Early evidence of Prunus and Prunus cf. amygdalus from Palaeolithic sites in the Khorramabad Valley,western Iran

Along with the early age obtained for the cultural remains attributed to anatomically modern humans from Kaldar Cave, the archaeological assemblages recovered from both Kaldar and Gilvaran Cave located in the Khorramabad Valley (Iran), have yielded charcoal remains that allow the identification of Prunus spp. These remains correspond to the Middle and Upper Palaeolithic, which are the earliest finds attesting to the presence of this taxa in the area. Our anatomical observation of the samples revealed the presence of Prunus spp. (plums) and Prunus cf. amygdalus (cf. almond). This also reflects specific plant communities in the area, characteristic of open forest growing in cool, dry conditions. These results provide new insights into the arboreal cover in this area during an Upper Pleistocene period. Furthermore, anthracological evidence together with other contextual materials provides new clues to assess how Neanderthals and early modern humans adapted to their surrounding landscape, and their relationship with their environment in this region and beyond.     

Alveolina-dominated assemblages in the early Eocene carbonates of Jaintia Hills,NE India: Biostratigraphic and palaeoenvironmental implications

Larger benthic foraminifera (LBF) are significant proxies in biostratigraphy and also act as excellent indicators of shallow-marine carbonate environments in fossil series. The Palaeogene LBF recorded from Meghalaya, NE India (eastern part of the relic eastern Tethys/Neo-Tethys) have high potential for dating shallow-marine sediments and documenting the multiple episodes of carbonate sedimentation that have contributed to the development of the Sylhet Limestone Group. Early Eocene witnessed the proliferation of LBF species worldwide, the phenomenon better known as the Larger Foraminiferal Turnover (LFT). Genera like Alveolina, Nummulites and Orbitolites with broad species complexes thrived as the dominant LBF amidst numerous other taxa on the verge of extinction or only surviving as stable forms. The current study emphasizes on the biostratigraphic and palaeoenvironmental account of the early Eocene Umlatdoh Limestone successions outcropping in the Jaintia Hills of Meghalaya, primarily based on the recorded species of Alveolina and other larger benthic foraminifera. Five species of Alveolina — A. oblonga, A. schwageri, A. cf. ruetimeyeri, A. aff. haymanensis and A. aff. varians are recorded in the evaluated sections that indicate an early Eocene age corresponding to the Shallow Benthic Zone 10. Major carbonate facies types in the present assessment include oolitic-smaller benthic foraminiferal -green algal grainstone–packstone, smaller miliolid-Alveolina grainstone, green algal-benthic foraminiferal grainstone, larger porcellaneous (Alveolina) grainstone-packstone, Alveolina-nummulitid grainstone-rudstone, and nummulitid grainstone-rudstone, which indicate a shallow marine, high-energy depositional environment ranging from shoal-sandy bars to a distal inner ramp setting.     

A new hawker dragonfly from the Middle Jurassic of China (Odonata: Aeshnoptera)

A new genus and species Linqibinia panae of paracymatophlebiid hawker dragonfly is described from the Middle Jurassic Haifanggou Formation (Inner Mongolia, China). Previously only known from Karatau in Kazakhstan, the discovery of another member of this family extends its range across Central Asia. It confirms that the Aeshnoptera was among the most diverse odonatan clades during the Middle-Late Jurassic.     

The Late Pleistocene faunal assemblage from Cava Muracci (Latium,Italy): Palaeoenvironmental implications for coastal central Italy during MIS 3

Palaeoenvironmental information on Marine Isotope Stage 3 (MIS 3) coastal Latium is sparse, mainly based on studies of isolated faunal assemblages or long pollen records from lake sediments, often of insufficient resolution to aid in palaeoenvironmental reconstruction. This study describes in detail the Late Pleistocene faunal assemblage from layers SU11 and SU12 of Cava Muracci (Cisterna di Latina, central Italy), the first of which is a partially-preserved hyena den. The first multi-disciplinary palaeoenvironmental reconstruction of coastal Latium between 34–44 ka BP, a critical time span for the presence of the latest Neanderthals and the arrival of Anatomically Modern Humans (AMH), is provided combining palaeoecological inferences from a previous pollen study of hyena coprolites with the palaeontological study described here. The results indicate a temperate climate and a landscape characterised by the coexistence of at least three habitats within a short distance between the coastline and the inland mountains, suitable for a wide variety of species.     

