New click beetles (Coleoptera: Elateridae) from the cretaceous of Russia and Kazakhstan

A new tribe, two new genera, and seven new species of click beetles are described: Pollostelaterini, trib. nov., Pollostelater baissensis, gen. et sp. nov. from the Lower Cretaceous of Transbaikalia (Baisa locality), five new species in the genus Cryptocoelus Dolin et Nel C. sinitshenkovae, sp. nov. (Romanovka locality), C. shcherbakovi, sp. nov., C. baissensis, sp. nov., C. lukashevichae, sp. nov., C. dolini, sp. nov. (all from Baisa locality), and Turonelater giganteus, gen. et sp. nov. from the Turonian of Southern Kazakhstan (Kzyl-Dzhar locality).     

A new genus of the subfamily Anaidinae (Coleoptera, Scarabaeoidea, Hybosoridae) from the Mesozoic of Transbaikalia

A new genus and species, Protanaides sibiricus gen et sp. nov., of the subfamily Anaidinae (family Hybosoridae) from the Lower Cretaceous Baisa locality in Transbaikalia is described. This find adds considerably to knowledge of the diversity of this subfamily in the Mesozoic of Asia and gives evidence of the relict status of the Recent Anaidinae.     

New bugs of the family Archegocimicidae (Hemiptera: Heteroptera) from the Upper Jurassic of Mongolia

Two new genera and five new species of Archegocimicidae are described from the Upper Jurassic of the Shar-Teg locality in Mongolia: Shartegocimex rasnitsyni, gen. et sp. nov., S. distans, sp. nov., Shartegocorpus paranotalis, gen. et sp. nov., Saldonia formosa, sp. nov., S. insolita, sp. nov.     

<Emphasis Type="Italic">Istchenkophyton filiciforme</Emphasis> gen. et sp. nov., a new small vascular plant with thick cuticle from the Devonian of Voronezh Region (European Russia)

Remains of a new small vascular plant, Istchenkophyton filiciforme gen. et sp. nov., are described from the Upper Givetian strata of Pavlovsk vicinity (Voronezh Region). The plant is characterized by a very thick cuticle (up to 300 μm), complex branching pattern, similar to that of a frond rachis, anomocytic stomata, and a well-developed vascular bundle with a central elongated in transversal section protoxylem strand and massive metaxylem composed chiefly of scalariform bordered-pitted tracheids. A reconstruction of the new plant is proposed; its ecology and systematic position are considered. In the light of new data relationships of problematic Devonian plants with thick cuticle are discussed.     

Foliar cuticle of Ruflorinia orlandoi nov. sp. (Pteridospermophyta) from the Lower Cretaceous of Patagonia

Foliar cuticle of a new species of Ruflorinia, R. orlandoi Carrizo et Del Fueyo, from the Lower Cretaceous Springhill Formation in Santa Cruz province, Argentina, is herein described with light and electronic (scanning and transmission) microscopy. The fronds of R. orlandoi nov. sp. are bipinnate and imparipinnate with a conspicuous main rachis bearing alternate to sub-opposite pinnae. Cuticular striations and hairs are present in both epidermises whereas stomata are only on the abaxial cuticle. Stomata have several papillate subsidiary cells and the guard cells are sunken. Ultrastructurally the adaxial and abaxial cuticles are composed of three and two cuticular layers, respectively. The cuticular characters here described are interpreted as a response to a possible adaptation to a wet climate as well as to the fluctuating environment produced by the gradual transition from fluvial to an estuarine and marine setting that characterizes the Springhill Formation. The discovery of this new species allows expanding the previous knowledge of the paleofloristic composition of the Springhill Formation with the addition of the first pteridospermous megafossil described for this formation. Moreover, the record of Ruflorinia orlandoi nov. sp. in the Springhill Formation highlights the importance of the Caytoniales during the Cretaceous in Patagonia.     

