Late paleozoic and Late Triassic limestones from north Palawan Block (Philippines): Microfacies and paleogeographical implications

Summary Selected Late Paleozoic and Triassic limestone exposures were studied on northern Palawan Island, Philippines, with regard to microfacies, stratigraphy and facies interpretation. Although some of the outcrops were already reported in literature, we present the first detailed microfacies study. Late Paleozoic carbonates in the El Nido area are represented by widley distributed Permian and locally very restricted Carbonifenous limestones. Of particular interest is the first report of Carboniferous limestones in the Philippines dated by fossils. Fusulinids indicate a ‘Middle’ Carboniferous (Moscovian-Kasimovian) age of the Paglugaban Formation only known from Paglugaban Island. The Permian Minilog Formation consists mostly of fusulinid wackestones and dasycladacean wacke-/packstones. Fusulinid datings (neoschwagerinids and verbeekinids) provide a Guadalupian (Wordian-Capitanian) age. The depositional setting of the Middle Permian carbonates corresponds to a distally steepened ramp with biostromes built by alatoconchid bivalves locally associated with richthofeniid brachiopods. Late Triassic limestones occur in isolated exposures on and around Busuanga Island (Calamian Islands). The age of the investigated carbonates is Rhaetian based on the occurrence ofTriasina hantkent Maizon. Microfacies data indicate the existence of reefs (Malajon Island) and carbonate platforms (Kalampisanan Islands, Busuanga Island, Coron Island). Reef boundstones are characterized by abundant solenoporacean red algae, coralline sponges and corals. Platform carbonates yield a broad spectrum of microfacies types, predominantly wacke- and packstones with abundant involutinid foraminifera and some calcareous algae. These facies types correspond to platform carbonates known from other parts of Southeast Asia (Eastern Sulawesi and Banda Basin; Malay Peninsula and Malay Basin). The Philippine platform carbonates were deposited on and around seamounts surrounded by deeper water radiolarian cherts. The new data on facies and age of the Philippine Permian and Triassic carbonates contradict a close paleogeographical connection between the North Palawan Block and South China and arise problems for the currently proposed origin of the North Palawan Block at the paleomargin of South China. We hypothesize that North Palawan was part of the Indochina Block during the Carboniferous and Permian, separated from the Indochina Block during the Middle Permian and collided with the South China Block in the Late Cretaceous.     

Geological significance of the discovery of Middle Triassic (Ladinian) radiolarians from the Hong Hoi Formation of the Lampang Group,Sukhothai Zone,northern Thailand

This paper presents a newly found fauna of Ladinian (Middle Triassic) radiolarians from the Hong Hoi Formation of the Lampang Group, Sukhothai Zone, northern Thailand. The Ladinian age determined by the radiolarian fauna concords with the age previously determined by mollusks. The study section consists of intercalated sandstone and shale in the lower part and a thick conglomerate in the upper part. The radiolarian-bearing siliceous beds are intercalated within the lower unit. The lithic sandstone of the lower unit consists mainly of volcanic rock fragments, quartz, and feldspar, whereas the thickly bedded conglomerate of the upper unit is characterized by abundant gravel-sized clasts of volcanic rock and limestone. These lithic features of the study section suggest that during deposition the Hong Hoi Formation was located near a supply of volcanic materials. A forearc basin close to the Sukhothai Arc would be the most suitable environment for the deposition. The Ladinian age determined by the radiolarian fauna supports the occurrence of intensive volcanic activity within the Sukhothai Arc during the Middle Triassic.     

Permian and Triassic radiolarians from chert breccia in the Nong Prue area,western Thailand: its origin and depositional setting in the Paleotethys

Bedded chert and siliceous shale successions previously regarded as the Silurian–Devonian rock units, distributed in the Nong Prue area, northwest of Kanchanaburi, western Thailand, yielded Lopingian (upper Permian) and Lower–Middle Triassic radiolarians. We found chert breccia layers in northern Nong Prue area, mainly consisting of angular to sub-angular chert clasts with matrices of silt-sized chert grains and clay minerals. We discriminated uppermost Pennsylvanian–Lopingian (upper Carboniferous–upper Permian) and Middle Triassic radiolarian-bearing chert clasts from four different levels of the chert breccia; 28 species of 15 genera with one radiolarian gen. et sp. indet. are identified. On the basis of sedimentary characteristics of the chert breccia, we suggest that the chert breccia is of sedimentary origin. The radiolarian assemblages reported here, together with previously known lithological and paleontological evidence, further indicate that the chert breccia was deposited in the Paleotethys with chert clasts derived from fine grained siliceous rocks in the continental margin to deep ocean basin of the Sibumasu Terrane.     