Sabellid and serpulid worm tubes (Polychaeta,Canalipalpata, Sabellida) from the historical stratotype of the Cenomanian (Late Cretaceous; Le Mans region,Sarthe, France)

The rich Cenomanian assemblage of tube-dwelling polychaete worms of the families Sabellidae and Serpulidae from Le Mans region (Sarthe) is described in detail for the first time in one place; the systematics and the palaeoecology of the species are discussed. The 19 species described belong to 12 genera: Glomerula Brünnich Nielsen, 1931; Filograna Berkeley, 1835; Protula Risso, 1826; Filogranula Langerhans, 1884; Serpula Linnaeus, 1758; Cementula Brünnich Nielsen, 1931; Neovermilia Day, 1961; Dorsoserpula Parsch, 1956; Mucroserpula Regenhardt, 1961; Placostegus Philippi, 1844; Cycloplacostegus Jäger, 2005; Pyrgopolon de Montfort, 1808. Two new species are described: Serpula? pseudoserpentina sp. nov. and Pyrgopolon (Septenaria) cenomanensis sp. nov.     

Toarcian and Aalenian (Jurassic) ammonites from the Shemshak Formation of the Jajarm area (eastern Alborz, Iran)

In the Jajarm area (eastern Alborz Mountains, NE Iran) the ?Upper Triassic — Lower Bajocian siliciclastic Shemshak Formation is up to 2000 m thick. Whereas the lower third of the formation is nearly exclusively non-marine, the upper two-thirds are fully marine. The middle part is characterized by several levels containing a relatively diverse and well preserved assemblage of ammonites of the Toarcian to Aalenian stages. Two sections of the ammonite-bearing strata, spaced 20 km, are presented and correlated by means of ammonite biostratigraphy. The ammonite fauna consists of 27 taxa, some of which are recorded for the first time from the Alborz Mountains. The ammonites are briefly described and their palaeobiogeographic context is reviewed. The ammonite fauna of the Shemshak Formation at Jajarm, as elsewhere in North and Central Iran, is exclusively Northwestern Tethyan in character and is closely related to the faunas of Northwestern and Central Europe.     

The Tournaisian (Early Carboniferous) of the Kahanag section (central Alborz; northern Iran); biostratigraphy with calcareous algae and foraminifers; palaeobiogeographic implications

