Miofloral succession and interpretation of the base of the permian system in the Eastern Shelf of North Central Texas,U.S.A.

The study includes palynological investigation of the Pennsylvanian (Upper Carboniferous) and lowermost Permian (Wolfcampian) strata of Texas, with special emphasis on the Pennsylvanian and Permian systemic boundary in the Eastern Shelf of north central Texas. The following palynozones have been recognized: (1) Grumososporites-Simozonotriletes Zone (Atokan); (2) Torispora-Thymospora Zone (Demoinesian); (3) Cristatisporites—Cadiospora arcuata Zone (Missourian); (4) Crucisaccites americanus Zone (Early Virgilian); (5) Guptaesporites honakerensis Zone (Late Virgilian); and (6) Nuskoisporites crenulatus-Hamiapollenites saccatus Zone (Early Wolfcampian). The base of the Permian Wolfcampian Stage has been placed at the base of Nuskoisporites crenulatus-Hamiapollenites saccatus Zone, which approximately coincides with the base of the Coleman Junction Limestone Member of the Putnam Formation of the Wichita Group. This conclusion may appear to be contradictory to fusulinid evidence because it places the base of the Permian System 182.88 m above the so-called fusulinid boundary. It also disagrees with the paleobotanical boundary. The discussion concludes that fusulinid and paleobotanical evidence is inconclusive.     

Revised Lithostratigraphy of the Sonsela Member (Chinle Formation,Upper Triassic) in the Southern Part of Petrified Forest National Park,Arizona

Background

Recent revisions to the Sonsela Member of the Chinle Formation in Petrified Forest National Park have presented a three-part lithostratigraphic model based on unconventional correlations of sandstone beds. As a vertebrate faunal transition is recorded within this stratigraphic interval, these correlations, and the purported existence of a depositional hiatus (the Tr-4 unconformity) at about the same level, must be carefully re-examined.

Methodology/Principal Findings

Our investigations demonstrate the neglected necessity of walking out contacts and mapping when constructing lithostratigraphic models, and providing UTM coordinates and labeled photographs for all measured sections. We correct correlation errors within the Sonsela Member, demonstrate that there are multiple Flattops One sandstones, all of which are higher than the traditional Sonsela sandstone bed, that the Sonsela sandstone bed and Rainbow Forest Bed are equivalent, that the Rainbow Forest Bed is higher than the sandstones at the base of Blue Mesa and Agate Mesa, that strata formerly assigned to the Jim Camp Wash beds occur at two stratigraphic levels, and that there are multiple persistent silcrete horizons within the Sonsela Member.

Conclusions/Significance

We present a revised five-part model for the Sonsela Member. The units from lowest to highest are: the Camp Butte beds, Lot''s Wife beds, Jasper Forest bed (the Sonsela sandstone)/Rainbow Forest Bed, Jim Camp Wash beds, and Martha''s Butte beds (including the Flattops One sandstones). Although there are numerous degradational/aggradational cycles within the Chinle Formation, a single unconformable horizon within or at the base of the Sonsela Member that can be traced across the entire western United States (the “Tr-4 unconformity”) probably does not exist. The shift from relatively humid and poorly-drained to arid and well-drained climatic conditions began during deposition of the Sonsela Member (low in the Jim Camp Wash beds), well after the Carnian-Norian transition.     

川南布拖县乌科志留纪早期介形类丽足介目(Myodocopida)

乌科剖面位于四川布拖县乌科乡境内。在此剖面的石门坎组上碎屑岩段中产有相当丰富的介形类丽足介目,特别是球茎介科的化石。经研究、鉴定,初步划分出2个属和6个种,其中包括3个新种(Entomozoe bispinosa sp.nov.,E.suboblongasp.nov.,E.wukeensis sp.nov.),1个比较种和2个未定种。在分析上述属、种,特别是相关种的地质、地理分布特征的基础上,并参考其他门类化石(如腕足类动物群)和岩石性质,提出石门坎组上碎屑岩段可能要低于笔石M.crispus带上部,大致相当于志留纪兰多维列世特列奇期早期。Entomozoe和球茎介科的最早代表推测是产自石门坎组上碎屑岩段Entomozoe的那些分子,如E.suboblongasp.nov.,而非E.tuberosa(Jones),1861(或其亲缘种E.aff.tuberosa)(Siveter et al.,1990,2006)。文中还就上述丽足介目动物群的生态环境进行了分析,指出石门坎组上碎屑岩段可能是在陆棚浅水到相对较深水、能量较低的、温暖的开阔海环境下沉积的。     