Mikrofazies karbonischer Kalkgerölle aus dem Paläozoikum des Rif (Marokko): Ein Beitrag zur Paläogeographie der westmediterranen Paläotethys im Karbon

Post-lower Serpukhovian conglomerates with abundant limestone clasts are intercalated within the Carboniferous flysch succession of the Ghomarides (internal zone of the Rif, northern Morocco). They are restricted to two ghomaridic nappes (Beni Hozmar, Akaili; Fig. 1). No limestone-bearing conglomerates are known from the third nappe (Kuhdiat Tizian). 83.4% of the limestone clasts derived from shallow-shelf environments and adjoining proximal slope environments of late Viséan (V3b gamma) to early Serpukhovian (Namurian E1–E2) age. Microfacies types of the inner shelf lagoon and of restricted shelf environments dominate. They indicate a complicated mosaic facies pattern (Fig. 2). 16.6% of the limestone clasts derived from distal slope environments and deep, open marine environments. They are of unknown age except for one Upper Devonian pebble from around the transition do IIα/do IIβ. All are supposedly of late Middle Devonian and Late Devonian age. Their sedimentology as well as the age and the microfacies of the limestone clasts indicate that the conglomerates of the Ghomarides are homologous to the Marbella Formation of the Malaguides (Betic Cordillera, Southern Spain). The conglomerate of Binifaillet (Minorca, Balearic Islands) is also similar. The source area of the conglomerates was the not any more existent Betico-Rifean Block south of the Malaguide-Ghomaride flysch trough. Owing to their facies development in Devonian and Carboniferous times, the nappes of the Ghomarides and the Malaguides can be arranged in a successively more distal position towards the source: nappe of Beni Hozmar—nappe of Kuhdiat Tizian—nappe of Akaili + Malaguides (Fig. 4). Minorca, which received its flysch sediments from the north, seems to be influenced only episodically by conglomerate inputs from the south. The lithofacies development of the Carboniferous in other Paleozoic massifs of the western Mediterranean clearly demonstrates that the Paleotethys of this region was framed by shallow-water carbonate shelves between early Viséan and Moscovian times. These shelves are not preserved in situ, but gravitative carbonate sediments, conglomerates, slide blocks and gravitational nappes with in the flysch troughs confirm their existence. The flysch troughs were situated at active continental margins. They developed initially in the northern Paleotethys realm (southern border of the Southern European plate) starting with the late Viséan; in the southern Paleotethys realm (northern border of the Betico-Rif block) they started in the lowermost Viséan. The appearance of the flysch troughs records the beginning subduction of the Paleotethys towards the north beneath the Southern European plate as well as towards the south beneath the Betico-Rifean block (Fig. 6). Flysch sedimentation was predated by pelagic sedimentation since the earliest Carboniferous. During the Viséan a successive deepening of the western Mediterranean Paleotethys from north to south was recorded in the facies development of the pelagic deposits. The deepening is genetically connected with the earlier start of the subduction at the Betico-Rifean block.     

Mid-eocene climatic change,Southwestern California and Northwestern Baja California

