基于最小限制水分范围评价不同耕作方式对土壤有机碳的影响

以2009年吉林省德惠市中层黑土上进行了8a的田间定位试验小区土壤为研究对象,对免耕和秋翻两种耕作方式及玉米-大豆轮作和玉米连作两种种植方式下耕层有机碳进行分析,分别采用加权平均和分层两种方法计算最小限制水分范围(LLWR),用其评价不同耕作方式对土壤有机碳的影响.结果表明,免耕在玉米-大豆轮作和玉米连作下0-5 cm土壤有机碳含量分别比秋翻增加了15.2％和11.5％ (P＜0.05).采用加权平均法计算的LLWR值为0.148-0.166 cm3/cm3,不同耕作方式下玉米-大豆轮作的LLWR高于玉米连作且在两种种植方式下均表现出免耕小于秋翻的特点;利用分层法计算得到的LLWR值介于0.130-0.173 cm3/cm3之间,玉米-大豆轮作和玉米连作下免耕0-5 cm LLWR均显著小于秋翻,而5-30 cm LLWR数值免耕大于秋翻(P＞0.05);玉米-大豆轮作下0-30 cm各层LLWR均高于玉米连作.由于LLWR可以评价不同耕作方式对土壤有机碳的影响,因此采用加权平均法计算的LLWR可以客观的反映不同耕作处理尤其是种植方式对土壤有机碳的影响;而采用分层法计算的LLWR则更清晰的刻画了土壤表层与亚表层固碳能力的差异.     

The sensitivity of shoot growth of corn to the least limiting water range of soils

The least limiting water range (LLWR), the range in soil water content within which limitations to plant growth associated with water potential, aeration and mechanical resistance to root penetration are minimal, has been proposed as an index of the structural quality of soils for crop growth. An hypothesis that is implicit in the proposed use of LLWR as an index of soil structural quality is that crop growth is negatively related to the proportion of the total number of measurements in which the water content falls outside the LLWR (p_out) and therefore given a certain climate positively related to the magnitude of the LLWR. The objective of this investigation was to test the hypothesis that plant response, specifically shoot growth rate of corn (Zea mays L.), can be functionally related to p_out and the LLWR of soils. The study was carried out on a farm with a side-by-side comparison of no-till and conventional-till. Thirty two paired sampling sites were located along the two transects. The LLWR and p_out were calculated for the 0–20 cm depth in each sampling site. Shoot growth rate (SGR) was measured during a 17 and 16 day period in 1992 and 1993, respectively that corresponded to the 10–11 leaf stage. Although the variation in p_out accounted for a larger percentage of the variation in SGR than did LLWR, the correlation between SGR and LLWR was high and justifies further studies to determine if crop yield can be related to LLWR under different soil and climatic conditions.     

Soil water availability for plants as quantified by conventional available water, least limiting water range and integral water capacity

There are different approaches to define the soil available water (SAW) for plants. The objectives of this study are to evaluate the SAW values of 12 arable soils from Hamadan province (western Iran) calculated by plant available water (PAW), least limiting water range (LLWR) and integral water capacity (IWC) approaches and to explore their relations with Dexter’s index of soil physical quality (i.e., S-value). Soil water retention and mechanical resistance were determined on the intact samples which were taken from the 5–10 cm layer. For calculation of LLWR and IWC, the van Genuchten-Mualem model was fitted to the observed soil water retention data. Two matric suctions (h) of 100 and 330 cm were used for the field capacity (FC). There were significant differences (P?<?0.01) between the SAW values calculated by PAW₁₀₀, PAW₃₃₀, LLWR₁₀₀, LLWR₃₃₀ and IWC. The highest (i.e., 0.210 cm³ cm^?3) and the lowest (i.e., 0.129 cm³ cm^?3) means of SAW were calculated for the IWC and LLWR₃₃₀, respectively. The upper limit of LLWR₃₃₀ for all of the soils was h of 330 cm, and that of LLWR₁₀₀ (except for one soil that was air-filled porosity of 0.1 cm³ cm^?3) was h of 100 cm. The lower limit of LLWR₃₃₀ and LLWR₁₀₀ for five soils was h of 15,000 cm and for seven soils was mechanical resistance of 2 MPa. The IWC values were smaller than those of LLWR₁₀₀ for two soils, equal to those of LLWR₁₀₀ for three soils and greater than those of LLWR₁₀₀ for the rest. There is, therefore, a tendency to predict more SAW using the IWC approach than with the LLWR approach. This is due to the chosen critical soil limits and gradual changes of soil limitations vs. water content in the IWC calculation procedure. Significant relationships of SAW with bulk density or relative bulk density were found but not with the clay and organic matter contents. Linear relations between IWC and LLWR₁₀₀ or LLWR₃₃₀ were found as: IWC?=??0.0514 + 1.4438LLWR₁₀₀, R ²?=?0.83; and IWC?=??0.0405 + 2.0465LLWR₃₃₀, R ²?=?0.84, respectively (both significant at P?<?0.01). Significant relationships were obtained between the SAW values and S indicating the suitability of the index S to explain the availability of soil water for plants even when complicated approaches like IWC are considered. Overall, the results demonstrate the importance of the choice of the approach to be used and its critical limits in the estimation of the soil available water to plants.     