Antlered richardiid flies: new species of Richardia (Tephritoidea: Richardiidae) with antler‐like genal processes

Within Richardiidae, Richardia infestans, from Colombia, is the only known species to have genal processes, which are structures formed by the extension of the cuticle on the lateral margin of the gena. In the present work, six new species, with different patterns of antler‐like genal processes, are described. An identification key and illustrations are provided. Richardia bella sp. nov. , R. modesta sp. nov. and R. vitta sp. nov. are recorded from Costa Rica; R. advena sp. nov. is recorded from Costa Rica and Panama; R. simplex sp. nov. is recorded from Panama; and R. ornatella sp. nov. is recorded from Ecuador.     

A new subfamily of ichneumonids from the Lower Cretaceous of Transbaikalia and Mongolia (Insecta: Hymenoptera: Ichneumonidae)

A new subfamily of ichneumonids, Palaeoichneumoninae, is described from the Lower Cretaceous of Transbaikalia (Baisa locality) and Mongolia (Bon Tsagan and Kholbotu Gol localities). The new subfamily is intermediate between the archaic subfamily Tanychorinae and the Recent Ichneumonidae. It includes 12 new species, described in three new genera: Palaeoichneumon freja gen. et sp. nov., P. ornatus sp. nov., P. danu sp. nov., P. micron sp. nov., P. mirabilis sp. nov., P. tenebrosus sp. nov., P. townesi sp. nov., Rudimentifera mora gen. et sp. nov., R. suspecta sp. nov., Dischysma maculata gen. et sp. nov., D. similis sp. nov., and D. ramulata sp. nov.     

Revision of the Indo‐African Pachycerus Schoenherr, 1823, with a description of four new species (Coleoptera: Curculionidae: Lixinae)

The Indo‐African species of the genus Pachycerus (Curculionidae: Lixinae: Cleonini) are revised. The identification of Cleonus senegalensis Gyllenhal, 1834 is discussed, its holotype rediscovered and the synonymy between C. senegalensis and Ammocleonus hieroglyphicus (Olivier, 1807) is confirmed. A neotype is established for Pachycerus opimus (Gyllenhal, 1834). Lectotypes for Pachycerus vestitus (Fåhraeus, 1834), Pachycerus badeni (Faust, 1888), and Pachycerus sellatus Faust, 1904 are designated. Pachycerus sahelicus sp. nov. (type locality: Senegal, Bambey), Pachycerus hippali sp. nov. (type locality: Saudi Arabia: Jeddah, Taif), Pachycerus barclayi sp. nov. (type locality: southern India: Manapparai), and Pachycerus simonae sp. nov. (type locality: Morocco, Western Sahara, Cap Boujdour) are described. Some remarks on taxonomy and biogeography of the species are added.     

甘肃酒泉盆地下白垩统Pseudofrenelopsis(掌鳞杉科)的发现及其意义

记述甘肃酒泉盆地早白垩世地层中发现的两种掌鳞杉科植物化石Pseudofrenelopsis dalatzensis(ChowetTsao,1977)Cao ex Zhou和P.gansuensissp.nov.。其中,归入P.dalatzensis的标本无论是小枝形态还是表皮构造都与产于吉林延吉盆地大砬子组的模式标本一致。定为新种的标本小枝较粗,可能只有一枚叶;节间角质层厚25—30μm,表面没有乳突和表皮毛,气孔器和表皮细胞均规则成行排列;气孔器圆形或椭圆形,较大,整体下陷。副卫细胞通常5—7个,少数可达8—9个。保卫细胞仅部分保存。已发现的Pseudofrenelopsis均分布于北半球,时代限于早白垩世,且多为早白垩世晚期,其中P.dalatzensis在模式标本产地与丰富的被子植物共生,应为早白垩世Aptian到Albian期。该种在酒泉盆地中沟组的出现说明该组的时代最高可到Aptian或Albian期。已有资料还表明,Pseudofrenelopsis主要生活于有季节变化、盐度较高或较干旱的环境中,这与酒泉盆地早白垩世沉积物和其他门类的化石所指示的环境特征相符合。     

Review of grylloblattid insects (Insecta: Grylloblattida) from the Solikamsk deposits of the Perm Region