The Tournaisian beds of the Kahanag section in central Alborz (northern Iran) are described. This Tournaisian is probably complete and continuous from MFZ1 to MFZ8 biozones; nevertheless, (1) the base is only lithostratigraphically characterized, (2) the biozones MFZ1 to MFZ5 cannot be differentiated, and (3) in topmost beds, Eoparastaffella simplex indicative of the basal Viséan have not yet found. This Tournaisian is subdivided into a local lower Tourmaisian (MFZ1-MFZ5) showing a poor assemblage of monolocular and bilocular foraminifers, and a local upper Tournaisian where the biozones MFZ 6 and 7 are precisely defined. In Kahanag, MFZ 6 is characterized by the FO of Eotextularia diversa; MFZ 7 is the range-zone of Darjella monilis; and the base of MFZ8 is marked by the appearance of Eoparastaffellina. The biozone MFZ 8 is particularly developed in Kahanag, and represents almost one third of the field-section; therefore, a local subdivision is proposed, especially in order to characterize an upper MFZ8 subbiozone. Additionally, some algae of the section are accurately described; especially a trio which might be characteristic of the latest Tournaisian in Iran, (1) the unilayered “Koninckopora” mentioned for the first time in Iran; (2) a new genus of dasycladaleans Kahanagella n. gen.; and (3) a new genus of palaeoberesellaceans, Oblikamaena n. gen. More classical Algospongia (incertae sedis algae) are briefly mentioned Issinella, Crassikamaena, Kamaena, Palaeoberesella, Exvotarisella, Pseudostacheoides, Sinustacheoides and Stacheoides. Regionally, the new genus Kahanagella might be an important palaeobiogeographic marker because it was also discovered in the Tabas Block, while it seems absent in the adjacent, well-known areas (Taurus, southern Urals, Kazakhstan). Perhaps, Kahanagella might permit to reconstruct the initial geography of the latest Tournaisian Iranian platforms. Oblikamaena elbursica n. gen. n. sp. is an endemic taxon for the moment but might become also palaeobiogeographically interesting. Finally, the whole foraminiferal and algal Tournaisian assemblage permits to propose a reconstruction of the initial palaeoposition of the Iranian terranes prior to the progressive assemby of the Cimmerian terranes.     

Test excavations and initial results at the Middle and Upper Paleolithic sites of Gilvaran,Kaldar, Ghamari caves and Gar Arjene Rockshelter,Khorramabad Valley,western Iran

This paper introduces the excavations in several Paleolithic sites in the Khorramabad Valley, Western Iran. Apart from the two well-known sites of Ghamari Cave and Gar Arjene rock shelter, first excavated by Frank Hole and Kent Flannery in the 1960s, the Gilvaran and Kaldar caves were excavated for the first time. Here we present the stratigraphy of these sites, general data from the lithic assemblages, and the identifications of a small part of the faunal remains. Preliminary results are showing that all of the sites were occupied from the Middle and Upper Paleolithic onward, and therefore provide great potential for the study of the transition between these cultural periods. Our preliminary techno-typological observations show that the lower levels of the Gilvaran and Ghamari sequences may represent an early phase of the Middle Paleolithic.     

Microfacies,microtaphonomic traits and foraminiferal assemblages from Upper Jurassic oolitic–coral limestones: stratigraphic fluctuations in a shallowing-upward sequence (French Jura,Middle Oxfordian)

The characterization and distribution of the microfacies and the microfossil assemblages of a Middle Oxfordian section from Jura Mountains composed by thick oolitic–coral limestones is analyzed. Six microfacies types (mainly grainstones) are differentiated mainly composed by ooids, intraclasts and bioclasts. Foraminiferal assemblages are dominated by agglutinated forms. Benthic microbial communities and sessile foraminifera are the main components of the encrustations. The whole set of microfossil assemblages is typical of shallow subtidal environments rich in “algae” (Cayeuxia, “Solenopora”, Thaumatoporella, Bacinella, Girvanella and Terquemella) and foraminifera such as Nautiloculina oolithica, Redmondoides lugeoni, Ammobaculites coprolitiformis, Troglotella incrustans and Rectocyclammina. The increasing upward record of debris of algae and Nautiloculina, and the decrease of serpulids, bryozoans, nodosariids and ophthalmidiids indicate a shallowing-upward trend. The stratigraphic distribution of microfacies and microfossil assemblages lead to differentiate two main successive phases. The first is a deeper subtidal environment in an open shelf, while the second is a shallow subtidal environment with evolution from winnowed to more restricted conditions. Microfabrics of radial to concentric ooids upwards in the section correspond to higher energy environments related to an oolitic shoal. This study shows how a very detailed analysis of microfacies, which integrates oolitic features, microfossil assemblages and microtaphonomy is potentially a useful tool for interpreting hydrodynamism and sequence evolution in marine carbonate shallow environments.     