Population genetics and the evolution of geographic range limits in an annual plant

Abstract Theoretical models of species' geographic range limits have identified both demographic and evolutionary mechanisms that prevent range expansion. Stable range limits have been paradoxical for evolutionary biologists because they represent locations where populations chronically fail to respond to selection. Distinguishing among the proposed causes of species' range limits requires insight into both current and historical population dynamics. The tools of molecular population genetics provide a window into the stability of range limits, historical demography, and rates of gene flow. Here we evaluate alternative range limit models using a multilocus data set based on DNA sequences and microsatellites along with field demographic data from the annual plant Clarkia xantiana ssp. xantiana. Our data suggest that central and peripheral populations have very large historical and current effective population sizes and that there is little evidence for population size changes or bottlenecks associated with colonization in peripheral populations. Whereas range limit populations appear to have been stable, central populations exhibit a signature of population expansion and have contributed asymmetrically to the genetic diversity of peripheral populations via migration. Overall, our results discount strictly demographic models of range limits and more strongly support evolutionary genetic models of range limits, where adaptation is prevented by a lack of genetic variation or maladaptive gene flow.     

Relationship between avian range limits and plant transition zones in Maine

Aim To determine if vegetation complexity associated with transition zones may be a contributing factor affecting bird species distributions in Maine, USA, and in increased numbers of bird species at about 45° north latitude in northeastern North America. Location Maine, USA; North America north of Mexico. Methods We delineated the ranges within Maine (86,156 km²) of 186 bird species and 240 woody plants using literature and expert review. Maps showing species richness and numbers of range limits, at 324 km² resolution, were developed for woody plants and groups of breeding birds: forest specialists, forest generalists, and those that used barren and urban habitats, early successional areas, and wetlands or open water. Two plant transition zones for Maine were identified previously, with the north–south transition zone mapped across eastern North America. Patterns in bird distribution maps were compared to woody plant maps and to transition zones. Results When the distributions of forest specialists were compared to the north–south vegetation transition zone in Maine, they were spatially coincident, but were not for other groups. Forest specialists had more species with range limits in the state (61%) than generalists (13%) or any other group. At a continental‐scale, the vegetation transition zone within eastern North America agreed fairly well with the areas of highest bird richness. Main conclusions A bird transition zone occurs in Maine and across eastern North America, akin to and overlapping the vegetation transition zone. Seasonality is likely the primary source of the inverse gradient in bird richness in the eastern USA, as reported by others. However, vegetation structure and habitat selection at very broad spatial scales appear to contribute to the reversed gradient. North of the vegetation transition zone, forest structure is simpler and coniferous forests more dominant, and this may contribute to reduced bird species richness. However, the northern (> 49°) typical gradient in bird species richness has been related to many hypotheses, and several are likely involved in the genesis of the gradient.     

