On the ecological scenario of the first hominin dispersal out of Africa

The archaeopaleontological site of Dmanisi in Georgia, dated to ～1.8 Ma, provides evidence on the first hominin dispersal out of Africa, while the sites of Barranco León and Fuente Nueva-3 in Spain, dated to ～1.4 Ma, record the earliest hominin settlements in Europe. However, a number of issues related to the dispersal route, the climatic conditions and the ecological scenario of this dispersal event are subject to debate. In a recent paper in L’anthropologie, Agustí and Lordkipanidze (2019) proposed an alternative scenario for the arrival of hominins in the Caucasus, which they conceived as a forest refugium area during the Early Pleistocene, and discarded that their dispersal coincided with that of other members of the Ethiopian and Asian faunas, like the sabertooth Megantereon whitei or the giant hyena Pachycrocuta brevirostris. Our review of these issues suggests that: (i) the elongated sabers and reduced postcanine teeth of African M. whitei limited the ability of this predator to process the prey carcass, which resulted in scavengeable resources for the Dmanisi hominins; (ii) the mass estimate in excess of 100 kg obtained for the trochlear perimeter of the distal humerus of the hyena from Dmanisi shows that it can be confidently ascribed to the genus Pachycrocuta; (iii) the postcranial anatomy of the Dmanisi hominins was not advantageous for scavenging tree-stored prey; (iv) the laterally flattened upper canines of M. whitei could not withstand the loads that would result from climbing a prey carcass into a tree; (v) paleobotanical analyses suggest a temperate grassland ecosystem in Dmanisi, not dominant forest conditions, with enhanced aridity in the level of hominin occupation; (vi) similarly, the low frequency of arboreal pollen in the Levantine Corridor at ～1.8 Ma points to more arid conditions than today in this area; (vii) many archaeopaleontological sites of the Rift Valley and its extension towards the Red Sea, the Levant and the Caucasus show evidence of tectonic, volcanic and/or hydrothermal events; and (viii) the delay of 400 ka in the arrival of hominins in Western Europe did not result from a lower availability of scavengeable resources.     

Paleoenvironmental reconstruction of the Early Pleistocene site of Quibas (SE Spain) using a rodent assemblage

In this paper we analyze the rodent assemblage from the Early Pleistocene site of Gruta1 (Quibas karstic complex, Murcia, SE Spain), providing taphonomic comments and paleoecological and paleoclimatic data. The studied assemblage includes 209 identified rodent teeth corresponding to at least four taxa. The karstic source of the site and the low presence of digested remains suggest that the accumulation is scatological in origin, with the influence of predators such as owls; the accumulation also shows certain evidence of slight hydrodynamic sorting. For the area around Quibas/Gruta1 the paleoecological study indicates a predominance of rocky areas (31.7%), forested environments (31.7%) and open dry meadows or shrublands (29.7%), which would indicate a mosaic forest environment, and to a lesser extent the presence of open humid land (5.4%) and areas along streams or ponds (1.5%). The distribution of the bioclimatic spectra yields the highest percentage for a Mediterranean climate (37.5%), while the climatic parameters calculated (MAT = 15 °C, MTW = 25.7 °C, MTC = 4.3 °C, MAP = 390 mm) suggest colder and slightly more humid conditions than today at the time of the deposition of the remains. The data also point to a relative decrease in temperature within the phase in which the site was produced, which possibly occurred during a relative cold period between MIS 36 and MIS 40 according to the age.     

Hominin occupations at the Dmanisi site, Georgia, Southern Caucasus: raw materials and technical behaviours of Europe's first hominins

Dmanisi is the oldest site outside of Africa that records unquestioned hominin occupations as well as the dispersal of hominins in Europe and Asia. The site has yielded large numbers of artefacts from several periods of hominin occupation. This analysis of Dmanisi stone tool technology includes a review of all the pieces recovered during the last 15 years of excavations. This lithic assemblage gives insights into the hominin behaviour at 1.7-1.8 Ma in Eurasia. Dmanisi hominins exploited local rocks derived from either nearby riverbeds or outcrops, and petrographic study provides data on patterns of stone procurement. Recent geological surveys and technological studies of the artefacts illustrate the roles of hominins in composing the assemblage. Dmanisi hominins selected two types of blanks, including cobbles and angular blocks, of basalt, andesite, and tuffs. Many complete cobbles, pebbles, and rolled blocks in basalt were unmodified, and geological analyses and surveys indicate that hominins brought manuports back to the site, suggesting a complex procurement strategy. Cores, flakes and debris show that all stages of flaking activity took place at the site. Numerous unifacial cores suggest that knapping was not very elaborate. Centripetal knapping is observed on some flake-cores. Knapping was influenced by the blank shape and natural angles. Most flaked objects were either cores or chopper-cores. Flakes predominate while flake tools are rare. The Dmanisi lithic assemblage is comparable to Oldowan sites in Africa in terms of reduction sequence, organisation of the removals, platform types, and the lack of retouched flakes. Dmanisi artefacts and may have been produced by the original hominins in Europe and Asia.     