The Solikamsk faunas of the Vorkuta series of the Pechora Basin and the Solikamsk Formation of the Perm Region of Russia are reviewed. New taxa are described from Tyul’kino locality (Solikamsk District): Artinska ufimica sp. nov., Sylvaprisca tyulkinensis sp. nov., Ufimoprisca gen. nov. (the type species Sylvaprisca alba Aristov, 2005), U. mica sp. nov., U. kamensis sp. nov. (family Lemmatophoridae), Sylvaella ovalis sp. nov. (family Liomopteridae), Ivakhosara prima sp. nov., and Megakhosarella prisca sp. nov. (family Megakhosaridae). Additionally, Uralioma sp. is described from Shchugor locality (Cherdyn’ District). Two species, Paraprisca solikamskensis Aristov, 2009 (family Lemmatophoridae) from Tyul’kino and Kolvidelia curta G. Zalessky, 1950 (family Ideliidae) from Pokcha locality (Cherdyn’ District)), are redescribed.     

Ichneumon wasps of the Khasurty locality in Transbaikalia (Insecta, Hymenoptera, Ichneumonidae)

One new genus and four new species of ichneumon wasps are described from the Lower Cretaceous of Buryatia (Khasurty locality): Amplicella mininae, sp. nov., A. shcherbakovi, sp. nov., A. bashkuyevi, sp. nov., Khasurtella buriatica, gen. et sp. nov. The species Tanychora sinensis Zhang, 1991 is transferred to the new genus Khasurtella. Similarity of the ichneumonid faunas of Khasurty and the Chinese localities Yixian and Laiyang indicates the age similarity between these localities. Identification key to genera and species of Tanychorinae is given.     

New beetle species of the formal genus <Emphasis Type="Italic">Artematopodites</Emphasis> (Coleoptera: Polyphaga), with remarks on the taxonomic position of the genera <Emphasis Type="Italic">Ovivagina</Emphasis> and <Emphasis Type="Italic">Sinonitidulina</Emphasis>

Three new beetle species of the formal genus Artematopoides, i.e., A. maximus sp. nov., A. crispulus sp. nov., and A. lepidus sp. nov., from the Mongolian Shar-Teg locality are described. Early Jurassic Ovivagina propinqua Zhang, 1997, O. prolixa Zhang, 1997, and O. insculpta Zhang, 1997 from the Junggar Basin (Xinjiang, China) and Middle Jurassic Sinonitidulina longa Hong, 1983 from the Zhouyingzi locality (Hebei, China) are transferred to Artematopoides, and the three species from Xinjiang are redescribed. Ovivagina immediata Zhang, 1997 is synonymized with O. insculpta Zhang, 1997. A new name, Artematopodites leskoviensis Yan, nom. nov., is proposed for A. longus Ponomarenko, 1990, which proved to be a junior secondary homonym of A. longa (Hong, 1983).     

On Cordaites neimengensis sp. nov from Lower Permian of Zhungeerqi,Inner Mongolia, China

The leaf cuticle of Cordaites neimengensis sp. nov was studied by light and scanning electron microscopy. The specimens were collected from kower Permian of Zhungeerqi, Inner Mongolia (Neimenggu), China. Comparing with other species of Cordaites, it was a new species of the genus, based on the structure of its epidermis. Cordaites neimengensis sp. nov (Pis. Ⅰ to Ⅲ; Text-fig.l,B) The specimen comprises two incomplete leaves over 16 cm in length. One is about 1.4 cm wide at the lower part and 0.3 cm wide at the upper part, with nearly 29 veins per cm and 3 to 5 interstitials. The width of the other leaf is about 7.5 cm at the lower part and 8.2 cm at the upper part with 25 veins per cm and 3 to 5 interstitials. Epidermis amphistomatic. Epidermal cells of upper cuticle aligned in longitudinal rows, nearly rectangular in shape, about 33.6 to 86.4 pm x 10.2 to 28. 8 pm in size. Stomatal apparatus consisting of 2 sunken guard cells surrounded by 2 lateral and 2 polar subsidiary cells. Stomata haplocheillic, about 56.2 pm x 45.5 pm in size, usually arranged in short chains, with one polar subsidiary cell (usually 23.1 to 25.4 pm wide, 34.1 to 39.2 pm long in size) shared with 2 consecutive stomata. The polar subsidiary cells round, oblong or rhomboid in shape. The guard cells reniform or bean-shaped, usually 9.7 to 11.6 pm wide, 23.4 to 29.1 pm long in size. Density and index of stomata about 18/mm2 and 3.2%, respectively. Epidermal cells of lower cuticle also nearly rectangular in shape, about 62.4 to 144 pm × 9.6 to 16.8 pm in size. Stomata on the lower cuticle, haplocheillic, 40.8 pm wide and 48 pm long in size, with a pair of sunken guard cells, which is bean-shaped and surrounded by 2 lateral and polar subsidiary cells. Stomata arranging in parallel bands, typically one, sometimes two. As for the latter, a single row of lateral subsidiary cells (9.2 to 14.4 pm wide, 33.6 to 62.4 pm long in size) is shared with 2 parallel rows of stomata and occasionally also in bands with two rows side by side. In view outside, small papillae on the outer periclinal wall as well as the lateral subsidiary cells. In view inside, some folds along the anticlinal wall but flat about the periclinal wall. Nonstomatic bands usually with 1 to 10 rectangular cell rows. Density and index of the stomata about 209/mm2 and 27.2 %, respectively. 1401%-type: 9342; locality: Heidaigou of Zhungeerqi; Inner Mongolia, North China; Age: Lower Pennian (Shanxi Formation).     