Evidence of synsedimentary tectonics in the Northern Tabas Block,East-Central Iran: The Callovian (Middle Jurassic) Sikhor Formation

Summary The Sikhor Formation (new) is a predominantly siliciclastic sediment package intercalated between the marly-silty Baghamshah Formation (below) and the calcareous Esfandiar Limestone and Qal'eh Dokhtar Limestone formations (above). All stratigraphic evidence points to an Early Callovian age of the formation. The Sikhor Formation is restricted to the southern and central Shotori Mountains and consists of two members: The Kuh-e-Neygu Member (new) is composed of fluvialdeltaic conglomerates, sandstones, and siltstones grading into marly silt of the Baghamshah Formation. The overlying Majd Member (new) is characterised by mixed siliciclastic-carbonate sediments that record the interfingering of carbonate ramp sediments with fluvialdeltaic sands and silts. Evidence of erosional truncation of the underlying Baghamshah Formation and confinement of the siliciclastic sediment to a comparatively narrow, NNW-SSE elongated strip suggest that the formation had its origin in the asymmetric uplift of a westdipping tilted fault block in the southern Shotori Mountains that shed its sediment predominantly in a northern and eastern direction. After erosional levelling, the former uplifted areas were overgrown by the highly productive Esfandiar Carbonate Platform. The Sikhor Formation thus is evidence of an extensional tectonic pulse in the early Callovian and underlines that this area of the Tabas Block was a tectonically highly unstable area during most of the Jurassic.     

Significance of Devonian-Carboniferous radiolarians from accretionary terranes of the New England orogen, eastern Australia

Radiolarians provide age constraints for many previously undated terranes in the New England Orogen (NEO), a tectonic collage developed along the eastern margin of Australia.Djungati terrane, the age range of which was previously unknown, contains two distinctive siliceous sedimentary lithofacies. The oldest is a thick sequence of red, ribbon-bedded cherts which probably accumulated in a deep ocean-floor setting far from land. Middle Silurian through Late Devonian radiolarians have been recovered from these cherts. Green tuffaceous cherts which contain a latest Devonian (Famennian) radiolarian fauna depositionally overlie the lower red ribbon-bedded chert sequence. These cherts are intercalated with volcaniclastic sediments and the fauna which they contain can be used to constrain the timing of accretion of older rocks into a subduction complex.Anaiwan terrane, which was also previously undated, contains thin ribbon-bedded cherts which are depositionally overlain by tuffaceous chert, siliceous siltstones and volcaniclastic sediments. Latest Devonian (?late Famennian) and Early Carboniferous radiolarians have been recovered from these cherts and tuffaceous siltstones.Radiolarians also occur in fine-grained siliceous sediments of the Yarrimie Formation, part of the Gamilaroi terrane. These radiolarians are of Late Devonian (Frasnian) affinity and their presence indicates that blocks of limestone, which contain Givetian conodonts and corals and were previously thought to indicate the age of the Yarrimie Formation, are allochthonous.     

Brachiopods from the Cisuralian–Guadalupian of Darvaz,Tajikistan and implications for Permian stratigraphic correlations

In this paper, we describe the upper Cisuralian Safetdara and Gundara formations of the Darvaz mountains, North Pamir, which were part of the Kunlun Arc, developed along the active Eurasian margin. The Safetdara Formation comprises massive limestones (mainly cyanobacterial, Tubiphytes and Archaeolithoporella boundstones) alternating with well-bedded bioclastic and oncoidal limestones and an interval of recessive shales. The formation crops out above the Chelamchi Formation consisting of turbiditic siltstones and sandstones with bioclastic silty limestones yielding massive limestone olistoliths. The Gundara Formation consists of fine sandstones at the base, followed by well-bedded marly bioclastic, oncoidal and microbial limestones, bearing a rich silicified brachiopod fauna in life-position. Two new taxa have been identified in this association: the cemented coralliform Gundaria insolita n. gen. n. sp. and the pedicle attached Hemileurus politus n. sp. The inferred environmental setting is that of shoal deposits of warm, shallow, high energy, clear marine waters for the Safetdara Formation. The agglutinated microbial reefs to cluster reefs of the Gundara Formation were probably growing in a muddier, quieter and probably slightly deeper setting.The foraminifers of the Brevaxina Zone suggest a Bolorian age for the top of the Chelamchi Formation, the Safetdara Formation and the base of the Gundara Formation. Kungurian conodonts have been found in the lower part of the Safetdara Formation. The biostratigraphic data from the sedimentary succession of North Pamir, integrated with those already obtained from Southeast Pamir, allow to refine the correlations between the Tethyan regional scale and the International Time Scale. In particular, it seems now clear that the Bolorian and the lower part of the Kubergandian correlate to the Kungurian.     