密云水库上游油松人工水源涵养林林下植物多样性与土壤理化特性

以密云水库上游丰宁县4种油松人工水源涵养林(油松×落叶松混交林、油松×蒙古栎混交林、油松×山杏混交林、油松纯林)为对象,研究林下植物多样性与土壤理化性质特征,分析林下植物多样性与土壤因子的相关关系。结果表明: 4种人工水源涵养林林下植物群落组成结构差异显著,油松×山杏混交林林下植物群落物种组成最为丰富,绣线菊、虎榛子和披针薹草为主要优势种。从物种丰富度指数、Simpson优势度指数、Shannon多样性指数、Pielou均匀度指数来看,油松×山杏混交林林下整体植物多样性水平最高,灌木层和草本层植物多样性分别以油松×蒙古栎混交林和油松×山杏混交林最高。4种人工水源涵养林之间除全磷外其他各理化指标均差异显著,油松×山杏混交林土壤理化性质总体最优,油松×蒙古栎混交林最差。土壤毛管孔隙度、pH、有机质为灌木层植物多样性的主要影响因子,土壤pH、毛管持水量为草本层植物多样性的主要影响因子。营造油松×山杏混交林更有利于提高林下植物多样性并促进土壤改良,土壤pH、有机质、毛管孔隙度、毛管持水量为影响研究区林下植物多样性的主导土壤因子。     

屏边空竹分布区海拔上下边界的土壤真菌群落特征

物种地理分布边界的形成一直是进化生物学中的一个重要课题。植物分布区边界如何影响土壤微生物尚不清楚。屏边空竹(Cephalostachyumpingbianense)是目前已知唯一能够在自然条件下四季出笋的珍稀竹种,仅分布于中国云南东南部,对研究竹类中的狭域分布种具有重要意义。为探究土壤真菌群落与屏边空竹分布边界间的联系,该研究测定了屏边空竹分布中心、分布边界以及分布区外的土壤理化性质,并利用基于真菌内转录间隔区(ITS)序列的IlluminaMiSeq高通量测序技术分析土壤真菌群落变化。主要结果:1)屏边空竹分布边界处土壤pH与速效磷含量显著低于其他位点。2)在分布中心,土壤真菌物种多样性最高,被孢霉属(Moritierella)相对多度显著高于其他位点;在分布边界处,土壤真菌物种多样性最低,担子菌门相对多度大于65.0%。3)土壤pH是驱动真菌群落变异的关键因素,与外生菌根真菌的相对多度负相关,与腐生真菌的相对多度正相关。综上,土壤酸化与磷缺乏可能是控制屏边空竹分布区域的重要土壤属性。土壤中的被孢霉具有溶解磷、缓解土壤酸化等功能,可能是屏边空竹重要的互利共生菌。     

芦芽山针叶林分布上下限土壤温度及含水量的季节差异

山地小气候特征对解释林线位置、形成机制以及生长-气候关系具有重要意义。由于高山气象数据匮乏,尤其是土壤水热数据的缺失,使以往对华北地区山地土壤的温湿度变化特征知之甚少。基于5个整年(2012—2016年)的连续监测,分析了华北芦芽山针叶林分布上下限土壤(10cm)温度和含水量的季节变化特征及差异。结果表明:(1)在芦芽山针叶林分布上限,北坡土壤10月末冻结,5月初解冻,南坡土壤冻结和解冻日均滞后于北坡,生长季内南北坡土壤均温、生长季长度无显著差异(122d,8.1℃和110d,7.6℃);(2)南北坡林线土壤含水量最低值都出现在冬季(1月),最高值则在秋季(10月和9月),并且南坡生长季土壤含水量(0.350 m³/m³)显著大于北坡(0.247 m³/m³);(3)与针叶森林的分布下限(2040 m a.s.l.)相比,林线土壤热量指标(年均温、生长季均温、最热月均温和生长季长度)均明显偏低,而土壤生长季内含水量显著偏大。研究结果揭示了亚高山区土壤冻融过程中温度和含水量的耦合关系,并进一步证实了芦...     

