Age,facies, and geometry of the Sandbian/Katian (Upper Ordovician) pelmatozoan-bryozoan-receptaculitid reefs of the Vasalemma Formation,northern Estonia

The Vasalemma Formation (early Katian, Late Ordovician) of northern Estonia consists of a succession of biodetrital grainstones up to 15 m thick with numerous intercalated reef bodies, which reach diameters of more than 50 m. Four dominant facies types are distinguished within the reef core limestones: (1) a bryozoan framestone—bindstone, (2) an echinoderm bindstone, (3) a receptaculitid-bryozoan-microbial framestone, and (4) a tabulate bafflestone. A linking theme between the different reef-core limestones is the presence of clotted microbial bindstone, which in some places contains spicules. Except for the tabulate bafflestone, all facies types occur in the youngest and oldest intervals of reef growth. Generally, a tendency can be observed with a dominance of echinoderm framestone low in the formation and at the base of individual reefs, towards a more receptaculitid dominated facies at the top of the formation. The reefs developed in a narrow, ca. 20-km-long and max. 5-km-wide band on a shallow NE–SW-directed platform in the central part of the North Estonian Confacies Belt. Reef growth can be constrained toward the latest Keila age, representing the rising limb and the peak interval of the Guttenberg Isotopic Carbon Excursion (GICE). Reef termination falls within a second-order sea-level lowstand, the Frognerkilen Lowstand Event, which led to partial subaerial exposure of the reefs. The dead reefs subsequently and rapidly drowned during the Nakkholm Drowning Event at the Oandu/Rakvere Stage. This timing is nearly equivalent to a phase of enhanced reef development elsewhere in Baltica and probably is related to locally increased nutrient availability during the GICE interval.     

Brachiopod morphology,life mode,and crushing predation in the Ordovician Arnheim Formation,Kentucky, USA

Predation on ancient shelled prey is an often-studied topic in paleoecology, but the early Paleozoic and the brachiopods that dominated the seafloor at that time are relatively underrepresented in the predation literature. We assessed predatory repair scar frequencies among the brachiopod genera from the Early Richmondian (Late Ordovician) Oregonia Member (Arnheim Formation) near Flemingsburg, Kentucky. We found higher repair frequencies on the concavo-convex Rafinesquina and Leptaena relative to the bi-convex genera. There were no trends in repair frequency through the stratigraphic section and no relationships between repair frequency and community diversity metrics. It is possible that concavo-convex brachiopods’ flat shape, thin shell profile, and free-lying (no pedicle attachment) lifestyle made them more likely or appealing targets of Ordovician crushing predators. It is also possible that concavo-convex brachiopods were better suited to survive crushing attacks than biconvex taxa. We also found differences in shell ornament that may influence the visibility of repair scars.     

Upper Ordovician bryozoans from the Xiazhen Formation of Yushan,northeastern Jiangxi,East China

The bryozoan fauna from the Xiazhen Formation (Katian, Upper Ordovician) of northeast Jiangxi Province, southeast China is reported here. Seventeen species of bryozoans belonging to fifteen genera and four orders are identified: Homotrypa yushanensis, Homotrypa sp., Prasopora yushanensis, Trematopora sp., Monotrypella sp., Rhombotrypa sp., Orbignyella sp., Constellaria jiangxiensis, Constellaria sp., Stictopora nicholsoni, Trigonodictya parvula, Ptilodictya ensiformis, Stictoporella sp., Pseudopachydictya sp., Nematopora sp., Arthrostylidae sp. indet., and Chasmatoporidae sp. indet. Four of these genera have been reported previously but nine genera (Trematopora, Monotrypella, Rhombotrypa, Orbignyella, Trigonodictya, Ptilodictya, Stictoporella, Pseudopachydictya, and Nematopora), one rhabdomesine and one fenestrate are found for the first time in the Late Ordovician strata of South China. Our palaeogeographical analysis suggests that the bryozoan association is typical for the Katian, which is mostly widespread in Laurentia, Siberia, Baltica and Mediterranean, and displays palaeobiogeographical relationships to the Laurentia–Siberia Province.     

An Early Ordovician Trilobite assemblage from the Lashkarak Formation, Damghan area, northern Iran

A low diversity Lower Ordovician trilobite assemblage is described from the middle part of the Lashkarak Formation at Simeh-Kuh, northwest of Damghan, northern Iran. Co-occurrence with conodonts of the lowermost Paroistodusproteus Biozone suggests an uppermost Tremadocian age. This assemblage includes four taxa, including one new genus and two new species, Damghanampyxginteri nov. gen. nov. sp. and Asaphellusfecundus nov. sp. The third diagnosable species from the formation is Taihungshaniamiqueli (Bergeron, 1894) which has previously been reported from southern France and the Turkish Taurides, suggesting north Gondwanan faunal affinities.     