A widespread paleosol of Paleocene and Early Eocene age occurs in southwestern California and northwestern Baja California. The dominant quartz-kaolinite mineralogy and cation-depleted chemistry of the buried soil indicate a humid, tropical paleoclimate similar to the modern equatorial belt. Although the Paleocene—Eocene paleomagnetic latitudes are similar to the modern latitudes (36–37°N to 32–33°N, respectively), rainfall was about 125–190 cm per year and average annual temperature was about 20–25°C in marked contrast to the present annual rainfall of 25 cm and average annual temperature of 16°C.A variety of indicators in the Late Eocene sedimentary succession suggests a change to a semi-arid paleoclimate. The nonmarine portions of the Late Eocene sedimentary record are dominated by a cobble conglomerate lithosome deposited in fluvial, alluvial fan and fan delta systems. Intertongued with the conglomerate is a sandstone lithosome deposited in flood-plain and nearshore marine environments. The conglomerate clasts were transported to the depositional site via a long-distance (200–300 km), moderate gradient, braided river system mostly by flash floods. Characteristic post-depositional, in situ fracturing of conglomerate clasts probably occurred due to salt crystallization.Within the flood plain sandstones, and to a lesser degree the conglomerates, are multiple well-developed caliche horizons of probable pedogenic origin. Clay minerals from the Late Eocene rocks are dominantly vermiculite and smectite with lesser chlorite and illite; this is in marked contrast to the kaolinite that comprises the underlying Early Paleogene lateritic paleosol.The character of the Late Eocene sedimentary succession indicates a semi-arid climate. Rainfall probably did not exceed 63 cm per year; it probably was seasonal and by occasional flash floods. This paleoclimate contrasts markedly with the earlier humid tropical paleoclimate and must indicate a widespread climatic change in late Middle Eocene time.     

The Upper Carboniferous Hamilton Fossil-Lagerstätte in Kansas: a valley-fill, tidally influenced deposit

A diverse assemblage of unusually well-preserved marine, euryhaline, freshwater, and terrestrial fossils (invertebrates, vertebrates, and plants) occurs within an Upper Carboniferous (Stephanian) Konservat Fossil-Lagerstätte near Hamilton, Kansas, USA. The Lagerstätte occurs within a paleovalley that was incised into the surrounding Carboniferous cyclothemic sequence during a time of low sea level, and was then filled-in during a subsequent transgression. The stratigraphically lowest and most voluminous facies within the valley is a cross-bedded, polymictic limestone conglomerate that contains caliche clasts and charcoal fragments as well as some normal-marine fossils apparently in situ. The origin of the conglomerate is enigmatic, but it was probably deposited by a migrating tidal channel. Overlying and interbedded with the conglomerate is an ostracode wackestone that contains plants (primarily seed ferns and ferns), eurypterids, shrimp, brachiopods, bivalves, and rare fish. The ostracode wackestone was deposited in a low-energy, marginal-marine environment. A thin sequence (<1 m thick) of interbedded laminated limestone and mudstone overlies the conglomerate in a small area. This facies contains a well-preserved mixed assemblage of terrestrial (conifers, insects, myriapods, reptile), freshwater (ostracodes), aquatic (amphibians, reptile), brackish or euryhaline (ostracodes, eurypterids, spirorbids, fish), and marine (brachiopods, echinoderms) fossils. Many of the vertebrates are articulated and show no evidence of preburial decay, scavenging, or predation. A few vertebrates exhibit signs of flotation. Most articulated vertebrate specimens exhibit soft-tissue preservation in the form of dark-brown to black early-diagenetic microbialite body outlines (‘skin preservation’) containing fossil bacteria. Rhythmic patterns of lamination thickness variation in the limestones and mudstones indicate that this facies was deposited in a tidal environment. High sedimentation rate and variable salinity (and therefore exclusion of bioturbators and invertebrate scavengers) are interpreted as key elements that led to the excellent preservation of the fossils in this ancient estuarine environment. □Lagerstätte, taphonomy, estuarine, tidal bedding, paleovalley, Carboniferous, Kansas.     

Late Carboniferous intra-arc sediments in the north Chilean Andes: Stratigraphy,paleogeography and paleoclimate

Summary Thick terrestrial Late Carboniferous to Triassic volcanosedimentary successions, a prominent feature of the Chilean and Argentinian High Andes, were formed on the active continental margin of Gondwanaland. Their stratigraphic position and the paleogeographic and paleoclimatic relations to neighbouring successions are poorly defined. A more precise age has been obtained for alluviolimnic intra-arc sediments (Miembro Medio), which are intercalated in the Late Carboniferous-Triassic volcano-sedimentary successions in the Salar de Atacama area of northern Chile. The ostracodesCarbonita cf.pungens andParaparchites sp., which occur in the lower part of the Miembro Medio, are of Late Carboniferous, probably Westphalian age. The diverse taphoflora, which occurs in a higher stratigraphic level than the ostracodes, includes sphenophytes, ferns, gymnosperms and pteridophylls, for which we assume a late Westphalian-Early Permian age. Considering radiometric data of under-and overlying volcanic rocks, a Westphalian-Stephanian (to? Early Permian) age is inferred for the Miembro Medio. Fauna and flora indicate that warm-humid and seasonal climatic conditions existed during the deposition of the lower fossiliferous part of the Miembro Medio. This coincides with the sedimentary paleoclimatic indicators of the Miembro Medio and the climate which was assumed to have predominated in wide parts of the Central and Southern Andes during the Latest Carboniferous.     