Crocodylian Tracks from Lower Oligocene Flysch deposits of the Barail Group,Manipur, India

A marine tidal delta siltstone from Gelmon locality in Northeast India preserved three crocodylian footprints and an elongate depression that appears to be a tail drag mark. Similar drag marks occur in nearby bedding surfaces. The discovery of crocodylian tracks from the basal part of Laisong Formation, Barail Group (Late Eocene–Early Oligocene age) of Manipur, India is noteworthy because of the age and the geographic location. Crocodylian tracks are rare in Cenozoic formations and they have not previously been reported from Asia. The footprints are herein named as a new ichnogenus and ichnospecies, Indosuchipes manipurensis.     

新疆若羌石灰华中的植物化石及古环境*

根据首次从富阳西湖组发现的孢子及其垂直分布特征,划分出下段 LH 组合和上段 DP 组合,分别归入晚泥盆世法门晚期(斯图年期)和早石炭世早期(杜内早期),即泥盆-石炭系界线通过西湖组内部,与江苏五通组类似.     

Facies pattern and sea-level dynamics of the early Late Cretaceous transgression: a case study from the lower Danubian Cretaceous Group (Bavaria,southern Germany)

The facies development and onlap pattern of the lower Danubian Cretaceous Group (Bavaria, southern Germany) have been evaluated based on detailed logging, subdivision, and correlation of four key sections using an integrated stratigraphic approach as well as litho-, bio-, and microfacies analyses. Contrary to statements in the literature, the transgressive onlap of the Regensburg Formation started in the Regensburg–Kelheim area already in the early Early Cenomanian Mantelliceras mantelli ammonite Zone and not in the Late Cenomanian. In the Early Cenomanian, nearshore glauconitic-bioclastic sandstones prevailed (Saal Member), followed by Middle to lower Upper Cenomanian mid-shelf siliceous carbonates intercalated with fine-sandy to silty marls (Bad Abbach Member). Starting in the mid-Late Cenomanian (Metoicoceras geslinianum ammonite Zone), a considerable deepening pulse during the Cenomanian–Turonian Boundary Event (CTBE) initiated the deposition of the deeper shelf silty marls of the Eibrunn Formation, which range into the early Early Turonian. During the CTBE transgression, also the proximal Bodenwöhrer Senke (ca. 40 km NE of Regensburg) was flooded, indicated by the onlap of the Regensburg Formation onto Variscan granites of the Bohemian Massif, overlain by a thin tongue of lowermost Turonian Eibrunn Formation. A detailed record of the positive δ¹³C excursion of the global Oceanic Anoxic Event (OAE) 2 has been retrieved from this shallow-water setting. An integrated approach of bio-, event-, carbon stable isotope and sequence stratigraphy was applied to correlate the sections and to decipher the dynamics of this overall transgressive depositional system. The Cenomanian successions show five prominent unconformities, which correlate with those being known from basins in Europe and elsewhere, indicating their eustatic origin. The rate of sea-level rise during the CTBE suggests glacio-eustasy as a driving mechanism for Late Cenomanian sea-level changes. The Regensburg and Eibrunn formations of the lower Danubian Cretaceous Group are highly diachronous lithostratigraphic units. Their regional distribution and northeast-directed onlap pattern onto the southwestern margin of the Bohemian Massif can readily be explained by the lateral movements of roughly coast-parallel (i.e., NW/SE-trending) facies belts of a graded shelf system transgressing on a northeastward-rising substrate. It took the Cenomanian coastline ca. 6 Ma to transgress from southwest of Regensburg to the topographically elevated granite cliffs southeast of Roding in the Bodenwöhrer Senke (=60 km distance).     