The Oldowan site of Muhkai II,layer 80 (northeastern Caucasus): Spatial structure and cultural and chronological attribution of the lithic assemblage

In 2004–2006 a group of Lower Palaeolithic sites was discovered in the northeastern Caucasus: Ainikab I–VI, Gegalashur I–III and Muhkai I–II. Since then, they have been the subject of comprehensive scientific investigations. One site which is undergoing active research is the multilayered site of Muhkai II. Multiple cultural layers at the site have been found in a 73 m thick sequence of Early Pleistocene sediments. The geological age of the layers encompasses the interval 2.1–0.8 million years ago, or the majority of the Early Pleistocene. Layer 80 of Muhkai II is located in the central part of the section at a depth of −33.6 to −34.0 m. Its significance lies in the fact that the site contains two cultural layers–the upper and the lower (main)–that are preserved in situ or have been subject to minimal fluvial processes. The finds consist of lithic artefacts and osseous faunal remains. According to field observations and microstratigraphic analyses, the cultural layers can be characterised as the ancient surfaces of an occupation area. Two stages of occupation can be defined in the lower cultural layer. Layer 80 of Muhkai II was a periodically visited location at the banks of a waterbody, where the acquisition and butchery of large mammals took place. The faunal collection contains 301 bones, deriving from four carnivorous and six herbivorous species of mammals. The total lithic collection contains 1094 finds. According to typological and technological criteria the lithic assemblage relates to the Oldowan and has analogies with Oldowan sites in Africa (particularly with sites of the Olduvai Gorge as well as others). At the same time, some specifics of the lithic assemblage reflect the uniqueness of the site and its functional type. Layer 80 of the Muhkai II site is estimated to date to within the chronological interval ∼2.1–1.7 mya.     

An interesting rock from Krapina

Symbolic items are seldom associated with Neandertals and, when they are, many paleoanthropologists consider them to be Neandertal imitations from modern Homo sapiens. At the Croatian site of Krapina dated to MIS 5e or about 130,000 years ago, among many items, a split limestone rock was excavated by Dragutin Gorjanović-Kramberger between 1899-1905. This brownish rock reveals many dark dendritic forms in cross- and longitudinal section. Of more than 1000 lithic items at Krapina, none resemble this specimen and we propose it was collected and not further processed by the Neandertals because of its aesthetic attributes. Along with other examples from sites in western and central Europe and the recent discovery of eight modified white-tailed eagle talons from Krapina, this unique item suggests that Neandertals were capable, on their own, of incorporating symbolic objects into their culture.     

The Kromdraai early hominin-bearing site. A review of recent findings

The Kromdraai archaeological site is located in a fossiliferous paleokarst situated in the UNESCO World Heritage Site referred to as the “Cradle of Humankind” in the Gauteng Province of South Africa. Kromdraai is noteworthy because it features among the three southern African early hominin-bearing sites considered to represent distinct temporal periods within the same stratigraphic succession. Kromdraai also yielded a partial skull and dentition (TM 1517) in 1938 that was designated as the holotype of a new genus and species, Paranthropus robustus. Although the hominin fossil assemblage collected from Kromdraai between 1938 and 2014 is rather paltry, morphometric and cladistic analyses of this material suggested that it represented a somewhat less-derived form of P. robustus than the considerably larger assemblage from the nearby site of Swartkrans. However, the geochronological and biotic relationships among the P. robustus-bearing sites in South Africa are not resolved. Since 2014, the previously unknown, albeit densely fossiliferous Unit P produced 51 individually catalogued hominin fossils (36 craniodental and 15 postcranial) that currently represent 13% of the faunal assemblage from this unit with a minimum number of 10 juvenile and 9 adult individuals. P. robustus and early Homo coexisted at the time of the accumulation of Unit P at Kromdraai, with a relative abundance of 89% and 11%, respectively. P. robustus and early Homo are associated with a highly diverse fauna that includes several water-dependent species, and a large variety of bovid and carnivore taxa. Biochronological data from Unit P and an interval of reversed polarity measured in younger deposits above it are interpreted in the context of the regional chronological framework to infer that it represents a deposit that was likely accumulated prior to 2 Ma.     