Five novel species of the anamorphic genus Candida in the Cyberlindnera clade isolated from natural substrates in Taiwan

Twelve strains representing five novel yeast species were isolated from natural samples distributed in mountain areas in Taiwan during 2007 and 2009. Sequence analysis of the D1/D2 domain of the large subunit (LSU) rRNA gene revealed that these species are members of the Cyberlindnera clade. These five new species have a greater than 1% difference from their closest relatives in the sequences of the D1/D2 domain of the LSU rRNA gene and were well separated from their closest relatives in terms of physiological characteristics. Moreover, a sexual state could not be found in these five novel yeast species. Therefore, the scientific names of Candida maesa sp. nov. (type strain GJ8L01^T), Candida takata sp. nov. (type strain EN25S01^T), Candida taoyuanica sp. nov. (type strain GY15S07^T), Candida hungchunana sp. nov. (type strain NC3W71^T) and Candida stauntonica sp. nov. (type strain GY13L05^T) were proposed to accommodate these yeasts.     

Study of the Histology of Leafy Axes and Male Cones of Glenrosa carentonensis sp. nov. (Cenomanian Flints of Charente-Maritime,France) Using Synchrotron Microtomography Linked with Palaeoecology

We report exceptionally well-preserved plant remains ascribed to the extinct conifer Glenrosa J. Watson et H.L. Fisher emend. V. Srinivasan inside silica-rich nodules from the Cenomanian of the Font-de-Benon quarry, Charente-Maritime, western France. Remains are preserved in three dimensions and mainly consist of fragmented leafy axes. Pollen cones of this conifer are for the first time reported and in some cases remain connected to leafy stems. Histology of Glenrosa has not previously been observed; here, most of internal tissues and cells are well-preserved and allow us to describe a new species, Glenrosa carentonensis sp. nov., using propagation phase-contrast X-ray synchrotron microtomography, a non-destructive technique. Leafy axes consist of characteristic helically arranged leaves bearing stomatal crypts. Glenrosa carentonensis sp. nov. differs from the other described species in developing a phyllotaxy 8/21, claw-shaped leaves, a thicker cuticle, a higher number of papillae and stomata per crypt. Pollen cones consist of peltate, helically arranged microsporophylls, each of them bearing 6–7 pollen sacs. The new high resolution tomographic approach tested here allows virtual palaeohistology on plants included inside a dense rock to be made. Most tissues of Glenrosa carentonensis sp. nov. are described. Lithological and palaeontological data combined with xerophytic features of Glenrosa carentonensis sp. nov. suggest that this conifer has been adapted to survive in harsh and instable environments such as coastal area exposed to hot, dry conditions.     

