Type specimens and identity of the described species of Conus.

During the third decade of the 19th century, 21 authors described 35 Recent and 11 fossil species of Conus , in 26 separate publications. This brought to 532 the number of species-group names in the genus introduced up to 1830. Only five fossil and seven Recent species are now considered valid. Two additional valid Recent species were described but were given preoccupied names. The remaining nominal species are concluded to be junior or contemporaneous synonyms (19), nomina dubia (11), of infrasubspecific rank (1), or not now considered Conus (1). At present 160 valid species of Conus are recognized from the species-group names described between 1758 and 1830. Of these, 138 are extant and 22 are known only as Tertiary fossils. Nine otherwise valid species, six Recent and three fossil, were described but were given preoccupied names.     

Chronological analysis of the species of Conus described during the 18th century

The 339 species-group names in the gastropod genus Conus proposed between 1758 and 1800 have been analyzed in chronological order, in order to (1) identify each nominal species, and (2) determine the correct name of each valid species. Additions and corrections are given to the author's series of papers on type specimens and identity of the described species of Conus. A table listing the 339 species-group names alphabetically indicates the present locations of the 124 known type specimens and the status of each name. At present, 123 valid species are recognized (118 Recent, 5 Tertiary), 159 names are synonyms of previously or simultaneously described species, 35 are nomina dubia, 19 are assigned to infrasubspecific rank, and 3 have been trandferred to genera in other families. Problems involving homonymy of 44 names have been resolved. The proportion of valid species to described species remained quite constant (about 80%) between 1758 and 1786, but then declined sharply to 40%. By 1800, although only 13% of all known species-group names in Conus had been introduced, they probably account for 25% of the known valid Recent species. Extension of the chronological study to species described through 1840 may establish the correct names of half the valid Recent species of Conus.     

Type Specimens and identity of the described species of Conus II. The species described by Solander, Chemnitz, Born, and Lightfoot between 1766 and 1786

Twenty-three species of Conus were described by authors other than Linnaeus between 1766 and 1786. Of these, 3 are Eocene species from England described by Solander in Brander (1766), and the remainder are Recent. Chemnitz (1777) described one valid species. Of the 10 species described by Born (1778), 7 are valid, 2 are junior synonyms of Linnaean species, and 1 is a nomen dubium. Solander in [Lightfoot] (1786) described 5 species, of which 3 are valid and 2 are junior synonyms of Linnaean species. Lightfoot (1786) described 4 species, of which 1 is valid and 3 are junior synonyms of Linnaean species.
Holotypes of 7 species and lectotypes of 3 species exist, and representatives of lecto-types of 10 species have been selected. All of these types are illustrated photographically.
Between 1758 and 1786, 60 species of Conus were described, of which 45 are now considered valid. Of these, 42 are Recent.     

The species of Hymenoptera described by Linnaeus in the genera Sphex, Chrysis, Vespa, Apis and Mutilla

Data are presented on the type-material representing the species described by Linnaeus in his genera Sphex, Chrysis, Vespa, Apis and Mutilla , The names here considered total 158; six are currently applied in the Hymenoptera Parasitica. Of the balance of 147, three are emendations and five are unavailable homonyms or names proposed in the synonymy of other species. Fifteen Linnaean names are here placed as synonyms. One name is attributed to an author other than Linnaeus, and live names remain species incertae sedis. The remainder, 118, are applied as valid names in die Hymenoptera Aculeata. Holotype specimens in London, Uppsala or Stockholm represent 60 names; 55 names are based on lectotype specimens of which 49 are here designated, two by other authors. One name is represented by a neotype specimen in Lund, and one by a lectotype figure. One non-aculeate species is based on syntypes. Specimens appear to be lost with respect to 25 names; three specimens in Uppsala may be the holotypes of a further three species.
Eight new combinations are made, and twelve new synonyms established; three further new combinations and three further new synonymies are suggested. A systematically arranged summary of species treated and of nomenclatural changes made is given. Nomenclatural changes affecting non-Linnaean names are included where relevant; lectotypes are designated for two non-Linnaean species.     

The Microlepidoptera described by Linnaeus and Clerck

Abstract The identities of the Microlepidoptera described by Linnaeus and Clerck have been critically reviewed. Evidence used to reassess usage of Linnaeus and Clerck names comes in part from the surviving material of these two authors, from their publications and those of subsequent revisers, and from contemporary collections. Results are presented in the form of an alphabetic catalogue of the species-group names used by Linnaeus and Clerck for Microlepidoptera, with a complete listing of material in the Linnaeus and Clerck collections. Lectotypes are designated for most of the species and new synonymies are proposed.     