Arachnids from the Carboniferous of Russia and Ukraine,and the Permian of Kazakhstan

New finds of Late Palaeozoic arachnids, based on three well-preserved carapaces from the Carboniferous of Russia and Ukraine and one complete, albeit poorly preserved, specimen from the Permian of Kazakhstan, are described. The spider genus Arthrolycosa is reported from the Late Carboniferous (Late Pennsylvanian: Kasimovian–Gzhelian) of Chunya in the Tunguska Basin of Siberia; it is the first find of a spider outside the Carboniferous tropics. Another fossil assigned to the same genus comes from the Late Carboniferous (Early Pennsylvanian: Bashkirian) of Kamensk–Shakhtinsky in the Donets Basin of Russia; it is probably the oldest fossil spider known. A thelyphonid (whip scorpion) carapace is described from the Late Carboniferous (Late Pennsylvanian: Kasimovian) of the adjacent Lugansk Province of the Donets Basin of Ukraine.     

A REVIEW OF TRIASSIC TETRAPOD TRACK ASSEMBLAGES FROM ARGENTINA

Abstract:  This paper contains the first comprehensive ichnotaxonomic review of the Triassic tetrapod track record in Argentina, including previous accounts and new material recently discovered, and an analysis of its composition and stratigraphic distribution. Triassic footprints have been recorded from three basins: the Ischigualasto-Villa Unión and Cuyo basins in north-west Argentina, and the Los Menucos depocentre in northern Patagonia. Most are in successions of Middle Triassic age; a lower number are from the Late Triassic, and there are two records from Early Triassic rocks. The known track types include: Brachychirotherium isp., cf. Brachychirotherium isp., Chirotherium barthii , Dicynodontipus ispp., Grallator isp., Rhynchosauroides isp., Rigalites ischigualastianus , Rigalites isp., Tetrasauropus isp., and bird-like, chirotheriid and unidentified tridactyl footprints. The ichnogenera Gallegosichnus Casamiquela, 1964, Calibarichnus Casamiquela, 1964, Palaciosichnus Casamiquela, 1964 and Stipanicichnus Casamiquela, 1975 are considered to be synonyms of Dicynodontipus (Hornstein, 1876). In addition, the abandonment of the following ichnogenera (and single ichnospecies) that are based on poorly preserved material is suggested: Ingenierichnus sierrai Casamiquela, 1964, Rogerbaletichnus aguilerai Casamiquela, 1964 and Shimmelia chirotheroides Casamiquela, 1964. At least eight Triassic ichnofaunas can be recognized. The most peculiar is that of the Late Triassic Los Menucos depocentre, which is characterized by the dominance of therapsid footprints ( Dicynodontipus ispp.). The track assemblages from the Cuyo Basin display the highest ichnodiversity, with five footprint types.     

Open-marine Hallstatt Limestones reworked in the Jurassic Zlatar Mélange (SW Serbia): a contribution to understanding the orogenic evolution of the Inner Dinarides

Varied west-transported and far-traveled Jurassic mélanges in southwestern Serbia represent a key to understand the geodynamic history and to solve paleogeographic questions and reconstructions in the Triassic–Jurassic passive and active margin arrangement of the Inner Dinarides. Of special interest are the carbonate-clastic radiolaritic mélange areas in the Zlatar Mountain below the Dinaridic Ophiolite nappe. The present study reports from a Middle Jurassic sedimentary mélange in the area of Vodena Poljana. Carbonate components and blocks of the mass-flow deposits consist exclusively of a reworked Middle/Late Anisian to Early Jurassic distal shelf succession. Ophiolite components from the Dinaridic Ophiolite nappe stack are missing in the spectrum. The underlying series of the Zlatar Mélange belong to Early/Middle Anisian shallow-water carbonates and to Late Anisian to Middle Jurassic deep-water sedimentary rocks of the Hallstatt facies zone. South of Vodena Poljana in the overlying ophiolitic mélange occur Late Triassic radiolaritic components from the sedimentary cover of the Late Triassic ocean floor, beside ophiolite clasts and limestone components from the continental slope. A comparison with preserved Hallstatt Limestone successions and Jurassic mélange complexes from the Eastern Alps, Western Carpathians, and Albanides strengthen the interpretation of a provenance of the Zlatar mélange from the distal passive margin facing the Neotethys Ocean to the east. An autochthonous Dinaridic Ocean west of the Drina-Ivanjica Unit cannot be confirmed.     

