Fluctuations morphologiques et hétérochronies chez Toxaster (échinides,Crétacé inférieur)

Morphological fluctuations and heterochronies of Toxaster (echinoid, Lower Cretaceous). Several steps of the diversification of the spatangoid Toxaster are described with an analysis of six species distributed over the north-west Tethyan margin, and ranging from the Lower Valanginian to the Barremian. Morphological variation was investigated with the analysis of two parameters (angles), which allowed recognition of three modules on the test. Comparison of ontogenetic trajectories of two reference species (T. granosus kiliani and T. retusus) suggests a peramorphic heterochronic pattern. However, no peramorphocline can be attested when comparisons are extended to adults of the six species. Evolution among these species is partially driven by allometric changes in one module, indicating an important influence of size. On the temporal and spatial scale of the study, the evolution of Toxaster appears globally quite conservative. To cite this article: É. François et al., C. R. Palevol 2 (2003).     

Succession des peuplements d’échinides du Crétacé inférieur dans le Haut Atlas atlantique (Maroc) ; révision systématique et intérêt stratigraphique

The rich echinoid fauna from the Lower Cretaceous of the Agadir, Haha and Essaouira basins has been revised taxonomically, based on new and abundant collections. Forty-seven species have been identified, of which 20 are reported for the first time from this region and a further three represent new species left in open nomenclature. Their stratigraphic ranges have been determined accurately by reference to the regional ammonite zonation. However, it is also possible to use a local biostratigraphical scheme based on the echinoid succession, where ammonites are rare. The Toxasteridae, which are very abundant and underwent rapid evolution, are the most useful biostratigraphically. The genus Toxaster appears in the Lower Berriasian and continues until the Gargasian passing through the following succession: T. africanus, T. granosus, T. kiliani, T. lorioli, T. obtusus, T. exilis, T. retusus, T. maurus, T. peroni, T. collegnoi. The presence of Eoholaster, Macraster, Douvillaster, Palhemiaster, Heteraster and Pygopyrina is signaled for the first time in the Atlantic High Atlas region.     

Late Viséan foraminifers and algae from the Cataloturan Nappe, Aladag Mountains, eastern Taurides, southern Turkey

The Aladag Mountains at the western end of the eastern Taurides, southern Turkey, are composed of nappe structures imbricated in the Late Cretaceous by ophiolite emplacement. The Cataloturan Nappe contains Mississippian (i.e., Early Carboniferous) deep-water (basinal-lower slope) facies in the lower part and shallow-water (shelf edge) facies in the upper part. These Mississippian strata belong to the Nohutluk Formation. The main outcrop occurs at Nohutluktepe where it is 460 m thick. Eighty taxa of algae, foraminifers, and calcareous microproblematica permit dividing the upper part of the formation into three informal assemblage biozones: (1) Howchinia bradyana longa - Lituotubella magna - Koktjubina (?) sp. biozone (late Viséan Cf6γ1), (2) Bradyina rotula - Euxinita tauridiana biozone (Cf6γ2), (3) Janischewskina typica - Biseriella aff. parva biozone (upper Cf6δ). The genus Euxinita is discussed and a new species E. tauridiana nov. sp. is described. A paleobiogeography, based on Euxinita, Koktjubina (?), Vissarionovella, Janischewskina and Hortonella, is reconstructed, where the Taurides are juxtaposed to the northwestern border of the Paleotethys, with possible direct connections with Russia, Kazakhstan, western Europe, and Morocco.     

Données paléoenvironnementales sur le Vraconnien/Cénomanien de la région de Tébessa (Atlas Saharien,nord-est Algérie). Caractérisation de l’OAE2