土壤水分胁迫对夏玉米植株性状整齐度的影响

利用大型防雨设施池栽,严格调控水量,研究夏玉米全程及阶段性土壤水分胁迫对植株性状整齐度的影响。结果表明,夏玉米生育全程水分胁迫,植株性状整齐度全面劣化,产量极低;苗期阶段水分胁迫导致大、小苗,壮、弱苗两极分化明显,株高整齐度显著下降,其负效应持续至生育后期,生育进程推迟,千粒重显著降低,对产量造成一定影响;穗期阶段水分胁迫对穗长、穗料数整齐度影响明显,产量降幅度较大;花粒期阶段水平胁迫对千粒重及穗     

Arctic and boreal plant species decline at their southern range limits in the Rocky Mountains

Climate change is predicted to cause a decline in warm‐margin plant populations, but this hypothesis has rarely been tested. Understanding which species and habitats are most likely to be affected is critical for adaptive management and conservation. We monitored the density of 46 populations representing 28 species of arctic‐alpine or boreal plants at the southern margin of their ranges in the Rocky Mountains of Montana, USA, between 1988 and 2014 and analysed population trends and relationships to phylogeny and habitat. Marginal populations declined overall during the past two decades; however, the mean trend for 18 dicot populations was ?5.8% per year, but only ?0.4% per year for the 28 populations of monocots and pteridophytes. Declines in the size of peripheral populations did not differ significantly among tundra, fen and forest habitats. Results of our study support predicted effects of climate change and suggest that vulnerability may depend on phylogeny or associated anatomical/physiological attributes.     

金矮生苹果土壤水分合理供给范围研究

对7年生田间和2年生盆栽金矮生苹果(Malus pumila)不同土壤水分(SWC)条件下水分利用效率(WUE)研究表明。SWC约为10％时,WUE最大值在所有处理中最高(230μmolCO₂·g^-1 H2O);在水分供应充足时,WUE最大值只有160μmolCO₂·g^-1 H2O左右,各SWC水平间差异不显著．WUE随SWC的变化与光照条件有关。     

岷江上游森林土壤大孔隙特征及其对水分出流速率的影响

大孔隙是森林土壤中常见的现象 ,对土壤壤中流的产生有重要的影响。但由于大孔隙研究方法的不成熟 ,各种方法得到的大孔隙半径范围并不一致。一般认为田间持水量和饱和含水量之间的土壤孔隙为大孔隙 ,利用水分穿透曲线和 Poiseuille方程研究了岷江上游不同植被下土壤的大孔隙状况 ,这种方法一方面与传统的研究方法相衔接 ,另一方面所得到的大孔隙与土壤水分运动有关 ,反映了土壤大孔隙的研究目的 ,因而是一种相对合理的研究方法。岷江上游几种主要植被下土壤大孔隙半径主要集中于 0 .3～ 2 .4 mm之间 ,平均在 0 .4 8～ 1.17mm之间 ,均值为 0 .84 mm,均方差为 0 .2 2 6 ;且随着剖面的发育表现出上部土层多 ,下部土层少的特点。同时 ,半径在 2 .4～ 1.4 m m之间的特大孔隙较少 ,<1.0 m m的小孔隙较多。大孔隙的平均半径对于水分出流速率有重要的影响 ,特别是半径 >1.4 mm的孔隙数量影响最大 ,虽然其数量仅占大孔隙数量的 5 %以下 ,但决定了稳定出流速率 70 %的变异。大孔隙所占过水断面的最高比例为 2 1.2 2 % ,最低为 2 .6 % ;在大孔隙所占过水断面的比例小于 2 0 %的条件下 ,稳定出流速率随大孔隙的增多而增大     

Mutation accumulation and the formation of range limits

The dynamics of range formation are important for understanding and predicting species distributions. Here, we focus on a process that has thus far been overlooked in the context of range formation; the accumulation of mutation load. We find that mutation accumulation severely reduces the extent of a range across an environmental gradient, especially when dispersal is limited, growth rate is low and mutations are of intermediate deleterious effect. Our results illustrate the important role deleterious mutations can play in range formation. We highlight this as a necessary focus for further work, noting particularly the potentially conflicting effects dispersal may have in reducing mutation load and simultaneously increasing migration load in marginal populations.     