西藏南部聂拉木地区晚泥盆世—早石炭世的介形类

本文系统描述西藏南部聂拉木地区晚泥盆世–早石炭世介形类共计15属24种。根据介形类的组合面貌及孢子、牙形刺的研究结果推断,亚里剖面上的亚里组时代为晚泥盆世–早石炭世,泥盆系-石炭系界线(D-CB)大致位于亚里东沟段顶部至沟陇日段。亚里组产出的介形类生态组合属于光滑速足目集群或艾菲尔生态型,指示亚里组在沉积时为远岸浅水沉积环境,且在上部腊扎蒲段水深变浅。亚里组介形类面貌与华南泥盆纪介形类相似程度高,与俄罗斯和欧美地区也有相似的属种,说明泥盆纪-石炭纪之交介形类的地理隔离并不显著,在各个板块之间有属种交流的可能。     

Middle and Upper Triassic radiolarite components from the Kcira-Dushi-Komani ophiolitic mélange and their provenance (Mirdita Zone,Albania)

The Middle Jurassic Kcira-Dushi-Komani Mélange in the northwestern Mirdita Ophiolite Zone contains blocks of Triassic oceanic crust, radiolarites and open marine limestones. We describe the microfacies and present biostratigraphic data from radiolarite clasts and blocks and from basalt-radiolarite blocks with preserved autochthonous sedimentary cover. The radiolarians yield Middle to Late Triassic (Late Anisian to Late Norian) ages for the oceanic realm from which the ophiolite and radiolarite blocks and clasts derived. Together with limestone blocks, the provenance areas of the components in the Kcira-Dushi-Komani Mélange are determined as the distal shelf and the ocean floor of the Neotethys. In the course of ophiolite obduction, the components were transported by mass movements into newly formed trench-like basins in front of the westward propagating nappe stack. These basins were later incorporated in the nappe stack forming the typical features of a syntectonic mélange. Our radiolarian biostratigraphic data confirm Late Anisian formation of the Neotethys Ocean, parts of which became closed in the Middle Jurassic. The data clearly speak in favour of one Neotethys Ocean to the east, from which the Mirdita ophiolites derived as far-travelled nappes.     

A sequence stratigraphic approach to a Middle Triassic shelf-slope complex of the Ligurian Alps (Ligurian Briançonnais,Monte Carmo-Rialto unit,Italy)

The Mesozoic sedimentary cover belonging to the Monte Carmo-Rialto unit of the Ligurian Briançonnais domain is composed of Scythian clastics and Anisian to Carnian carbonate rocks over 300 m thick. This paper focuses on the stratigraphy of this carbonate complex, its environmental significance, and its evolution in light of dynamic stratigraphy. Our facies analysis of limestones and dolomites of the Triassic complex allowed us to reconstruct an environmental model. Data support a distally steepened carbonate ramp of Anisian age evolving to a more diversified Ladinian platform with an oolitic sand-bar belt separating the lagoon from the slope. The Monte Carmo-Rialto slope facies are the only witnesses of deep sedimentation in the Triassic terrains of the Ligurian Briançonnais domain, otherwise represented by shallow-water carbonate deposits. On the basis of facies succession, we have identified nine medium-scale cycles (third-order sequences) in the study area, comparable to those evidenced in the Briançonnais s.s. domain by the French authors. Small-scale cycles analysis evidenced mainly shallowing-upward trends in the examined sequences; although a few evidences of transgression-related deposits (deepening upward cycles) have been found at the base three sequences, they have been mostly obliterated by dolomitization and masked by local tectonics. For this reason, we can undoubtedly distinguish only the part of each sequence belonging to HST, while the TST, though present, still remains a partition that cannot be precisely characterized. In the same way, LSTs are not present in the Monte Carmo-Rialto unit, due to the original relative landward position of the examined area. Sequence stratigraphy analysis indicates different long-term dynamics for the two evolutionary stages of the Triassic Ligurian platform: a general landward backstepping to moderate progradation during the Early Anisian and true progradation during the latest Anisian and Ladinian. In addition, a good fit with the sequences proposed by the SEPM chart has been found, indicating a correspondence for the third-order sequences of the Middle Triassic.     