Stratigraphy and microfacies of the oligocene sequence at Gabal Bu Husah, Marada Oasis, South Sirte Basin, Libya

Summary Stratigraphic and microfacies investigations carried out on the Oligocene sequence exposed at Gabal Bu Husah, northwest of Marada Oasis, south of the Sirte Basin, Libya, showed that the Oligocene sequence conformably overlies the Late Eocene Thamat Formation and unconformably underlies the Early to Middle Miocene Marada Formation (Qarat Jahannam Member). The lithostratigraphic studies of the Oligocene sequence revealed the presence of two rock units, from base to top: 1- Umm Ad Dahiy Formation (Early Oligocene, Rupelian) and 2- Bu Hashish Formation (Late Oligocene, Chattian). The Oligocene sequence yields a rich foraminiferal assemblage with fifty-one benthonic and large formaminiferal species. The foraminiferal analysis allowed to subdivide the sequence into three local foraminiferal assemblage zones, arranged stratigraphically from base to top: 1)Elphidium minutum zone comprising the Umm Ad Dahiy Formation. 2)Miogypsinoides complanatus/Nonion granosus assemblage zone covering the lower two thirds of the Bu Hashish Formation. 3) Zone with abundantNummulites spp., including the upper part of the Bu Hashish Formation. The paleoenvironmental significance of the recorded species is described and discussed. The microfacies studies led to the recognition of eleven microfacies types. These microfacies indicate that the lower part of the Umm Ad Dahiy Formation was deposited in a shallow warm marine environment, but the conditions changed to fluviomarine in the uppermost part. The Bu Hashish Formation was deposited in a shallow-marine, inner-shelf environment (as indicated by the operculinid limestone) but a probable hiatus in its middle part is indicated by the presence of a nummulitic coquina and gypsum beds formed in a lagoonal environment. After returning to a shallow marine environment at the end of Oligocene the marine sedimentation ended.     

Conodontes de la coupe de Reun Ar C'Hrank en Lanvéoc (Rade de Brest) emsien du massif Armoricain (France) corrélations biostratigraphiques

In the Reun ar C'Hrank section (Rade de Brest), the Conodonts from just above the base of the Beg an Arreun Formation of the Troaon Group in the Armorican Massif indicate the Polygnathus laticostatus Zone, which belongs to the highest Zlichvian and the base of the Dalejan. Species of Icriodus from the same beds indicate a correlation with the lower and middle part of the Hierges Formation, which represents the Upper Emsian in the Ardennes. Conodonts from Le Fret Formation are tentatively assigned an Upper Emsian/Dalejan age.     

新疆昆仑山黄羊岭—木孜塔格一带早二叠世的(竹蜓)

本文报道了新疆昆仑山黄羊岭—木孜塔格地区下二叠统黄羊岭群的分布、层序划分及对比;首次对黄羊岭群所含?类进行了研究和描述。共描述早三叠世?类15属26种,其中4个新种,1个新亚种,2个未定种。 黄羊岭群下亚群和华南的栖霞组大致可以对比,黄羊岭群上亚群和华南茅口组大致可以对比。黄羊岭群的早二叠世?类,属典型的特提斯海暖水?类,生物地理分区上属特提斯大区的华夏-特提斯区。     

Facies features and paleoenvironmental reconstruction of the Early to Middle Devonian syn-rift volcano-sedimentary succession (Padeha Formation) in the Eastern-Alborz Mountains,NE Iran

The Padeha Formation in the Eastern-Alborz Mountains (northeast Iran) is an Early to Middle Devonian syn-rift succession. Siliciclastic rocks (conglomerates, sandstones, mudstones, and red paleosols) to non-marine carbonates (yellowish calcretes, dolocretes, stromatolites, and sandy bivalve packstone) associated with volcaniclastic rocks (basalts or andesites and tuffs) are present in this succession. Facies analysis led to the recognition of three facies associations that are deposited in three terrestrial environments (alluvial fan, distal fan, and palustrine/lacustrine). Its sedimentary fill pattern and association with mafic volcaniclastic rocks indicate that this formation is related to the initial phase of a rift basin (intracratonic rift). Paleosols of this formation are good indicators for recognition of paleoenvironmental factors (climatic changes from semi-arid to sub-humid with annual fluctuation, small and short gross-like vegetation cover, subaerial exposure, very shallow lake, volcanic source rocks, and early meteoric diagenesis).     