A taxonomic revision of the genera Phrygionis Hübner and Pityeja Walker (Geometridae: Ennominae, Palyadini)

Phrygionis and Pityeja belong to the Palyadini, a tribe of neotropical Ennominae. The moths of both genera bear striking wing markings and mere variants were frequently described as species with the consequence that species diversity was overestimated. Numerous taxonomic changes are made in this work based on the study of primary types and much other material. Variation and distribution is recorded for each species and subspecies. Phylogenetic relationships within Phrygionis are examined, and comments are made on the evolution of wing pattern within this genus.
Fifteen full species are recognized in this study (13 in Phrygionis and two in Pityeja ), of which four (all in Phrygionis ) are described as new; 19 species-group (specific and subspecific) names refer to valid taxa. Twenty-four species-group names are synonymized (19 in Phrygionis and five in Pityeja ) and five are recombined with different genera (four with Phrygionis and one with Pityeja ). Three generic names are synonymized (one in Phrygionis and two in Pityeja ).     

Typification of the Linnean species of Saxifraga

WEBB, D. A., 1987. Typification of the Linnean species of Saxifraga. Types (mainly lectotypes) arc assigned to all the species of Saxifraga described by Linnaeus. There are 38 names in all, of which one is now assigned to Bergenia, one is known to be a hybrid, three have been rejected in the past as nomina ambigua, one was later reduced to synonymy by Linnaeus himself, while 32 are accepted as valid species today. Three have been previously typified by Pugsley, one by Steam and one by the author; for the others the types are newly proposed here. Nineteen names are typified from the Linnean herbarium in London (LINN), three from the Linnean herbarium in Stockholm (S), seven from the Burser herbarium at Uppsala (UPS), one from the Tournefort herbarium in Paris (P), and eight from illustrations cited by Linnaeus in synonymy. Lists are appended of all specimens of the genus in the Linnean herbaria in London and Stockholm and the Burser herbarium at Uppsala, and a determination is given for each specimen, though in some cases these can only be tentative. The specimens include 15 species which Linnaeus did not describe, though in the case of five he probably regarded them as varieties.     

First reliable record of a fossil species of Anthomyiidae (Diptera), with comments on the definition of recent and fossil clades in phylogenetic classification

All previous records of fossil Anthomyiidae are shown to be unsubstantiated. A female anthomyiid of a new genus and species is hereby described from a piece of Dominican amber (Upper Eocene-Oligocene). Character analysis suggests that the fossil, Coenosopsites poinari gen. & sp. nov. , belongs to a Neotropical clade with two recent genera, Phaonantho Albuquerque and Coenosopsia Malloch. Evidence for a sister-group relationship between Coenosopsites poinari and the genus Coenosopsia is provided. Clades are the only acceptable units of phylogenetic classification. Combining fossil and recent clades in phylogenetic classification requires them to be temporally delimited. Proper application of phylogenetic definitions is essential for this purpose. It is proposed that the units of phylogenetic classification should be taxa for recent clades and plesia for fossil clades. A taxon is defined as node-based with reference to its recent species, while a plesion is defined as apomorphy-based. The term lineage is proposed for a recent clade defined as stem-based with reference to its recent sister group. Individual recent species represent clades that can be incorporated into phylogenetic classification as minimal taxon units. Individual fossil species may not represent clades and thus do not count as proper units of phylogenetic classification. However, the names of fossil species are readily construed also to signify plesia with the fossil species as their only known component. As such, they are proper units of phylogenetic classification.     

Species-level phylogeography and evolutionary history of the hyperdiverse marine gastropod genus Conus

Phylogenetic and paleontological analyses are combined to reveal patterns of species origination and divergence and to define the significance of potential and actual barriers to dispersal in Conus, a species-rich genus of predatory gastropods distributed throughout the world's tropical oceans. Species-level phylogenetic hypotheses are based on nucleotide sequences from the nuclear calmodulin and mitochondrial 16S rRNA genes of 138 Conus species from the Indo-Pacific, eastern Pacific, and Atlantic Ocean regions. Results indicate that extant species descend from two major lineages that diverged at least 33 mya. Their geographic distributions suggest that one clade originated in the Indo-Pacific and the other in the eastern Pacific + western Atlantic. Impediments to dispersal between the western Atlantic and Indian Oceans and the central and eastern Pacific Ocean may have promoted this early separation of Indo-Pacific and eastern Pacific + western Atlantic lineages of Conus. However, because both clades contain both Indo-Pacific and eastern Pacific + western Atlantic species, migrations must have occurred between these regions; at least four migration events took place between regions at different times. In at least three cases, incursions between regions appear to have crossed the East Pacific Barrier. The paleontological record illustrates that distinct sets of Conus species inhabited the Indo-Pacific, eastern Pacific + western Atlantic, and eastern Atlantic + former Tethys Realm in the Tertiary, as is the case today. The ranges of <1% of fossil species (N=841) spanned more than one of these regions throughout the evolutionary history of this group.     