The micropalaeontological content of five sections, located in Northeastern Algeria (Saharan Atlas) was investigated by means of washing/counting of microfossils in marly levels, and microfacies analyses of calcareous levels. In these levels of Upper Cretaceous age, hundred species of foraminifera were identified but only about 15 species of ostracoda and about ten of radiolaria. This work allows, first of all, to establish a rather precise stratigraphic frame, in particular by means of studying planktonic foraminifera. Two to five biozones were defined, between the Vraconnian (Th. Appenninica biozone) and the early Turonian (Whiteinella archaeocretacea biozone, then Helvetoglobotruncana Helvetica biozone for certain sections). Secondly, the quantitative analyses led on foraminifera allowed the definition of palaeoenvironment. The ratio P/P+B, generally very high, coupled with a little diversified benthonic microfauna, indicates a calm and deep environment, of external platform or slope type. Furthermore, at numerous levels, various indications give evidence of the existence of a strong surface productivity (presence of upwellings), responsible for the proliferation of radiolaria (late Vraconnian/early Cenomanian especially) or of globular planktonic foraminifera (hedbergellids/heterohelicids); being both associated with low-oxygen deep waters. Two anoxic events were also revealed, the first one at the end of Vraconnian (OAE1d) and the second at the end of Cenomanian (OAE2). This last event in particular was characterized on all the sections, in a more or less detailed way, thanks to the identification of certain indicator: Heterohelix “bloom”, “filament” event, disappearance of rotaliporids, presence of “blackshales” strongly enriched in organic matter (Bahloul levels).     

Identification des zones biostratigraphiques méditerranéennes dans le Pliocène du Cap-Bon (Tunisie)

The Nabeul-Hammamet series analysis allows to recognize the six biozona known in the Mediterranean realm, as specified by Cita (1975a). These are: the Sphaeroidinellopsis subdehiscens BANNER & BLOW acme biozone equivalent to MPL1; the Globorotalia margaritae margaritae BOLLI & BERMUDEZ interval zone (= MPL2); the Globorotalia margaritae BOLLI & BERMUDEZ, Globorotalia puncticulata (DESHAYES), concurrent range zone (= MPL3); the Sphaeroidinellopsis subdehiscens BANNER & BLOW interval zone (= MPL4); the Globigerinoides elongatus (D’ORBIGNY) interval zone (= MPL5); the Globorotalia inflata (D’ORBIGNY) interval zone (= MPL6). There is no similarity between these biozones and those established by Berggren and al. (1995) for the atlantic and the indo-pacific realms. The MPL1, MPL2, MPL3 and MPL4 zones are especially englobed in argillaceous and yellow sandy lithological units specified in north-eastern Tunisia, such as ‘Argiles des Potiers’, ‘Sables Jaunes de Nabeul’ and ‘Argiles de Sidi Barka’. These units are together equivalent to the Raf-Raf Formation developed towards the North, in the Bizerte area. Besides, the MPL5 and the MPL6 zones are included in sands and sandstones composing the Hammamet unit, equivalent to the Porto-Farina Formation.     

Pliocene ostracods of Southwestern Europe

This paper analyzes the lithostratigraphic features and ostracod associations of the Pliocene deposits located in the western sector of the Guadalquivir basin (SW Spain). A regressive cycle is defined by the vertical distribution of five main facies (FA-1 to FA-5), which were deposited in upper bathyal to fluvial palaeoenvironments. The broad palaeontological record is utilized to draw a general outline of the food chain in each palaeoenvironment. A comparison with the Pliocene ostracod distribution of southeastern Spain, southwestern France and Portugal permits to define a common ostracod biozonation for southwestern Europe during this period. Four main marine biozones are distinguished: (a): Pontocythere elongata biozone (infralittoral); (b): Costa edwardsii biozone (shallow circalittoral); (c): Bosquetina carinella biozone (deep circalittoral); and (d): Henryhowella asperrima biozone (upper bathyal). Freshwater, brackish and lower bathyal/abyssal species are very scarce in this area during this period.     

Calcareous and siliceous phytoplankton stratigraphy of Neogene marine sediments in central Crete (Greece)

Fine laminated diatomaceous beds in outcrops of the Finikia unit (Estavromenos and Athanatoi sections) in the northern part of the Heraklion district in Crete, have yielded a total of 55 marine diatom and 24 silicoflagellate taxa. The diatom assemblages, characterized by species of low latitudes, belong to the upper part of the Nitzschia jouseae biozone (early-late Pliocene). The silicoflagellate associations belong to the local Dictyocha fibula ausonia and Dictyocha hellenica subzones. These subzones correspond to the subzones CN11b (Discoaster asymmetricus) and CN12a (D. tamalis) of calcareous nannofossils respectively, as well as to the Globorotalia puncticulata/G. bononiensis biozones of planktic foraminifera. The sparse silicoflagellate association from fine sandy marls of the Panassos section in the central part of the Heraklion district (Aghia Varvara unit, upper Tortonian-Messinian) is assigned to the local biozone Distephanus speculum minutus. The latter corresponds approximately to the biozone C9 (Discoaster quinqueramus) of calcareous nannofossils and to the Globorotalia conomiozea zone of planktic foraminifera. No discoidal diatoms could be found in the samples from Panassos-section. The presence of silicoflagellates but simultaneous absence of diatoms is probably due to greater opal dissolution within the fine sandy marls of the Panassos section.     