Les industries archaïques au Proche-Orient

The Middle East (from the Mediterranean coast to Iran and from the Red Sea to the Black Sea) is located at the crossroads of African, Asian and European continents. It is a compulsory route well-trodden during the dispersal of the first humans out of Africa. Recent discoveries, mainly in Syria, Jordan and Israel, show that the first settlements in this area date back to over 2 million years (Ma). The location of the deposits containing archaic industries of the Oldowan type in the broad sense, often called “Core and Flake Industries”, indicates that several roads have been repeatedly used to connect Egypt and the Arabian Peninsula with the Euphrates and the Tiger basins. From south to north, if the coastal road and the tectonic depression of the Dead Sea, the Jordan Valley and the Beqaa were privileged places of very old settlements, the desert roads through Jordan and Syria were also readily used even before 2 Ma. We can observe at least three successive stages of archaic industries. During the oldest one, from about 2.5 to 1.8 Ma, choppers, chopping-tools, cores and flakes are prevailing without traces of intentional retouching (Sites of Aïn Al Fil in Syria, of the Zarqa Valley in Jordan in particular). This period might be named Pre-Oldowan or Lower Oldowan. From approximately 1.8 to 1.3 Ma, a similar culture developed with the additional presence of regular polyhedrons and sometimes a rough bifacial shaping as well as the beginning of rare intentional retouching other than that of use-wear; we will call it Upper Oldowan (Lower G-Hummal in Syria, Bizat Ruhama and Lower Ubeidiya in Israel). Starting around 1.3 Ma, coarse bifaces begin to appear, always knapped with a stone hammer. This technology points to the transition towards Acheulean industries. The first bifaces appeared within a very Oldowan technological background (Hummal, Ubeidiya). However, in the Levant, sites often considered as “pre-Acheulean” because they lacked bifaces, though they belonged to more recent periods, younger than 1.3 Ma, have been termed “Tayacian” in order to underline their difference from Acheulean properly speaking. They seem contemporary with the first Acheulean phases and could correspond to a final Oldowan. The question remains, however, whether these are non-Acheulean cultures or “Acheuleans without bifaces”.     

Early human occupations at the westernmost tip of Eurasia: The lithic industries from Menez–Dregan I (Plouhinec,Finistère,France)

The stratigraphical complex preserved at the site of Menez–Dregan I (Brittany, France) displays an alternating sequence of 17 occupation levels and of 4 marine deposits, between ca. 450 and 150,000 years (stratigraphical units 9a and 3b). The lithic industry retrieved at the site is extremely abundant, with more than 153,000 artefacts over 3 cm in length, as well as millions of knapping debris, and corresponds to a regional facies of the Acheulean, with heavy-duty tools essentially represented by choppers, as well as a few handaxes and cleavers. This paper presents a first synthesis of the data produced for almost 30 years on the site of Menez–Dregan I. The typological study of the entire lithic collection is now complete and allows us to assess the different human occupations that occurred at this site during the Lower Palaeolithic.     

The oldest human fossil in Europe,from Orce (Spain)

The Orce region has one of the best late Pliocene and early Pleistocene continental paleobiological records of Europe. It is situated in the northeastern sector of the intramontane Guadix-Baza Basin (Granada, Andalusia, southern Spain). Here we describe a new fossil hominin tooth from the site of Barranco León, dated between 1.02 and 1.73 Ma (millions of years ago) by Electron Spin Resonance (ESR), which, in combination with paleomagnetic and biochronologic data, is estimated to be close to 1.4 Ma. While the range of dates obtained from these various methods overlaps with those published for the Sima del Elefante hominin locality (1.2 Ma), the overwhelming majority of evidence points to an older age. Thus, at the moment, the Barranco León hominin is the oldest from Western Europe.     