海南岛早志留世晚期腕足类Xinanospirifer的发现--兼论南好组

海南岛保亭县毛感乡南兵至南好公路边南好组以往被确认为下石炭统岩关阶 ,并认为与其下的上志留统足赛岭组呈角度不整合接触。著者最近在该剖面南好组中发现兰多维列世特里奇期晚期 (LateTelychian)Xi nanospirifer腕足动物群和三叶虫Latiproetuscf.latilimbatus,证明久归于下石炭统岩关阶南好组的地质时代应改归于早志留世 (Llandoverian) ;海南岛地区在早志留世明显属于扬子地台区的范畴 ;从地质时间上还暗示南好组与其下伏的足赛岭组不可能存在角度不整合接触 ;     

浙江桐庐早二叠世船山组介形类化石

研究报道的介形类化石系采自浙江省桐庐县瑶林镇沈村剖面早二叠世船山组,计14 属4 亚属22 种,包括4新种和4未定种。其中12属4亚属和19种产自船山组第三段,并被归为Hollinella(Praehollinella) ema ciata Basslerella ola 组合,代表早二叠世船山世隆林期。牙形刺化石主要产自船山组第一段,带化石Streptogna thodus elegantulus 及其共生的其它牙形刺化石,指明船山组第一段的时代为中、晚卡西莫夫期到早格泽里期。根据介形类动物群生态组合和岩石特征,推测船山组第三段是在温暖的近岸浅水且为低能的开阔海环境下沉积的。     

Lithostratigraphy, petrography and facies analysis of the Late Cenozoic sediments in the foreland basin of the West Kunlun

The Yecheng profile, lying in the southwest Tarim Basin at the northern foot of the West Kunlun Mountains, comprises 4.5 km of conformable Miocene to Pliocene strata. The lower part of the section, the Wuqia Group, is composed of interbedded red mudstone and pale-coloured fine sandstone with a thickness of 1700 m. The Artux Formation is 800 m thick and composed of mudstone, sandstone with thin gravel and conglomerate beds. The upper part of the section, known as the Xiyu Formation, consists of 2000 m of cobble and boulder conglomerate intercalated with massive siltstone lenses.Compositional study of the sandstones in the Wuqia Group and Artux Formation indicates that they were sourced from low relief areas of the Kunlun region and probably further south from Tibet. The provenance of the conglomerate in the Xiyu Formation is the West Kunlun Mountains. Compositional trends within the conglomerate indicate that Upper Palaeozoic marine, and Mesozoic to Tertiary terrestrial silicic rocks were eroded first, along with the Proterozoic to Lower Palaeozoic Proto-Tethys metasedimentary rocks. Erosion into deeper levels of the Kunlun Mountains provided igneous and high-grade metamorphic sediment, which first appears 640 m above the base of the Xiyu Formation.Lithofacies change from fine-grained mudstone and sandstone to coarse clasts coincides with the onset of aeolian sedimentation, indicating major shift of regional palaeoclimatic regime. Although climatic changes may have played an important role in controlling the sedimentary regime worldwide, our study of the lithostratigraphy and petrography of the Yecheng section suggests that the lithofacies change recorded the progressive unroofing history of the source rocks in the West Kunlun Mountains.     

湖北宜昌地区晚奥陶世三叶虫动物群及分带序列

从湖北宜昌黄花场和普溪河桥宝塔组剖面获得的新材料,极大地丰富了华南晚奥陶世三叶虫动物群内容,包括22属、1亚属、31种之多。新材料保存完美,很大程度上完整了有关三叶虫的形态特征。RemopleuridesamphitryonoidesLu,1975一种由于新发现了活动颊、唇瓣、胸部及尾部,证明它确实是Remopleurides的成员,排除了对该种归属的怀疑。文中建立了1新属和2新种:Pentacopygepuxiheensisgen.etsp.nov.,Lamproscutellum?limbatumsp.nov.。同时,以桨肋虫类为基础在宝塔组Nankinolithus带之下进一步划分出4个三叶虫带,从下到上依次为Pentacopygepuxiheensis间隔带,Hexacopygeyichangensis间隔带,Remopleuridesamphitryonoides延限带(普溪河段,卡拉道克中期),和Dislobosaspisguizhouensis延限带(临湘段下部,卡拉道克晚期)。     

Facies analysis of the old Red Sandstone of Spitsbergen (Wood Bay Formation): Reconstruction of the depositional environments and implications of basin development