新疆轮南油田上奥陶统良里塔格组生物类群和古生态地层学*

南疆塔里木板块北部轮南油田在上奥陶统良里塔格组沉积期显示大致由东向西加深的陆表海缓坡地貌,19口钻井岩芯灰岩中可鉴定出钙质微生物Renalcis、Girvanella、Wetheredella、Garwoodia、Phacelophyton yushanensis、Hedstroemia?和Halysis;钙藻类Vermiporella、Dasyporella、Plexavaria和Solenopora;后生动物类型含海绵、珊瑚、苔藓虫、三叶虫、腕足动物、棘皮动物、腹足类、介形类等。不同钻井之间的古生态地层单元对比显示生物礁和灰泥丘主要发育于轮南油田西部井区,而东部井区则多为台内碎屑滩相沉积;海底地貌差异,特别是向西水深加大可视为控制轮南油田良里塔格组生态组合展布的主要因素。     

Some ostracodes from the Gur'yanovka Formation (Upper Ordovician) of northeastern Gorny Altai

Ostracodes from the Gur'yano-ka Formation (Upper Ordo-ician, Sandbian and Katian stages) of Gorny Altai, Russia, are studied for the first time. Some localities with fossil ostracodes of this age are described briefly. Ele-en new species of the genera Ballardina, Eurychilina, Laccochilina, Bolbina, Retiprimites, Pseudozygobolbina, Bullaeferum, Soanella, Egoro-ella, and Allertemmysa are described and figured. The pre-iously described species Leperditella aff. par-ipunctata V. Ivanova, 1959, Grammolomatella valdari V. Ivanova et Melnikova 1977, Longidorsa? baltica Meidla, 1996, and Steusloffina cuneata (Steusloff, 1895) and ostracodes in open nomenclature, Bollia sp., Klimphores sp., Rectella sp., Longiscula sp., are only figured.     

Cryptostomid Bryozoans From The Sassito Formation,Upper Ordovician Cool‐Water Carbonates of the Argentinean Precordillera

Abstract: Two new bryozoan species are described from the Upper Ordovician Sassito Formation of the Argentinean Precordillera: Moyerella  spinata sp. nov. and Phylloporina  sassitoensis sp. nov. The bryozoans are found in cool‐water carbonates. The Silurian genus Moyerella is reported the first time in the Ordovician, showing palaeobiogeographic connections with Estonia and Siberia.     

江西玉山上奥陶统凯迪阶三衢山组的丛花海绵(Corymbospongia)*

串管海绵在中–晚奥陶世经历了的首次辐射演化事件, 在北美、新南威尔士、哈萨克斯坦、西伯利亚和中国西北等地的上奥陶统地层中皆有报道, 但研究程度较低。本文系华南奥陶系串管海绵首次系统古生物学的报道, 详细描述江西玉山上奥陶统凯迪阶三衢山组的Corymbospongia (丛花海绵属)化石, 并基于C. amplia Rigby, Karl, Blodgett & Baichtal, 2005和C. mica Rigby & Potter, 1986的标本展示该属化石外壁的微孔等特征性结构, 这2个种具有跨板块的生物地理分布。综合串管海绵产出层的碳酸盐岩微相以及伴生钙藻和珊瑚化石等群落分子的指相参照, 识别C. amplia和C. mica均具有明显的生态专属性: C. amplia为造礁生物, 在小型微生物岩中出现; C. mica则代表适应于更浅水生境的平地群落代表。本研究将为从底栖生物群落角度开展扬子区生物地层学及古地理重建提供进一步的依据。     

Upper Ordovician sequences of western Estonia

The Upper Ordovician (uppermost Caradoc-Ashgill) section of western Estonia consists of a series of seven open-shelf carbonate sequences. Depositional facies grade laterally through a series of shelf-to-basin facies belts: grain-supported facies (shallow shelf), mixed facies (middle shelf), mud-supported facies (deep shelf and slope) and black shale facies (basin). Locally, a stromatactis mud mound occurs in a middle-to-deep shelf position. Shallow-to-deep shelf facies occur widely across the Estonian Shelf and grade laterally through a transitional (slope) belt into the basinal deposits of the Livonian Basin.

Each sequence consists of a shallowing-upward, prograding facies succession. Sequences 1 (Upper Nabala Stage) and 2 (Vormsi Stage) record step-wise drowning of underlying shelf units (lower Nabala) that culminated in the deposition of the most basinal facies (Fjäcka Shale) in the Livonian Basin. Sequences 3–6 comprise the overlying Pirgu Stage and record the gradual expansion of shallow and middle-shelf facies across the Estonian Shelf. The Porkuni Stage (sequence 7) is bracketed by erosional surfaces and contains the shallowest-water facies of the preserved strata. The uppermost part of the section (Normalograptus persculptus biozone) is restricted to the Livonian Basin, and includes redeposited carbonate and siliciclastic grains; it is the lowstand systems tract of the lowest Silurian sequence 8. Sequence 7 and the overlying basinal redeposited material (i.e., the lowstand of sequence 8) correspond to the latest Ordovician (Hirnantian) glacial interval, and the bracketing unconformities are interpreted as the widely recognized early and late Hirnantian glacial maximums.