A chronological framework for a long and persistent archaeological record: Melka Kunture, Ethiopia

New ⁴⁰Ar/³⁹Ar geochronological data for several volcanic ash horizons from Melka Kunture, Ethiopia, allow for significantly more precise age constraints to be placed upon the lithostratigraphy, archaeology and paleontology from this long record. Ashes from the Melka Kunture Formation at Gombore yielded the most reliable age constraints, from 1.393 ± 0.162 Ma² (millions of years ago) near the base of the section to 0.709 ± 0.013 Ma near the top. Dating the Garba section proved more problematic, but the base of the section, which contains numerous Oldowan obsidian artifacts, may be >1.719 ± 0.199 Ma, while the top is securely dated to 0.869 ± 0.020 Ma. The large ignimbrite from the Kella Formation at Kella and Melka Garba is dated to 1.262 ± 0.034 Ma and pre-dates Acheulean artifacts in the area. The Gombore II site, which has yielded two Homo skull fragments, ‘twisted bifaces,’ and a preserved butchery site, is now constrained between 0.875 ± 0.010 Ma and 0.709 ± 0.013 Ma. Additional ashes from these and other sites further constrain the timing of deposition throughout the section.Integration with previously published magnetostratigraphy has allowed for the first time a relatively complete, reliable timeline for the deposition of sediments, environmental changes, archaeology, and paleontology at Melka Kunture.     

Cold-climate rodent indicators for the Late Pleistocene of Central Iberia: New data from the Buena Pinta Cave (Pinilla del Valle,Madrid Region,Spain)

In the Iberian Peninsula, the Late Pleistocene record of small mammal indicators of cold climates is largely restricted to two sets of sites at the eastern and western ends of the Pyrenees. Some assemblages from other sites at the Peninsular centre have, however, recently yielded such taxa. This work describes the remains of three such rodent species from the Buena Pinta Cave, a site in the Sierra de Guadarrama mountains in the Spanish Central System. Excavation campaigns in the cave have taken place every summer since 2003. Thermoluminescence analyses of sediments from levels 2–5 of the site suggest an age corresponding to the middle of the Late Pleistocene, within Marine Isotope Stage 4 or the beginning of Marine Isotope Stage 3. Sieve-washing and picking out of the small fossils contained in the sediments of levels 2–5 yielded several thousand small mammal teeth and other remains, dominated by Microtus arvalis. Smaller numbers of remains belonging to other rodents typical of cold climates were also identified, such as Microtus oeconomus, Microtus gregalis and Chionomys nivalis. Thus, the small mammal record of the Buena Pinta Cave shows that rodent indicators of cold climates reached the centre of the Iberian Peninsula during the mid-Late Pleistocene, i.e., well before the Last Glacial Maximum. These findings represent one of the southernmost Pleistocene records for M. oeconomus in Europe, and the most southerly for M. gregalis.     

Spatial behaviours of Early Oldowan toolmakers in the Shungura Formation (Lower Omo Valley,Ethiopia): Proposal for an integrated approach

The spatial relationships of the Early Oldowan toolmakers with their environment have been so far addressed through raw material procurement analyses and the characterization of hominid habitat. This paper proposes to integrate these two approaches into a broader spatial analysis encompassing archaeological and environmental data (palaeontological, geological and isotopic data) from Member F and lower Member G of the Shungura Formation (Lower Omo Valley, Ethiopia). Heterogeneity in data resolution induces a multiscale approach with three levels of analysis. The level of occurrence complex allows focusing on the characterization of archaeological occurrences and on their environmental settings. The level of “study area” allows working on hominid habitats and on their raw material procurement behaviours. Finally, at the Shungura Formation scale, we can address temporal issues related to the evolution of spatial behaviours between Member F and the lower part of Member G, ca. 2.3 to 2 million years (Ma).     

Epipaleolithic human appendicular remains from Ein Gev I,Israel

The Upper Paleolithic (Early Epipaleolithic/Kebaran; ∼ 19,000 cal BP) human skeleton, from Layer 3 of Ein Gev I on the western flanks of the Golan Heights adjacent to the Sea of Galilee, retains sufficient limb remains to permit assessment of its body size and proportions, as well as diaphyseal reflections of skeletal hypertrophy. The individual was of modest stature but average mass for a later Upper Paleolithic individual, providing it with the body mass-to-stature body proportions characteristic of later Upper Paleolithic and more recent circum-Mediterranean humans. The humeri exhibit unexceptional diaphyseal asymmetry and robustness for an Upper Paleolithic human, and the femur exhibits similar relative diaphyseal hypertrophy. The humeral midshafts are relatively round, but the femoral and tibial midshafts are pronounced anteroposteriorly. As such, Ein Gev 1 provides additional paleobiological data on the appendicular remains of these Southwest Asian humans prior to the increasing sedentism of the terminal Pleistocene.     