Stratigraphie, paléoécologie et foraminifères du paléogène des Alpes Maritimes et des Alpes de Haute-Provence (Sud-Est de la France)

Paleogene sedimentation in the Alpes Maritimes and Alpes de Haute Provence regions is composed of one complete sedimentary sequence. This sequence begins with the continental Formation from Microcodium and continues with the Cerithium layers, the Calcaires Nummulitiques and the Marnes Bleues, which are overlaid by the turbidites of the Grès d’Annot Formation. Sedimentation starts in the Nummulites perforatus zone close to the base of ‘the Truncorotaloides rohri zone (P 14) and ends in the Cassigerinella chipolensis-Pseudohastigerina micra zone (P 18) and the NP21 zone in the upper part of the Grès d’Annot. More biomarkers were used in order to define a more detailed local biozonation (biozones AMP 1 to AMP 7). Four local zones were also defined by the last occurrence of Nummulites millecaput and N. perforatus-N. ptukhiani and then by the first appearance of N. retiatus (AMGF 1-4). The evolution of benthic foraminiferal assemblages shows an increase in bathymetry from the internal platform in the Cerithium layers to the calcareous platform with large foraminifers, then to the external platform and the deep offshore environment of the Marnes Bleues and Grès d’Annot. Low faunal diversity in the Grès d’Annot together with the predominance of agglutinated species indicates a poorly oxygenated, organic rich and turbidite environment. Seven steps (SD 1 to SD 7) in the Eocene marine transgression are shown from East to West by detailed local biozonation and sequence analysis. Grès d’Annot sedimentation is also diachronous, beginning within the Truncorotaloides rohri zone (P 14, AMP 1) in the East and ending within the Pararotalia opima opima zone (P 20, middle Rupelian) in the West (Barrême). Small foraminifer Paleogene fauna from the Alpes Maritimes and Alpes de Haute Provence was studied from 400 samples. It is composed of 378 species. Two new taxa are proposed: Fissurina niceana n. sp. and Globocassidulina alpina n. sp. The species from the Escarène and Gorbio neighborhood described by M. von Hantken (1884) were re-examined.     

Matière organique sédimentaire et paléoenvironnements du Tithonien inférieur quercynois (France)

The Solen 98 well corresponds to the limestones of the Lower Tithonian Cazals Formation (Gigas Zone). The iterative succession of six sedimentary terms expresses a cyclic peritidal dynamic. Limited by two emersion surfaces, each sequence evolves from an upper subtidal lagoon to a tidal flat, upper intertidal or supratidal environment, and ends with open sea depositional bioclastic and oolitic shoals deposits. Hierarchical ascendant classification applied to palynological data define 6 palynofacies types associated with different depositional facies. Type 1, characterizing open marine deposits, shows a diversified and balanced assemblage. The blade-shape woody particles are abundant and the amorphous organic matter is absent. Types 2 and 3 are linked to lagoonal and skeletal shoals deposits. Then microfossil population is dominated by Corculodinium or long-spine Micrhystridium. The Shannon-Weaver and the equitability indices are moderate. Type 4 is associated with the upper tidal flat, lagoonal and skeletal shoal deposits. When microfossils are present, the algal assemblages are more balanced than in type 5. This type, observed in all the palaeo-environments except the open marine, is enriched in elements attributed to the Hyalinsphaeridia complex. The marine component assemblages are balanced. The amorphous organic matter is relatively abundant and the oxydized woody particles absent. Type 6, mainly composed of amorphous organic matter and phytoclasts, is principally associated with the stromatolitic facies of tidal flat deposits. The example of the Solen 98 well, shows that hierarchical ascendant classification method is well suited for identification of palynofacies     

Distribution de Megaporellaboulangeri Deloffre et Beun, 1986 (algue dasycladale) dans le Jurassique du bassin atlantique marocain

The recent biostratigraphic and micropaleontologic works permitted to specify the distribution of Megaporellaboulangeri Deloffre and Beun in the Jurassic of the Moroccan Atlantic basin. The lower limit of the extension of this dasycladale alga, earlier attribued to the Early Kimmeridgian by Deloffre and Beun (1986), is currently placed in the Early Callovian, on the basis of the age provided by Megaporellaboulangeri biozone. The Otternstellaarabica biozone dates the upper limit of the extension of the species of the Early Oxfordian. Megaporellaboulangeri proliferates in the borders of the basin where the internal lagoons facies dominate, however it is absent in the western parts with open sea facies.     