The sequences appear correlative to Upper Ordovician sequences in Laurentia. Graptolite biozones indicated that the Estonian sequences are equivalent to carbonate ramp sequences in the western United States (Great Basin) and mixed carbonate-siliciclastic sequences in the eastern United States (Appalachian Basin–Cincinnati Arch region). These correlations indicate a strong eustatic control over sequence development despite the contrasting tectonic settings of these basins.     

Upper Ordovician microphytoplankton of the Bill's Creek Shale and Stonington Formation,Upper Peninsula of Michigan,U.S.A.: Biostratigraphy and paleogeographic significance

An abundant, diverse, and well-preserved organic-walled microphytoplankton assemblage is described from the Upper Ordovician Bill's Creek Shale and the lower Stonington Formation (Bay de Noc Member) in the Upper Peninsula of Michigan, U.S.A. Based on graptolite and conodont evidence, the Bill's Creek Shale and Stonington Formation are Richmondian (=Ashgill) in age. The assemblage is dominated by acritarchs, which comprise 29 species (including the enigmatic palynomorph Gloeocapsomorpha prisca) assigned to 20 genera. The prasinophyte phycomata are represented by undifferentiated species of Leiosphaeridia and Tasmanites. In addition, chitinozoans are abundant, and scolecodonts and graptolite fragments are common. Paleontologic-palynologic and sedimentologic evidence indicates that the Bill's Creek Shale was deposited in a low-energy, shallow, nearshore marine environment. The overlying Bay de Noc Member of the Stonington Formation also accumulated in a low-energy, normal marine environment, but in a more offshore, somewhat deeper water setting. Both formations experienced minor transgressive and regressive episodes as indicated by fluctuations in the composition of the palynoflora. The combined Bill's Creek/Stonington acritarch assemblage closely resembles those described from the Richmondian-aged Maquoketa Shale (Missouri and Kansas), Sylvan Shale (Oklahoma), and Vauréal Formation (Anticosti Island, Québec, Canada). The overall composition of the acritarch assemblage from these four formations reflects a distinctive, recognizably Laurentian character. Nonetheless, many of the Bill's Creek/Stonington acritarchs have been reported from Upper Ordovician localities elsewhere, providing additional evidence for Late Ordovician cosmopolitanism of the marine microphytoplankton community. Additionally, the restricted stratigraphic range of many of the taxa further enhances their biostratigraphic application, both regionally and globally, and reaffirms the Richmondian (=Ashgill) age of the Bill's Creek Shale and Stonington Formation.     

Oldest known Dicoelosia and Epitomyonia,deep water brachiopods from the Beiguoshan Formation (Middle Katian,Upper Ordovician), Shaanxi,north China

Abstract: A diverse brachiopod fauna from a relatively deep water carbonate facies of the Upper Ordovician Beiguoshan Formation (uppermost Caradoc – lower Ashgill, middle Katian) is characterized by small shells and contains the oldest known Dicoelosia and Epitomyonia, two diagnostic taxa of deep water brachiopod palaeocommunities during the Late Ordovician and Silurian. Three new species are recognized: Dicoelosia cordiformis sp. nov., Dicoelosia perbrevis sp. nov. and Epitomyonia fui sp. nov. These pioneer forms of the family Dicoelosiidae show a relatively high degree of morphological plasticity. The shells of Dicoelosia from the Beiguoshan Formation range from the typical slender‐lobed form with a concavoconvex profile to the strongly equibiconvex, fat‐lobed morphotype that was not known previously until the late Silurian. The Beiguoshan dicoelosiids point to an important attribute of the deep water brachiopods: small generalists with high morphological plasticity, which make them ideal candidates as progenitors for the evolution of shallow water brachiopod faunas in shelf and platform depositional environments.     

Upper Tremadoc–Lower Arenig? Anisograptid–Dichograptid fauna from the Cabitza Formation (Lower Ordovician,SW Sardinia,Italy)

The discovery of the anisograptid graptolite Araneograptus murrayi (Hall, 1865), the dichograptids Clonograptus (Clonograptus) cf. rigidus (Hall, 1858) and Clonograptus (Clonograptus) cf. multiplex (Nicholson, 1868) and of two unassigned species of Didymograptus in the Late Tremadoc–Early Arenig? sediments of the Fluminese area (SW Sardinia) adds new data on the biostratigraphy and palaeoecological setting of the Cabitza Formation, as well as on the paleobiogeographical distribution of these taxa.     