华北铜川组爬行动物化石的首次发现及其地层意义

１９９２年,中国科学院古脊椎动物与古人类研究所从人民来信中得知,在山西永和县桑壁镇附近的河沟中发现了陆生四足动物化石。不久古脊椎所的专业人员就到化石产地进行了实地调查,并发掘出了几具保存较好的爬行动物化石。化石保存在坚硬的细砂岩中,已经修理出其中一个头骨（如图１所示）。经初步鉴定,这批化石是一类初龙形类动物,与产自新疆吐鲁番盆地中三叠世克拉玛依组的达板吐鲁番鳄（Ｔｕｒｆａｎｏｓｕｃｈｕｓｄａｂａｎｅｎｓｉｓ）（Ｙｏｕｎｓ,１９７３）关系较近,但是其内颈动脉孔位于基蝶骨侧面,而不是位于腹面,这表明…     

The Permian–Triassic transition and the onset of Mesozoic sedimentation at the northwestern peri-Tethyan domain scale: Palaeogeographic maps and geodynamic implications

The main aim of this paper is to review Middle Permian through Middle Triassic continental successions in European. Secondly, areas of Middle–Late Permian sedimentation, the Permian–Triassic Boundary (PTB) and the onset of Triassic sedimentation at the scale of the westernmost peri-Tethyan domain are defined in order to construct palaeogeographic maps of the area and to discuss the impact of tectonics, climate and sediment supply on the preservation of continental sediment.At the scale of the western European peri-Tethyan basins, the Upper Permian is characterised by a general progradational pattern from playa-lake or floodplain to fluvial environments. In the northern Variscan Belt domain, areas of sedimentation were either isolated or connected to the large basin, which was occupied by the Zechstein Sea. In the southern Variscan Belt, during the Late Permian, either isolated endoreic basins occurred, with palaeocurrent directions indicating local sources, or basins underwent erosion and/or there was no deposition. These basins were not connected with the Tethys Ocean, which could be explained by a high border formed by Corsica–Sardinia palaeorelief and even parts of the Kabilia microplate. The palaeoflora and sedimentary environments suggest warm and semi-arid climatic conditions.At the scale of the whole study area, an unconformity (more or less angular) is observed almost everywhere between deposits of the Upper Permian and Triassic, except in the central part of the Germanic Basin. The sedimentation gap is more developed in the southern area where, in some basins, Upper Permian sediment does not occur. The large sedimentary supply, erosion and/or lack of deposition during the Late Permian, as well as the variable palaeocurrent direction pattern between the Middle–Late Permian and the Early Triassic indicate a period of relief rejuvenation during the Late Permian. During the Induan, all the intra-belt basins were under erosion and sediment was only preserved in the extra-belt domains (the northern and extreme southern domains). In the northern domain (the central part of the Germanic Basin), sediment was preserved under the same climatic conditions as during the latest Permian, whereas in the extreme southern domain, it was probably preserved in the Tethys Ocean, implying a large amount of detrital components entering the marine waters. Mesozoic sedimentation began in the early Olenekian; the ephemeral fluvial systems indicate arid climatic conditions during this period. Three distinct areas of sedimentation occur: a northern and southern domain, separated by an intra-belt domain. The latter accumulated sediments during the Early–Middle Permian and experienced erosion and/or no-deposition conditions between the Middle–Late Permian and the beginning of Mesozoic sedimentation, dated as Anisian to Hettangian. At the top of the Lower Triassic, another tectonically induced, more or less angular unconformity is observed: the Hardegsen unconformity, which is dated as intra-Spathian and is especially found in the North European basins. This tectonic activity created new source areas and a new fluvial style, with marine influences at the distal part of the systems. During the Anisian and Ladinian, continental sedimentation was characterised by a retrogradational trend. In other words, the fluvial system evolved into fluvio-marine environments, attesting to a direct influence of the Tethys Ocean in the southern and northern domains. Both at the end of the Olenekian (Spathian) and during the Anisian, the presence of palaeosols, micro- and macrofloras indicate less arid conditions throughout this domain.     

