Direct and Indirect Track Features: What Sediment Did a Dinosaur Touch?

Studies of dinosaur tracks have benefited from a distinction between true tracks and those made in subsurface layers—undertracks. However, the straightforward definition of true tracks becomes problematic when dealing with deep tracks, which often perforate or incise surface layers rather than simply distort them. Deep tracks from the Late Triassic of Greenland were made by theropods moving their feet through a volume of sediment along a complex three-dimensional trajectory. I suggest that designating different portions of the track as direct or indirect features is fruitful for reconstructing foot motion. Identifying which sedimentary grains, rather than which layers, were touched by the foot avoids dismissing deep tracks as undertracks and overlooking a valuable source of kinematic data.     

The First Iguanodontid Dinosaur Tracks from the Swiss Alps (Schrattenkalk Formation,Aptian)

Until now dinosaur tracks from Switzerland were only known from Triassic and Late Jurassic strata. We report here for the first time the occurrence of ornithopod tracks from the Schrattenkalk Formation (Late Aptian) from the Swiss Central Alps. The locality is situated in an abandoned quarry on the shore of Lake Lucerne close to the village of Beckenried. The steeply inclined surface has more than 50 tracks (in three trackways) of ornithopod dinosaurs that are attributed to iguanodontids. Three trackways can be followed for distances of 25 to 35 m. The lengths the footprints (mean: 30 cm) point to animals ranging in size of from 4 to 6 m, with estimated hip heights between 1.8 and 2 m (hip height 6 FL) and 1.4 to 1.7 m (hip height 5 FL). One of the trackways shows two succesive manus impressions, indicating facultative quadrupedal gait. The track bearing layer consists of shallow water micrites with traces of emersion, and it is overlain by bioclastic grainstones. Previously the Upper Schrattenkalk Member in the Helvetic realm was thought to have formed on a large shelf far away from any continents. The present discovery will shed new light on the paleogeographic position of the Helvetic nappes.     

Two new dinosaur tracksites from the Lower Cretaceous Jiaguan Formation of Sichuan Basin,China: specific preservation and ichnotaxonomy

Two new dinosaur tracksites are reported from the Lower Cretaceous Jiaguan Formation in the Sichuan Basin, Qijiang District of Chongqing. These are the Gaoqing-Yongsheng and the Huibu tracksites, which represent the 13th and 14th reports from this formation. The Gaoqing-Yongsheng tracksite reveals the trackway of a large biped (ornithopod) in association with isolated sauropod tracks and large indeterminate undertracks with radial cracks. These features are preserved as natural casts with pebble infillings in a coarse, cross bedded and very thick bedded sandstone sequence. The Huibu tracksite reveals isolated theropod tracks and ornithopod tracks, the latter having a quadripartite, Caririchnium-like morphology, preserved in a thin bedded sandstone sequence with intercalated mudstone.     

Dinosaur Tracks from the Cedar Mountain Formation (Lower Cretaceous), Arches National Park,Utah

The seventh and largest known dinosaur tracksite from the Cedar Mountain Formation is reported from two important stratigraphic levels in the Ruby Ranch Member within the boundaries of Arches National Park. Previous reports of sites with a few isolated tracks are of limited utility in indicating the fauna represented by track makers. The Arches site reveals evidence of several theropod morphotypes, including a possible match for the coelurosaur Nedcolbertia and an apparently didactyl Utahraptor-like dromeosaurid. Sauropod tracks indicate a wide-gauge morphotype (cf. Brontopodus). Ornithischian tracks suggest the presence of an iguandontid-like ornithopod and a large ankylosaur. Dinosaur track diversity is high in comparison with other early Cretaceous vertebrate ichnofaunas, and it correlates well with faunal lists derived from skeletal remains, thus providing a convincing census of the known fauna.     

Telling Apart Ornithopod and Theropod Trackways: A Closer Look at a Large,Late Jurassic Tridactyl Dinosaur Trackway at Serwah,Republic of Yemen

A large bipedal tridactyl dinosaur trackway from the Late Jurassic of Serwah, near Madar, Arhab district, Republic of Yemen, has been attributed to an ornithopod trackmaker. As the distinction between theropod and ornithopod dinosaurs can pose a challenge, we present additional data to support and reconfirm the previous attribution.     

Behaviour of Dinosaurian Track-Makers in the Winton Formation (Cretaceous,Albian–Cenomanian) at Lark Quarry,Western Queensland,Australia: Running or Swimming?