Les kystes de dinoflagellés des Argiles Ostréennes du Bassin de Paris, France

Palynological studies realized from a borehole drilled on the site of ANDRA (Agence Nationale pour la gestion des Déchets RAdioactifs; Aube Centre near Soulaines, Paris Basin, France) show a sedimentary discontinuity between, on one hand, the Argiles à Plicatules (lower Aptian in age) and Argiles Ostréennes (Lower Barremian/Upper Barremian in age) and, on the other hand, between these same Argiles Ostréennes and the Calcaire à Spatangues (Lower Hauterivian). These conclusions argue against the chronostratigraphical subdivision recently proposed on the basis of sequence stratigraphical and sedimentological interpretations.     

Depositional model and paleodepth reconstruction of a coral-rich,mixed siliciclastic-carbonate system: the Burdigalian of Capo Testa (northern Sardinia,Italy)

This study presents a detailed facies analysis and paleodepth reconstruction of a coral-rich mixed siliciclastic-carbonate system Burdigalian in age, outcropping in the northern sector of Sardinia (Capo Testa). Excellent exposures of continuous sea-cliff outcrops around the southwestern and northeastern area of Capo Testa promontory allowed us to: (1) trace stratigraphic surfaces; (2) document stratal geometries; (3) discern details of the lithofacies and, (4) reconstruct the paleodepths of the different depositional environments. A total of seven sedimentary facies has been recognized and interpreted: siliciclastic conglomerate and coarse bioclastic sandstone (F1), fine- to medium-grained hybrid sandstone (F2, scleractinian coral domestone (F3), bioclastic packstone to floatstone with platy Porites (F4), red algae floatstone to rudstone (F5), larger benthic foraminifers (LBF) bioclastic rudstone floatstone in a packstone matrix (F6), molluscan floatstone in a bioclastic packstone matrix (F7). The investigated system is characterized by nearshore to shoreface deposits with a conspicuous terrigenous content that grades seaward into deeper zones where coral patch-reefs developed in association with adjacent areas colonized by seagrass meadows. The more distal facies are constituted by scattered encrusting tabular colonies of Porites in growth position occurring in a deeper and lower-energy environment. The paleodepth interval that is observed in the Capo Testa outcrop ranges from 0 to 50 m.     

Biostratigraphie,paléoécologie et zones d’acmé des foraminifères planctoniques au passage Crétacé-Paléogène dans la Téthys (Tunisie et Espagne) et l’Atlantique (France)