Late Pleistocene proboscidean ivory artifacts from the Hiscock site,NY

Here is described the corpus of proboscidean ivory artifacts – the fruit of 30 years excavating – within the Palaeo-American component at the Hiscock site, western New York State. Several types and varieties are newly recognized and unique for the North American archaeological record. They allow us to forge linkages between the Old and New Worlds.     

Ostracods from the marginal coastal Lower Cretaceous (Aptian) of the Central Tunisian Atlas (North Africa): Paleoenvironment,biostratigraphy and paleobiogeography

This work provides the first detailed taxonomic study of ostracod species from the Lower Cretaceous (Aptian) marginal coastal deposits of the Central Tunisian Atlas, the Kebar Formation, as well as biostratigraphic, paleoecological and paleobiogeographic implications. The ostracod fauna provides new insights into the depositional environment and biostratigraphic framework of the Kebar Formation, and is represented mainly by freshwater and brackish-water species among them, Perissocytheridea tunisiatlasica nov. sp., is newly described. The environmental setting of this formation comprises marginal-littoral conditions in its lower part, thus not exclusively non-marine/continental as assumed previously. The ostracod associations from the studied locality, Jebel Ksaïra, indicate a minimum age of Early Aptian as recently attributed to the lower member of the Kebar Formation based on charophytes (presence of Late Barremian to Early Aptian Globator maillardii var. biutricularis Vicente and Martín-Closas, 2012), whereas an Early Albian age had been previously assigned to the Jebel Kebar site. The relative sea-level fall documented in the lower member of the Kebar formation at Jebel Ksaïra might correspond to the 3rd order cycle major eustatic sea-level fall event starting at the base of the Aptian. Paleobiogeographically, the non-marine ostracod fauna newly discovered in the Kebar Formation shows some affinities to contemporaneous faunas of southern and western Europe, e.g. that of the uppermost Weald Clay Group of southern England (Cypridea fasciata subzone of Horne, 1995), West Africa, and possibly to eastern South America (Brazil). Hence, the studied ostracods further support the hypothesis of supraregional faunal exchange by passive ostracod dispersal during the Early Cretaceous – between Europe and Northern Africa on the one hand, and eastern South America/western Africa and North Africa on the other hand. This leads to the hypothesis that the Peri-Tethyan islands could have worked as effective bridges for non-marine ostracods to become widely dispersed passively by “island-hopping” of larger animals and thus, ultimately, facilitated intercontinental faunal exchanges between South America and Europe – potentially even Asia – via North Africa during the Early Cretaceous.     

Palaeomagnetic analysis of the Sterkfontein palaeocave deposits: implications for the age of the hominin fossils and stone tool industries

Palaeomagnetic analysis was conducted on speleothems from Members 1-5 at Sterkfontein Cave, South Africa. Palaeomagnetic analysis of siltstone and speleothem from the bulk of Member 4 indicate a reversed magnetic polarity that dates the deposits and its Australopithecus africanus fossils to between 2.58 and ∼2.16 Ma. Further confirmation of this age comes in the form of two short normal polarity events correlated to the Rèunion (∼2.16 Ma) and Huckleberry Ridge (∼2.05 Ma) events in speleothem capping the bulk of Member 4 and coeval with deposition of the final phase of Member 4, including A. africanus fossil Sts 5. At ∼2.16-2.05 Ma, Sts 5 is the youngest representative of A. africanus yet discovered. Palaeomagnetic analysis of the Silberberg Grotto deposits identifies a single short geomagnetic field event in flowstone overlying the StW 573 Australopithecus fossil, which is suggested to represent the Rèunion event at ∼2.16 Ma. This further supports the uranium lead age estimates of 2.3-2.2 Ma for the StW 573 fossil. Based on a reversed polarity for the deposits below the skeleton it cannot be older than 2.58 Ma. If StW 573 is considered to be a second species of Australopithecus then this indicates that two species of Australopithecus are present at Sterkfontein between 2.6 and 2.0 Ma. All of the Member 5 deposits date to less than 1.8 Ma based on a comparison of palaeomagnetic, faunal, and electron spin resonance age estimates. The StW 53 fossil bearing infill (M5A) is intermediate in age between Member 4 and the rest of Member 5 (B-C) at around 1.78-1.49 Ma. The rest of Member 5 (B-C) containing Oldowan and Acheulian stone tools and Homo and Paranthropus fossils was deposited gradually between 1.40 and 1.07 Ma, much younger than previously suggested.     