Paleoecological and microenvironmental aspects of the first European hominids inferred from the taphonomy of small mammals (Sima del Elefante,Sierra de Atapuerca,Spain)

Unit TE9 of the Sima del Elefante (Sierra de Atapuerca, Burgos, Spain), where the remains of Homo sp. have been discovered (1.2–1.3 Ma), is also a level rich in small mammals. The taphonomic study of these small vertebrates sheds light on the landscape that provided the setting for the activities of these early hominids and allows us to describe what the cave was like during the formation of the level. Small mammal predators identified in the study indicate that during this period the Sierra de Atapuerca was part of a large biome consisting of semi-open riparian forests with meadows nearby. Postdepositional alterations reveal that the production of fossils took place outside the cave, being transported inside by water currents. During the formation of TE9, the cave presented conditions of high humidity, which made it difficult for hominids to establish occupations inside, although these conditions became somewhat less severe in TE9c, the sublevel where human remains are found.     

Avian fossil assemblages at the onset of the LGM in the eastern Alps: A palaecological contribution from the Rio Secco Cave (Italy)

The avian fossil assemblages from the late Pleistocene deposits of the Rio Secco Cave (north-eastern Italy) is presented herein. We studied the layers that date back to the end of MIS3 and the beginning of MIS2, which also contain evidence of Gravettian frequentation dated to 33.5–30 ka cal BP. The systematic analysis revealed the presence of 18 species and other supraspecific taxa that supported palaeoenvironmental reconstructions. Taxa indicate that, at the onset of LGM, site surroundings were characterised by conifer or mixed forests, open grasslands, slow-flowing water bodies and mountain meadows with rocky outcrops, as indicated by Lagopus muta. Today, this environment is found above the tree line (beyond 1500–2000 m) and cannot be detected near the site, located at 580 m asl. Noteworthy, is also the finding of the second Italian late Pleistocene fossil record of Picus canus.     

Live and dead benthic foraminiferal assemblages from coastal environments of the Aegean Sea (Greece): Distribution and diversity

Benthic foraminiferal composition assemblages and their temporal changes, ecological indices and foraminiferal densities are used to compare three coastal environments with different physicogeographical features in the Aegean Sea (coastal environment of Avdira–Vistonikos Gulf and Kitros–Thermaikos Gulf and open lagoonal environment of Vravron–South Evoikos Gulf). Three main foraminiferal assemblages have been recognized: a) “Assemblage A”; high degree of similarity between living and dead foraminiferal species, dominated by Ammonia beccarii, Elphidium spp. and relatively abundant and diverse miliolids, b) “Assemblage B1”; intermediate degree of similarity between live and dead assemblages, characterized by highly-abundant and well-diversified foraminiferal assemblages including the algal symbiont bearing Peneroplis pertusus together with Ammonia tepida and several small epiphytic rotaliids and miliolids, and c) “Assemblage B2”; absence of living individuals, strongly dominated by the opportunistic species A. tepida. Our results suggest a good comparison between living and dead assemblages from different coastal environments in the Aegean Sea, however the prevailing environmental conditions (vegetation cover, hydrodynamics, fresh water influx) have a strong impact on the taphonomic processes.     

Civettictis braini nov. sp. (Mammalia: Carnivora), a new viverrid from the hominin-bearing site of Kromdraai (Gauteng,South Africa)

A new Plio-Pleistocene viverrid species is described based on two newly discovered maxillae (KW 10141 and KW 10383) from the recent excavations at the hominin-bearing site of Kromdraai (Gauteng, South Africa). This major site allows us to address the conundrum of Paranthropus and Homo origins in South Africa and presents a highly diverse carnivore spectrum (at least 22 species) including herpestids and viverrids. Civettictis braini nov. sp. is a viverrid species comparable in size to the extant African civet Civettictis civetta (Schreber, 1776). However C. braini nov. sp. differs significantly from the extant species in its dental proportions. Its canine and three premolars (P1–P3) are relatively robust, while its carnassials (P4) and two molars (M1, M2) are extremely reduced. This new species supplements our knowledge on carnivore taxonomic diversity and paleoecology in Southern Africa about 2 millions of years ago.     