Biostratigraphical high-resolution analyses and quantitative data confirm that deposition is continuous across the K-Pg transition in several sections in Tunisia (El Kef, stratotype section) and Spain (Agost and Caravaca sections) located in the Tethyan realm and the Bidart sections in the Atlantic realm, without any relevant hiatus. The Upper Maastrichtian assemblages of planktic foraminifera from these sections are largely dominated by small biserial heterohelicids. They are associated to common species having planispiral test (i.e. globigerinelloids), trochospiral test (i.e. hedbergellids, rugoglobigerinids globotruncanids), to rare triserial heterohelicids (i.e. guembelitriids) and trochospiral species showing tubulospines (i.e. schackoinids). Stratigraphical ranges of these diverse taxa through the late Maastrichtian in the Tethyan and Atlantic realms show very few changes in the planktonic foraminiferal assemblages and most of the species are present in the Abathomphalus mayaroensis biozone. By our high-resolution sampling and the intensive research for the A. mayaroensis index species in the uppermost Maastrichtian samples, we confirm that this species is omnipresent up to the top of the Maastrichtian. Therefore, A. mayaroensis is present in almost all samples which are late Maastrichtian in age, but this species became very scarce in the uppermost Maastrichtian samples. This scarceness could be due to a climate cooling. A sharp decrease in relative abundance of the deep dwellers species, like as Abathomphalus intermedius and A. mayaroensis as well as in other keeled globotruncanids is observed at the studied sections from the Tethyan realm (indicative of low latitude) across the latest Maastrichtian. At the K/Pg boundary, all the globotruncanids disappeared. They are considered specialists living in tropical-subtropical deep seawater habitat. At this boundary, large and ornate heterohelicids also disappeared. Therefore, all the studied sections show that about 90 % of the Maastrichtian species became extinct according to a catastrophic mass extinction pattern. Only about 10 % crossed the K/Pg boundary and survived during the earliest Danian. The minor difference in the number of disappeared taxa is related to their latitude location or environment paleodepth. The changes in the species relative abundance, observed in the successive planktic foraminiferal assemblages, make it possible to recognize the Acme-stage 0 typical of the upper Maastrichtian interval. It is characterized by the highest species richness of Globotruncanids and heterohelicids specialists of tropical to subtropical marine conditions, the Acme-stage 1 typical of the Guembelitria cretacea Zone, and in particular of the Hedbergella holmdelensis Subzone dominated by “opportunists” species belonging to Guembelitria, the Acme-stage 2 which corresponds to the Pv. eugubina Zone dominated mainly by specimens belonging to Palaeoglobigerina and Parvularugoglobigerina and the Acme-stage 3 which characterizes mainly the Ps. pseudobulloides Zone dominated by biserial species belonging to Chiloguembelina and Woodringina.     

Biostratigraphical value of some Strunian (Devonian, uppermost Famennian) Productidina, Rhynchonellida, Spiriferida, Spiriferinida brachiopods

The space of Devonian time considered in this paper corresponds to the uppermost part of the Famennian or the latest Famennian as a fourfold subdivision of the stage, usually called by many authors Strunian in neritic facies and Wocklumeria Stufe or Wocklumian in pelagic facies. Here, we examine the biostratigraphical value of certain brachiopod genera and species as bio-markers of the uppermost Famennian throughout the world (Europe, the former USSR, Middle East, Asia, North America, North Africa, and Australia). We have focused our study for species which stratigraphical range has been firmly established in correlation with the conodont biozones (Upper expansa, Early, Middle and Late praesulcata), the ammonoid biozones (do VI = “Wocklumeria Stufe”), or the foraminifera biozones (kobeitusana Zone). Other data for which the stratigraphical range is not based on standard biozonation are provided in the annex. The brachiopod bio-markers discussed in this paper belong to 14 Productidina genera (Acanthatia, Araksalosia, Ericiata, Hamlingella, Mesoplica, Nigerinoplica, Orbinaria, Ovatia, Rugauris, Semiproductus, Sentosia, Spinocarinifera, Steinhagella, Whidbornella), 7 Rhynchonellida genera (Araratella, Centrorhynchus, Hadyrhyncha, Megalopterorhynchus, Novaplatirostrum, Rozmanaria, Tchanakhtchirostrum), 13 Spiriferida genera (Brachythyris, Cyrtospirifer, Dichospirifer, Eochoristites, Imbrexia, Parallelora, Prospira, Rigauxia, Sphenospira, Tenisia, Toryniferella, Tylothyris, Voiseyella) and one Spiriferinida genus (Syringothyris). Other orders have not been studied in this paper. The main features of the uppermost Famennian brachiopod taxa represented in this paper are commented. All these taxa are listed as completely as possible throughout the world (with complements in the annex). A quick summary of the geographic distribution of the more represented taxa is given in conclusion.     

Échinodermes de l'Ordovicien supérieur (Ashgill) de Sardaigne et d'Algérie

Research on the influence of sea level variations on the benthic faunas have been carried out in the Upper Ordovician of Sardinia. Study of the depositional facies and sequence analysis of the upper part (Lower Ashgill) of the Portixeddu Formation led to the identification of the sedimentary environments. Cystoids and crinoids are associated to bryozoans and brachiopods in most levels. The numerical analysis of associations and megaguilds shows that crinoids and cystoids have a higher frequency in the proximal and median facies of the upper offshore. The columnal association characterized by Conspectocrinus celticus and the coronoid Mespilocystites tregarvanicus has been discovered in the upper part of the formation. This material and complementary samples from Upper Ordovician of Sardinia and Kabylia (Algeria) bring additional data on the systematic and show the wide distribution of this fauna outside of the Ibero-armorican domain. The distribution of this echinoderm association supports a palaeogeographical position of the Ibero-armorican domain and Sardinia within the north gondwanan margin during the Lower Palaeozoic.     