Morphometry,biogeography and ecology of Calcidiscus and Umbilicosphaera in the South Atlantic

Here we present morphometrical evaluation, biogeographical distribution patterns and ecological information for five coccolithophore taxa (Calcidiscus leptoporus, C. leptoporus small, C. quadriperforatus, Umbilicosphaera foliosa and U. sibogae). This information is based on data obtained from surface sediments from the South Atlantic. The three Calcidiscus taxa can easily been distinguished by a combination of size and qualitative characters of their distal shields. Mostly encountered in the temperate to sub-polar regions C. leptoporus is the most abundant taxon and exhibits a negative correlation to temperature and salinity. Both, C. leptoporus small and C. quadriperforatus reach their maximum abundances also at higher latitudes and in the SW-African upwelling area. Their distributions therefore suggest preference for nutrient-enriched waters, which is also indicated by CCA. The two circular Umbilicosphaera species exhibit significant differences in coccolith morphology and show little overlap in size. Highest abundances are encountered in sub-tropical latitudes and are mainly derived from U. sibogae. In contrast, U. foliosa is present in very low abundances. Both species exhibit a preference for warm and oligotrophic conditions. However, U. foliosa increases in relative proportion to U. sibogae at the southernmost locations and in the Benguela upwelling. This could be interpreted as an affinity for slightly cooler and nutrient-enriched environments.     

New anthropological data from Cussac Cave (Gravettian,Dordogne, France): In situ and virtual analyses of Locus 3

Cussac Cave presents a unique combination of parietal art and several hundred parts of scattered human remains, dated to the Middle Gravettian (29–28,000 cal BP). The cave is protected as a National Heritage site. As a result, only noninvasive bioanthropological analyses are allowed, consisting of in situ observations and the study of 3D models obtained by photogrammetry. Here we present the first results of these analyses of the human remains from Locus 3. Only 65 of the 106 human skeletal fragments and bones could be firmly identified. Virtual analyses were carried out on 3D models of 16 skeletal elements so that osteometric data could be provided. Despite the limitations inherent in studying commingled remains and those specific to Cussac Cave, the search for virtual pair-matching, articular congruence, and osteometric sorting allowed the allocation of twelve bones to three individuals, one late adolescent and two adults.     

Découverte des sites du Paléolithique inférieur au Nord de l’Arménie

For a long time, the scientific community assumed that the Acheulean culture was expressed on the territory of the Armenian Plateau as well as in the neighboring regions of the Caucasus only by its late phase; therefore, it appeared in the second half of the Middle Pleistocene. In recent years, the Armenian-Russian mission has discovered and studied much older Acheulean industries sites, located in northern Armenia (the Lori intermountain Depression). These industries, represented by archaic type tools (large hand axes, picks, choppers, chisel-like tools, scrapers, points, etc.), are discovered in three deposits of origin of proluvial genesis. In the Karakhach site, this type of industry is deposited in the lower levels of volcanic tuff and below; the U/Pb study of this level of tuff proposes a series of dates, assigned to the time interval between 1.944 + 0.046 and 1.75 + 0.02 Ma. The paleomagnetic study demonstrated the inverse polarity on the tuff and the normal polarity of the underlying deposits; in correlation with other dating, this fact allows to attribute the Acheulean layers of the site of Karakhatch to the Lower Pleistocene, in particular to the Oldoway episode and to the Upper Matuyama time period. The estimated age and the techno-morphological characteristics of the tools indicate the Lower Acheulean period. The dating of the Muradovo site does not seem possible, however its very old industries and the archaeological layers, where they were discovered, find equivalents in the layered layer, surmounted by tuffs, of the Karakhatch site. The Kutran I site presented a paleosol sequence with similar Acheulean tools (hand axes, picks, choppers, etc.). Its oldest layer is older than 1.5 Ma, the upper layer is attributed to the early Middle Pleistocene; this fact means that it is possible to speak of the Lower Acheulean and of its transition to the Middle Acheulean period. The specific character and the age of the Lower Acheulean of Armenia admit that it could have formed independently of the Lower Acheulean of Africa, whose estimated age does not rise before 1.76 Ma. It should also be noted that on the neighboring territory of Georgia about the same time when appeared the Acheulean culture in Armenia, the Oldowan Dmanisi site already existed.