Armored dinosaurs(Ornithischia: Ankylosauria)) from the Middle and Upper Jurassic of England

Hitherto the earliest positive record of ankylosaurs(armored dinosaurs) has been from beds well up in the Lower Cretaceous; in fact, however, specimens referable to the ankylosaurian family Nodosauridae are present in the Middle and Upper Jurassic of England: from the Middle Callovian [partial mandible Sarcolestes leedsiLydekker]], the Upper Oxfordian [femur Cryptodraco eumerus (Seeley)), maxilla Priodontognathus phillipsii (Seeley))], and the Upper Tithonian [caudal vertebra, tooth]. The Tithonian tooth and those of Priodontognathus are large and similar to those of the nodosaurids Priconodon and Sauropelta (Lower Cretaceous, U.S.A.). The incomplete mandible of Sarcolestes is similar to that of Sauropelta with a dermal scute fused to the lateral surface, and a tooth row extending to the anterior end of the jaw; an unusual feature is the caniniform first tooth. The quadrupedal ankylosaurs and stegosaurs probably represent separate evolutionary lines that extend back at least into the Lower Jurassic, and both lines probably evolved from ornithopod dinosaurs that were bipedal. Nodosaurid ankylosaurs occur in Europe from the Middle Jurassic to Late Cretaceous and probably reached North America via a filter route in the early Cretaceous.     

Hrabalicrinus zitti gen. et sp. nov., and other Upper Jurassic crinoids (Echinodermata,Crinoidea) from the Brno area (Czech Republic)

Jurassic (Oxfordian) crinoids from the Brno area (Czech Republic) are described in details for the first time. A rich crinoid assemblage consisting of cups, isolated cup elements, brachial plates, columnals, pluricolumnals, and cirrals is assigned to isocrinids Isocrinus amblyscalaris (Thurmann), Balanocrinus subteres (Münster), B. pentagonalis (Goldfuss), Isocrinida indet., comatulid Hrabalicrinus zitti gen. et sp. nov., cyrtocrinids Lonchocrinus sp., Cyrtocrinus cf. nutans (Goldfuss), Pilocrinus moussoni (Desor), Tetracrinus moniliformis Münster, Cyrtocrinida indet., and millericrinids (Millericrinida indet.). Crinoids already mentioned from the Jurassic strata of the Czech Republic are subsequently revised. Apart from the isolated remains of Isocrinida, Millericrinida, and Thiolliericrinida, the presence of any other taxon reported from this area should be treated with extreme caution. A rare example of non-regenerative columnal healing (the so-called callus) in I. amblyscalaris is also described. Based on sedimentology and microfacies, the Jurassic limestones were deposited in various palaeoenvironments of upper carbonate platform and shelf lagoon (0 to > 50 m palaeodepth).     

Thyreophoran vertebrae from the Callovian (Middle Jurassic) of the Vaches Noires cliffs (Normandy,France), with remarks on the dinosaur assemblage from the Vaches Noires

Two associated incomplete thyreophoran dorsal vertebrae from the Callovian Marnes de Dives Formation of the Vaches Noires cliffs, on the Normandy coast, are referred to an indeterminate stegosaur that appears to be different from Lexovisaurus, previously reported from the Callovian of western France. These vertebrae are the first evidence of thyreophorans from the Vaches Noires and complement the dinosaur assemblage from this locality, which hitherto consisted of several theropod taxa and an indeterminate sauropod. The dinosaur record from the Vaches Noires is heavily dominated by theropods and this imbalance is difficult to explain. A possible explanation may be that the dinosaur sample from the Vaches Noires is too small to be statistically significant and representative of the original faunal assemblage from which it is derived.