Toarcian, causing the formation of two distinct palaeostructural units: the ‘Tunisian trough’ and the ‘Tunisian ridge’ carbonated platform, both belonging to the Maghrebian passive margin of the Tethys. We must insist on the part played, as soon as the Upper Toarcian, by the polyphased tilted-blocks tectonics on the deposition of the different conglomeratic units of the Bou Kornine Jebel.     

Nouveautés sur la séquence du Pléistocène final et l’Holocène initial dans le versant méditerranéen de la péninsule Ibérique à travers l’industrie lithique

This paper explores the cultural sequence for the period 12,500–9000 BP in the Mediterranean coast of the Iberian Peninsula. According to the traditional division by Fortea, this period starts with the end of the Upper Magdalenian, followed by two facies: the Microlaminar Epipalaeolithic and the Geometric Epipalaeolithic (Filador type), (related to Sauveterrian facies). The data collected in recent years suggest a change or amendment to this previous proposal, which now would be divided into four phases: Final Upper Magdalenian, Early Epimagdalenian, Recent Epimagdalenian and Microlaminar Sauveterroid. This paper discusses each of these periods examining the evolution of the stone tools and concluding that in the Mediterranean coast of the Iberian Peninsula the Magdalenian tradition continues with little changes until the advent of the Mesolithic.