History of a Late Carboniferous phylloid algal bank complex in northeastern New Mexico

The growth history of a phylloid algal biohermal complex of Late Carboniferous (Westphalian) age, outcropping within the La Pasada Formation in northeastern New Mexico, U.S.A., is described in relationship to paleogeography, biostratigraphy, and microfacies associations. This phylloid algal biohermal complex occurs within typical Late Paleozoic cyclical sediments, in a paleogeographic setting along a narrow shelf margin bordering a deep geosynclinal trough to the northwest. Fusulinid foraminifers suggest that the bioherm is of Middle Pennsylvanian (lower Des Moines) age, whereas the overlying sediments are of middle Des Moines age. Both field and petrographic evidence suggest that the bioherm grew upward from relatively shallow water depths, into extremely shallow water where it probably was subaerially exposed. Biohermal growth appears to have been initiated on a 'hard-ground' surface on which lithified clasts and abundant clusters of robust linoproductid brachiopods served as a foundation. Four major microfacies have been identified from the biohermal complex. These are: (I) phylloid algal wackestones-packstones (mound proper), (2) marly limestones (sediments laterally adjacent to and overlying mound), (3) pelletal-foraminiferal wackestone (partially capping the mound), and (4) crinoidal packstones-grainstones (post-mound). Comparison is made with other surface and subsurface occurrences of phylloid algal complexes of similar age in the southwestern United States.     

Facies and biota of Anisian to Carnian carbonate platforms in the Northern Calcareous Alps (Tyrol and Bavaria)

Summary During the Middle and early Late Triassic carbonate ramps and rimmed platforms developed at the northwestern margin of the Tethys ocean. In the Northern Calcareous Alps, Anisian stacked homoclinal ramps evolved through a transitional stage with distally steepened ramps to huge rimmed platforms of Late Ladinian to Early Carnian age. Middle Triassic to early Late Triassic facies and biota of basin, slope and platform depositional systems are described. Special emphasis is given to foraminifers, sponges, microproblematic organisms and algae. The Ladinian to early Carnian reef associations are characterized by the abundance of segmented sponges, microproblematica, biogenic crusts and synsedimentary cements. Among the foraminifers, recifal forms likeHydrania dulloi andCucurbita infundibuliformis (Carnian in age) are reported from the Northern Calcareous Alps for the first time. Some sphinctozoid sponges likeParavesicocaulis concentricus were known until now only from the Hungarian and Russian Triassic.     

The first tunicate with a calcareous exoskeleton (Upper Triassic,northern Italy)

The first tunicates with a calcareous exoskeleton are reported from Late Triassic buildup‐slope deposits of the Dolomites. Although examples of this group have been known since the early 1900s from the middle–upper Permian of eastern Asia and Sicily as Khmeria, they were erroneously attributed to rugose corals. These early representatives are small, double‐valved, conical skeletons, which evolved into multi‐plated capsules with up to 35 opercula. The latter are joined along zigzag margins, which in life could probably be opened for the atrial and branchial siphons. The construction and shape of these skeletons distinguish them from plants or other invertebrate phyla, while they share several similarities with living tunicates, specifically to sessile ascidians. Apart from a soft‐bodied genus from the lower Cambrian of China, ascidians are known only from isolated spicules, which occur sporadically from the Lower Jurassic onwards. The calcareous skeleton of these Late Triassic tunicates consists of aragonitic fibres, which form spherulitic or clinogonal microstructures. It seems that the stellate aragonitic spicules of Jurassic to Recent ascidians are a vestige of Permian–Triassic ancestors, which after the Carnian lost the ability to construct compound solid skeletons but partly still retain a soft double‐valved or multi‐operculate cellulose‐like tunic. The following taxa are described as new: Order Khmeriamorpha with the genera Khmeria Mansuy and Zardinisoma gen. nov., and the following species: Khmeria stolonifera (late Permian), Khmeria minima (Late Triassic), Zardinisoma japonicum (late Permian), Z. cassianum, Z. pyriforme, Z. polyplacophorum and Z. pauciplacophorum (all Late Triassic).