A remarkable assemblage of dinosaur trackways in the Winton Formation (Albian–Cenomanian) at Lark Quarry, in western Queensland, Australia, has long been regarded as evidence of a stampede involving small theropods, whose tracks were classified in the ichnogenus Skartopus, and small ornithopods, whose tracks represented a second ichnogenus, Wintonopus. However, one recently-published study has claimed that existing interpretation of Lark Quarry is incorrect: it maintains that all the track-makers were ornithopods, that the ichnogenus Skartopus is a variant form of Wintonopus, and that most of the Lark Quarry track-makers were not running but more probably swimming downstream in a current of water. Those iconoclastic claims are investigated here and shown to be untenable. They derive from overgeneralized interpretation of the Lark Quarry track assemblage and invoke ad hoc auxiliary hypotheses which are either untestable or demonstrably incorrect. Closer inspection of the evidence underpinning those claims corroborates the existing interpretation of Lark Quarry as the site of a dinosaurian stampede and confirms the validity of the original distinction between theropod tracks (Skartopus) and ornithopod tracks (Wintonopus).     

Reconstruction of Middle Jurassic Dinosaur-Dominated Communities from the Vertebrate Ichnofauna of the Cleveland Basin of Yorkshire,UK

Globally, skeletal remains of dinosaurs are particularly rare throughout much of the Middle Jurassic. Thus, other sources of evidence, and most importantly ichnofaunas, are important indicators of the contemporary terrestrial vertebrate communities. The outcrops of the Ravenscar Group (Aalenian—Bajocian) within the Cleveland Basin of Yorkshire, UK, which have recently been recognised as a megatracksite of global significance, provide one such major source of ichnofaunal information of this age. A comprehensive database on the variety and occurrence of dinosaur and other vertebrate traces within the Ravenscar Group has been built from a long-term and detailed study of the sequence. Thirty different and distinct morphotypes of vertebrate traces have been recognised and are being analysed and further differentiated morphometrically. Some of the morphotypes represent behavioural, preservational and perhaps ontogenetic variants of other morphotypes, but nevertheless the range of quadrupedal and bipedal prints allows an overall fauna of sauropod, stegosaurian, ornithopod and theropod dinosaurs along with crocodiles, pond turtles and fish to be reconstructed. The distribution and abundance of prints and print types within the succession shows evidence of environmental control on the behaviour and distribution of the vertebrates. Case studies highlight both the advantages and disadvantages of this type of data in reconstructing palaeocommunities.     

Conservation of Leafhoppers in Floodplain Grasslands – Trade-off between Diversity and Naturalness in a Northern German National Park

The impacts of mowing and of flooding on the leafhopper communities of a river flood plain were investigated. Samples were taken by a motor-driven suction apparatus. In 2001 leafhoppers were collected in a variety of sites differing in land use (fallows, mown sites) and in flood intensity (high, medium, low, none). In 2002 samples were only taken in fallows subject to different flooding regimes.In fallows, more species (43) were collected than in mown sites (33). Flooding had an effect only in fallows. Here, the most species-rich (29) communities occurred in sites not subject to flooding, whereas fewest species (21) were found in sites subject to regularly occurring long lasting winter floods. Mown sites were dominated by pioneer species. In fallows, the communities differed in respect to flood intensity. In fallows that were subject to summer and winter floods pioneer species prevailed. In contrast, in fallows that were flooded a long time during winter but not in summer, communities of very specialised species were found which were not very species-rich.For the conservation of the typical leafhopper communities of floodplain grassland, management by mowing should be at least reduced if not totally stopped and natural flooding dynamics should be restored.     

广东省南雄盆地白垩系—第三系交界恐龙绝灭问题

广东省南雄盆地中的红层可划分为三个群五个组,大致代表了晚白垩世—始新世的沉积.根据绝对年龄、古地磁测定结果和脊椎动物化石组合性质的综合分析,位于地磁极性带 29R 上部的坪岭组和上湖组之间的分界线被确定为 K/T 界线.对晚白垩世恐龙蛋的研究表明,不同\"种\"的恐龙蛋是在地磁极性带 29R 的中、下部,也就是说在白垩系—第三系交界之前20～30万年期间绝迹的.而且在这一时期内,所有已发现的蛋壳中,绝大多数蛋壳的厚度和显微结构都显示出明显的病理特征,例如根据随机取样统计,Macroolithus yaotunensis 蛋壳异常结构的出现率,最高可达75%.产生病态恐龙蛋壳的生理机制可以根据发生在现生鸟类的相同病理特征来解释.进一步分析恐龙蛋壳的微量元素和稳定同位素组成,结果显示, Pb, Cu, Mn 等9种元素丰度变化在这一时期达到最大峰值, δ~(18)O 也出现正异常.在这一基础上提出,微量元素的污染和气候突然的变化妨碍了正常蛋壳结构的形成,导致了恐龙的绝灭.这一绝灭过程大约经历了20～30万年.     

A theropod tooth from the Late Triassic of southern Africa

An isolated, large recurved and finely serrated tooth found associated with the prosauropodEuskelosaurus from the Late Triassic part of the Elliot Formation is described here. It is compared to the Triassic thecodonts and carnivorous dinosaurs and its possible affinity is discussed. The tooth possibly belongs to a basal theropod and shows some features similar to the allosauroids. This tooth is of significance, as dinosaur remains except for some footprints and trackways, are poorly known in the Late Triassic horizons of southern Africa.