The graptolite correlation of the North American Upper Ordovician Standard

The North American Upper Ordovician reference standard, the Cincinnatian Series, contains rich shelly microfossil faunas in its type area in the Cincinnati Region but graptolites are uncommon in most of its shallow-water calcareous sediments. Consequently, the graptolite correlation of this key sequence has remained uncertain, in part, even controversial. A review of both previously published recently discovered graptolite Occurrences in the type Cincinnatian, combined with data from the important graptolite successions in Oklahoma New York-Quebec, has not only clarified the graptolite correlation of the Cincinnatian but also added new data on the morphology taxonomy, the vertical horizontal distribution, of several taxa. The information now at hand indicates that the Edenian Stage correlates with the C. spiniferus Zone, the Maysvillian Stage with the C. pygmaeus lower middle P. manitoulinensis Zone, the Richmondian Stage with the upper P. manitoulinensis , the D. complanatus , possibly part of the C. inuiti Zone. Comparison between graptolite conodont biostratigraphic evidence reveals no apparent conflict. Correlations are proposed between Upper Ordovician North American stages, graptolite conodont zones, successions in Texas, Oklahoma, Sweden, European graptolite zones, British series.     

The trace fossil Thalassinoides from the Upper Ordovician of Tuva

Traces of Thalassinoides (the tunnels of unknown burrowing organisms) are described from carbonates of the Khondelensky layers of the Upper Ordovician of Tuva. Hitherto, this fossil was unknown in the Ordovician of the USSR. They demonstrate great similarity with Thalassinoides from coeval deposits of the Great Basin, USA. The traces are assumed to have been made in terrigenous-carbonate sediments deposited on the areas of a gently sloping shelf in quiet water, below wave base. □ Upper Ordovician, terrigenous-carbonate sediments, burrows.     

The trace fossil Sinusichnus from the Upper Ordovician of Anticosti Island,eastern Canada

Branched, sinusoidal burrows assigned to Sinusichnus cf. seilacheri Knaust et al. 2016; occur in Upper Ordovician (Hirnantian) carbonates on Anticosti Island. These are abundant and monospecific at the base of the Oncolite Bed of the uppermost Ellis Bay Formation, marking a regional discontinuity surface during the multiphase Ordovician mass-extinction event. In contrast to Mesozoic and Cenozoic records of Sinusichnus, commonly attributed to the activity of decapod and isopod crustaceans, the burrow systems from Anticosti Island were probably produced by other arthropods of unknown affinity. A combined dwelling, locomotion and feeding behaviour of their trace maker is assumed. This is the oldest record of the ichnogenus Sinusichnus, previously only known since the Triassic, and its first evidence from North America.     

Paleoecology of Sphenothallus on an Upper Ordovician hardground

A hardground from the Upper Ordovician Dillsboro Formation near Dillsboro, Indiana, U.S.A., preserves an assemblage of encrusting and boring fossils on both top and bottom surfaces. The slab is inferred to have been an undercut ledge, and the dominant fossils of the assemblage, holdfasts of the tube-building worm Sphenothallus and trepostome bryozoans, are prevalent on both sides. The clumping of Sphenothallus holdfasts has been statistically demonstrated using a nearest-neighbor technique. Sphenothallus has also been shown to withstand overgrowth in interactions with bryozoans.     

新疆柯坪和甘肃平凉上奥陶统底部附近的牙形刺

新疆柯坪大湾沟和甘肃平凉官司庄剖面是Nemagraptusb gracilis带及上奥陶统底界的重要剖面,研究这两剖面的牙形刺具有十分重要的意义。本文描述了采自新疆柯坪大湾沟和甘肃平凉官司庄剖面上奥陶统底界附近的牙形刺共13属18种。     

Ordovician (Caradoc) Gastropoda of the Katkoyeh Formation, Kerman Province, Iran

Seven taxa of gastropods are described from the Ordovician (Caradoc) upper Katkoyeh Formation of the Kerman region, east-central Iran. Three are named species and four are under open nomenclature. The most abundant is the minute bellerophontiform Tritonophon peeli Horný, 1977, indicative of a shallow water plectonotid community. Shell material is not preserved in this species, but a small, simple bulbous protoconch is present. Two new species are Slehoferia pachyta and Nonorios kleistos, both with prominent thick-shelled conchs. Shell repair is documented in both of these forms, interpreted as a result of failed predation. Other micromorphic species occur in the samples, but only Tropidodiscus sp. and Nonorios? sp. are named. The Kerman assemblage is comparable directly with similar, closely coeval faunas in Bohemia, France, Portugal, Morocco, and Italy, corresponding with the Palaeozoic Mediterranean Province of northern peri-Gondwana.