Interactions between soil habitat and geographic range location affect plant fitness

Populations are often found on different habitats at different geographic locations. This habitat shift may be due to biased dispersal, physiological tolerances or biotic interactions. To explore how fitness of the native plant Chamaecrista fasciculata depends on habitat within, at and beyond its range edge, we planted seeds from five populations in two soil substrates at these geographic locations. We found that with reduced competition, lifetime fitness was always greater or equivalent in one habitat type, loam soils, though early-season survival was greater on sand soils. At the range edge, natural populations are typically found on sand soil habitats, which are also less competitive environments. Early-season survival and fitness differed among source populations, and when transplanted beyond the range edge, range edge populations had greater fitness than interior populations. Our results indicate that even when the optimal soil substrate for a species does not change with geographic range location, the realized niche of a species may be restricted to sub-optimal habitats at the range edge because of the combined effects of differences in abiotic and biotic effects (e.g. competitors) between substrates.     

Soil erosion alters soil chemical properties and limits grassland plant establishment on an oceanic island even after goat eradication

Soil erosion after vegetation degradation induced by disturbance by feral goats, an invasive mammal, can lead to loss or accumulation of soil at a local scale and can alter soil chemical properties. This alteration of soil properties can in turn affect the establishment of plant species. We evaluated relationships among the extent of soil erosion, soil chemical properties, and the distribution of plant species in grassland ecosystems after goat eradication on Nakodo‐jima Island in the northwestern Pacific. In 105 1 × 1–m quadrats, changes were measured in the position of topsoil over 2 years. Surface soils were sampled for analysis of chemical properties, and plant species in the quadrat were recorded. Changes in the position of topsoil were related to the area of bare ground. Soil loss occurred at sites where areas of bare ground were extremely large. Significantly higher values of soil exchange acidity and smaller amounts of available phosphorus, total carbon, and total nitrogen were detected in soils at sites with large soil losses. Most of the 11 dominant plant species were absent from sites with large losses of soil. The presence of eight species was significantly negatively related to soil exchange acidity, and three species were significantly positively related to available phosphorus. Our results indicated that exposure of subsoils at the soil surface after vegetation degradation can increase soil loss, which can alter soil chemical properties, and this alteration can continue to limit the establishment of plant species, even long after goat eradication.     

Comparative resistance of the soil and the plant to water transport

The resistances to liquid water transport in the soil and plant were determined directly and simultaneously from measurements of soil, root, and leaf water potentials and the flux of water through the soil-plant system to the sites of evaporation in the leaf. For soybean (Merr.) transporting water at a steady rate, water potential differences between soil and root were smaller than between root and leaf over the range of soil water potentials from −0.2 to −11 bars. As soil water was depleted, water flow through the soil and plant decreased to one-tenth the maximum rate, but both the soil resistance and plant resistance increased. The plant resistance remained larger than the soil resistance over the entire range of soil water availability. Previous suggestions that the soil is the major resistance have ignored the increase in plant resistance and/or assumed root densities that were too low.     

Woody plant encroachment alters soil hydrological properties and reduces downward flux of water in tallgrass prairie

Plant and Soil - Plant and soil interact to shape ecosystem properties, processes and services provided. Changes in ecosystem productivity, biogeochemical cycling and plant herbivore interactions...     

上海外环线绿地群落凋落物对土壤水分物理性质的影响

选取上海外环线绿化带8个人工植物群落为对象,以裸地为对照,研究了凋落物量、土壤孔隙性状和持水特性,探讨了城市绿地群落凋落物对土壤水分物理性质的影响.结果表明:各群落0～10 cm土壤的最大持水量、毛管持水量、最小持水量、质量含水量、体积含水量、毛管孔隙度、非毛管孔隙度、总孔隙度和土壤通气度均显著高于裸地(P<0.05),而容重均显著低于裸地(P<0.05);10～20 am土壤水分物理性质差异不显著;凋落物年凋落量与绿地群落0～10 cm土壤毛管孔隙度呈显著负相关,而与绿地群落0～10 cm土壤持水特性相关性均不显著;凋落物现存量与绿地群落0～10 cm土壤总孔隙度和通气度呈显著正相关,与0～10 cm土壤容重呈显著负相关;而凋落物年凋落量、现存量与绿地群落10～20 cm土壤水分物理性质相关性均不显著;绿地群落凋落物蓄积能有效改善绿地群落内表层土壤(0～10 cm)水分物理性质,主要是改良土壤孔隙状况;在绿地养护中应保留绿地群落凋落物.