A new xeromorphic conifer from the Cretaceous of East China

Newly discovered coniferous shoots from the Cretaceous of Nanjing are attributed to the xeromorphic coniferous genus Glenrosa Watson and Fisher, which is found in China as well as in Eurasia for the first time. G. nanjingensis sp. nov. is known from shoots bearing scale-like leaves with thick epidermal cuticle and hypodermis. Stomata are confined to the abaxial surface deeply sunken in stomatal chambers which are filled with papillae. The cuticle, which has been sectioned for TEM, consists of an outer pale amorphous cuticle proper, usually with a peripheral polylamellate zone, and an inner dark cuticular layer containing reticulate fibrillae. G. nanjingensis sp. nov. bears a close resemblance to the type species G. texensis (Fontaine) Watson and Fisher from North America in gross morphology and especially in the characteristic stomatal chambers, but differs in phyllotaxis and in having the hypostomatic leaf with an obtuse apex. G. nanjingensis sp. nov. is believed to have grown near inland marshes or water bodies under hot and arid climates.     

Origin and development of stomatal microanatomy in two species ofEucalyptus

Summary Development of the stomata ofEucalyptus orbifolia (in which they are relatively superficial) andE. incrassata (in which they are deeply sunken) is described from light microscopy of thin sections of resin-embedded material. The envelope of the guard mother cell is retained intact while in the daughter cells (guard cells) the inner and outer thickenings are formed. The mother cell envelope may even remain discrete and intact during early stages of formation of the separation spaces, precursors of the future stomatal pore, between the thickenings. Remnants of the guard mother cell wall may be retained as parts of at least the inner stomatal ledges. Likewise, remnants of the wall which divides the mother cell persist on the maturing guard cells.Sudan III-positive materials, probably cutin, are removed from the cuticle over the mother cell soon after it is formed. The cuticle above the guard cell is finally perforated by enzymic attack forming, inE. incrassata, a large cavity outside the developing stoma into which the outer stomatal ledges grow as extensions of the upper guard cell walls.The termostiole is suggested for the aperture in the cuticle. The flanges of cuticle seen in section to bound it are termedostiolar ledges. The ostiolar ledges are to be distinguished from the outer stomatal ledges, which develop from the upper thickenings of the guard cell initials. The distinction is clear inE. incrassata (and other species with deeply sunken stomata) but not in mesophytic plants or species with superficial stomata such asE. orbifolia in which the outer stomatal ledges are fused with the cuticle.Growth of the outer stomatal ledges inE. incrassata involves transport of wall materials through an annular space, the equivalent of an ectocythode.The relevance of the observations to stomatal development in other genera is discussed.     

New species of Lonchaeidae (Diptera: Schizophora) from Japan and a re‐evaluation of genus Setisquamalonchaea Morge

Three new species of Lonchaeidae are described from material reared from fig syconia in Japan: Silba inubiwa sp. nov., S. ishigaki sp. nov. and S. japonica sp. nov. Based on the taxonomy of these and similar species the boundaries of the genus Silba are re‐evaluated, and, as a result, it is considered that Setisquamalonchaea Morge is synonymous with Silba Macquart. A revised key and generic notes on the tribe Lonchaeini are provided.     

The Strongylophthalmyiidae (Diptera) of Papua New Guinea,with descriptions of five new species and a world checklist

Eight species of the Strongylophthalmyiidae are reported from Papua New Guinea. Of these, five species (Strongylophthalmyia gigantica sp. nov., S. papuana sp. nov., S. rubella sp. nov., S. sedlaceki sp. nov. and S. shatalkini sp. nov.) are described as new to science. Strongylophthalmyia puncticollis Frey is recorded for the first time from Papua New Guinea. Faunistic remarks, a key to the species of the Strongylophthalmyia in Papua New Guinea and a world checklist are provided.     

Ammonites of Tethyan origin from the Ryazanian of the Russian platform: Genus <Emphasis Type="Italic">Subalpinites</Emphasis> Mazenot

The ammonite genus Subalpinites is known from the Berriasian of southern and central Europe, North Africa, Mangyshlak, and Ryazanian Stage of Central Russia. New species are described: S. gruendeli sp. nov., S. faurieformis sp. nov., S. remaneiformis sp. nov. These species and also S. krischtafowitschi Mitta characterize the lower subzone of the Riasanites rjasanensis Zone in the Moscow Region. The upper subzone of this zone in the Ryazan Region contained members of Subalpinites identified in open nomenclature.