Summary The object of this paper is to provide a facies guide that helps subsequent workers recognizing different facies and palaeoenvironments of the terrestrial old Red Sandstone exposed in central N Spitsbergen. Sediments of the Early Devonian Wood Bay Formation represent continental molasse deposition under arid to semi-arid climatic conditions in three main depositional environments (rivers, alluvial plains, perennial lakes) represented by characteristic lithologies, sedimentary structures and microfacies. River channel deposits, consisting of conglomerates, very coarse-to very fine-grained sandstones and sandy siltstones are generally arranged in successive fining-upward sequences marked by specific sedimentary structures in a particular order. The sediments reflect the bed, mixed and suspended load of low sinuosity braided to high sinuosity meandering rivers. Adjacent overbank areas (levees, crevasse splays) and extensive alluvial plains are represented by reddish-brown, fine-grained sandstones to sandy siltstones and silty mudstones. The sediments are the accumulated, mixed and suspended load of floodwaters, which recurrently inundated the lowlands. Various lithofacies types, facies associations and sequences reflect the local formation of palaeosols, ephemeral flood lakes and boggy areas with persistent high water tables. Multicoloured, calcareous successions and single lime-stone beds intercalated in the red beds represent the depositional products of perennial, low relief, low-energy lakes, which developed in topographic depressions of the alluvial plains. The sediments reflect deposition in deep-water, littoral and palustrine sub-environments, forming regressive, shallowing-upward sequences in the central lake areas. The overall arrangement of the various facies reflects a terrestrial basin that developed from a steep-sloped, high-energy, river-dominated molasse trough into a level, lowenergy coastal plain, marked by broad mud flats and the formation of extensive perennial lakes.     

Martharaptor greenriverensis, a New Theropod Dinosaur from the Lower Cretaceous of Utah

Background

The Yellow Cat Member of the Cedar Mountain Formation (Early Cretaceous, Barremian?) of Utah has yielded a rich dinosaur fauna, including the basal therizinosauroid theropod Falcarius utahensis at its base. Recent excavation uncovered a new possible therizinosauroid taxon from a higher stratigraphic level in the Cedar Mountain Formation than F. utahensis.

Methodology/Principal Findings

Here we describe a fragmentary skeleton of the new theropod and perform a phylogenetic analysis to determine its phylogenetic position. The skeleton includes fragments of vertebrae, a scapula, forelimb and hindlimb bones, and an ischium. It also includes several well-preserved manual unguals. Manual and pedal morphology show that the specimen is distinct from other theropods from the Cedar Mountain Formation and from previously described therizinosauroids. It is here named as the holotype of a new genus and species, Martharaptor greenriverensis. Phylogenetic analysis places M. greenriverensis within Therizinosauroidea as the sister taxon to Alxasaurus + Therizinosauridae, although support for this placement is weak.

Conclusions/Significance

The new specimen adds to the known dinosaurian fauna of the Yellow Cat Member of the Cedar Mountain Formation. If the phylogenetic placement is correct, it also adds to the known diversity of Therizinosauroidea.     

Probable predation on Upper Cambrian trilobites and its relevance for the extinction of soft-bodied Burgess Shale-type animals

The lower Rabbitkettle Formation of northwestern Canada is a monofacial Upper Cambrian unit of variably calcareous, argillaceous siltstone and fine-grained sandstone with rare bioclastic grainstone, deposited on a gentle slope below fair-weather wave base with no discernible fluctuation in water depth. The trilobite fauna is a mixture of pandemic agnostoids and Laurentian polymeroids, including protaspides and meraspides, and individuals are disarticulated, non-abraded and mostly oriented convex-upward. Bioclasts are interpreted as in situ elements affected only by weak bottom currents and storm-induced turbulence. A major proportion of the larger (≥5 mm across) polymeroid cranidia and pygidia in the lower part (Marjuman) of the formation are broken; large thoracic segments are often broken at the axial furrow and some broken free cheeks occur, but essentially no broken agnostoids or hypostomes were observed. Trilobites are not broken in upper beds (Steptoean), above the base of the Glyptagnostus retculatus Zone. Physical breakage cannot be dismissed entirely, but most damage is interpreted to be due to size-selective predation, possibly through lethal blows similar to those delivered by some extant stomatopod crustaceans. A possible culprit may be an animal akin to Yohoia , known from the Middle Cambrian Burgess Shale. The distribution of attacked trilobites serves as a proxy for the presence and disappearance of soft-bodied carnivores. In the Rabbitkettle Formation, it suggests that Burgess Shale-type animals may have persisted into the Late Cambrian but suffered extinction at the Marjuman-Steptoean 'biomere' event when most trilobite species vanished.