Catalog of the adelgids of the world (Hemiptera,Adelgidae)

A taxonomic and nomenclatural Catalogue of the adelgids (Hemiptera: Adelgidae) is presented. Six family-group names are listed, five being synonyms of Adelgidae. Twenty-two genus-group names, of which nine are subjectively valid and in use, are presented with their type species, etymology, and grammatical gender. One hundred and six species-group names are listed, of which 70 are considered subjectively valid.     

Discovery in the Linnaean collection of type-material of insects described by Johann Reinhold Forster, with notes on the Hymenoptera

Type-material of 100 insect species described by J. R. Forster has always been thought lost. We present evidence that syntypic specimens of some of these species were sent by Forster to Linnaeus in 1772. Specimens of at least 27 species survive in the Linnaean collection now in Burlington House, and are listed. The identity of each of the 18 species of Hymenoptera described by Forster is examined in detail, particularly the nine represented by specimens. Lectotypes are designated for two species; four new synonymies and two new combinations are established; and three names remain nomina dubia within their genera.     

Type specimens of Mitridae (Mollusca:Gastropoda) described by Linnaeus in the genera Voluta, Buccinum and Bulla between 1758 and 1771

From the 16 species of Mitridae described by Linnaeus between the years 1758-71 11 species are considered valid and available for taxonomic use; four specific names have been suppressed by the International Commission on Zoological Nomenclature for taxonomic usage (1968). One specific name, i.e. Voluta mitra episcopalis Linnaeus, has appeared in various combinations in malacological literature over the years. From the style of listing in the 10th edition of the Systerna Naturae , it is evident that 'episcopalis' has to be treated as of infraspecific rank; there is no evidence that 'episcopalis' is even a valid subspecies of Voluta mitra.
Three holotypes and five lectotypes designated in this paper are extant in the Linnaean collection; illustrations representing three Linnaean species have been selected as lectotypes.     

Two ants (Insecta: Hymenoptera: Formicidae: Formicinae) from the Late Pliocene of Willershausen, Germany, with a nomenclatural note on the genus Camponotites

Two species of the genus Camponotites (Formicidae, Formicinae) are described from the Late Pliocene deposits of Willershausen, Lower Saxony, northern Germany: C. silvestris Steinbach, 1967, and C. steinbachi n. sp. The generic name Camponotites has been established for fossil (Tertiary) ants independently by Steinbach (Bericht der Naturhistorischen Gesellschaft zu Hannover 111:95–102, 1967) and by Dlussky (Trudy paleontologi?eskogo instituta, akademiâ nauk SSSR, 1981), each for materials of different stratigraphical and geographical origin. Though poorly described, Camponotites Steinbach, 1967, and the single included (type) species C. silvestris Steinbach, 1967 (a monotypic species from the Late Pliocene of Willershausen), were based upon indication in the sense of the ICZN. Therefore, both the generic and specific names are valid and available. Camponotites Dlussky, 1981 (and its type species C. macropterus Dlussky, 1981) were certainly introduced correctly and are therefore available, too; but due to its homonymy the generic name is not valid. The revision shows that in this rare case both generic names are not only homonyms but also synonyms.     

New species of Crotonia (Acari: Oribatida) from South Africa

Four new species of oribatid mites of the genus Crotonia , ( C. pauropelor sp. nov., C. dicella sp. nov., C. lyrata sp. nov. and C. ecphylla sp. nov. ) are described from soil and plant litter from South Africa. Crotonia pauropelor, C. dicella and C. ecphylla are allied to the 'capistrata' species-group, previously known only from Queensland, Australia, in that they possess the full complement of c setae on the notogaster. They differ in having eight pairs rather than nine pairs of genital setae. In addition, C. ecpylla is unusual in the genus Crotonia in that it appears to possess two pairs of setae on each of the second epimera. Crotonia lyrata has only two pairs of c setae and is allied to the 'cophinaria' species-group. This is only the second published record of the genus Crotonia from South Africa, and the first of any named species. A key to the African Crotonia species is provided, which includes the only two previously known species C. alluaudi and C. rothschildi (Berlese, 1916).     

A review of the moth tribe Palyadini with the description of a new genus (Geometridae: Ennominae)

The tribe Palyadini Guenée is revised at the generic level. It includes about 115 described species, and occurs in tropical America with representatives from Florida to Argentina. There are six genera: Palyas Guenée, Phrygionis Hübner, Pityeja Herrich-Schäffer, Argyrotome Warren, Opisthoxia Hübner, and a new genus Ophthalmoblysis. Ophthalmophora Guenée and Argyroplutodes Warren are treated as junior synonyms of Opisthoxia. Checklists of the species-group names in each genus are included.