Early Cretaceous stratigraphy and SHRIMP U‐Pb age constrain the Valanginian–Hauterivian boundary in southern Tibet

Wan, X., Scott, R., Chen, W., Gao, L. & Zhang, Y. 2011: Early Cretaceous stratigraphy and SHRIMP U‐Pb age constrain the Valanginian–Hauterivian boundary in southern Tibet. Lethaia, Vol. 44, pp. 231–244. The Late Jurassic to the Early Cretaceous marine strata are extensively distributed in southern Tibet. In Gyangze, the strata are divided into the Weimei and Jiabula formations. In Nagarze, they are divided into the Weimei and Sangxiu formations. Previous work has reported diverse ammonite species of Haplophylloceras and Himalayites in the Weimei Formation, and a few species of Spiticeras in the lower Jiabula and Sangxiu formations. The present study has found the bivalve Inoceramus and nannofossil assemblages in the lower Jiabula and Sangxiu formations. The nannofossil assemblage of Nannoconus steimannii steinmannii, N. steinmannii minor and Watznaueria barnesae indicates Berriasian age, and the Calcicalathina oblongata–Speetonia colligata assemblage is Valanginian in age. Numerical ages for the Jiabula and Jiabula‐goukou sections in Gyangze have been interpolated by comparing the fossil ranges with ages calibrated in other sections. The correlation experiment plots fossil ranges in the two sections to the CRET1 Database. The estimated rate of sediment accumulation of the lower Sangxiu Formation is 22.6 m/myr. The Jurassic–Cretaceous (J/K) boundary is at the bottom of the Jiabula Formation in Gyangze, and the base of the Sangxiu Formation in Nagarze. The boundary is marked by the appearance of the ammonite Spiticeras and the nannofossil assemblage of Nannoconus st. steinmannii–N. st. minor–Watznaueria barnesae. The radiometric age in Tibet is the first to be integrated with upper Valanginian fossils. The volcanic rocks of the upper Sangxiu Formation are dated at 136 ± 3.0 Ma deduced from zircon SHRIMP age of rhyolite. By consideration of the rate of sediment accumulation of the underlying sedimentary deposits, the J/K boundary in the Gyangze–Nagarze area is approximately 145 Ma as suggested by the newly issued International Stratigraphic Chart, and the Valanginian/Hauterivian boundary lies between 134 Ma and 136 Ma. □ Biostratigraphy, graphic plot, Jurassic/Cretaceous boundary, nannofossil, SHRIMP U‐Pb age, Southern Tibet, Valanginian/Hauterivian boundary.     

Rongeurs et lagomorphes du Miocène inférieur de Béon 2 (MN4, Montréal-du-Gers,SW France)

About 200 micromammal isolated teeth (rodents, lagomorphs) originating from the middle Orleanian locality of Béon 2 (Montréal-du-Gers, SW France) are described. The rodent fauna is dominated by myomorphs (7 species), including the cricetid Democricetodon aff. hispanicus Fahlbusch, the melissiodontid Melissiodon sp., the glirids Peridyromys murinus (Pomel), Pseudodryomys aff. ibericus, Pseudodryomys aff. simplicidens, and Glirudinus modestus (Dehm), and the eomyid Ligerimys aff. florancei. Sciuromorphs are represented only by Heteroxerus rubricati Crusafont, Villalta and Truyols. Within lagomorphs, 2 dental morphs referred to the ochotonid Prolagus Pomel are identified in Béon 2. They correspond to Prolagus oeningensis (König) and P. aff. vasconiensis. On a biostratigraphical point of view, this study confirms the location of Béon 2 at the early part of the MN4; this locality is older than other regional sites such as Pellecahus, Béon 1, La Romieu, and Bézian (MN4b), and it is coeval to Artenay, in the Loire basin. In particular, glirids and lagomorphs from Béon 2 testify to close relationships with micromammal localities referred to the MN3 biozone.     

Biostratigraphie des brachiopodes et âge carixien (Pliensbachien inférieur) des calcaires à grands bivalves en Algérie occidentale

Recent researches on the Lower Jurassic of Western Algeria allow to establish a lithostratigraphic standard correlating the different members and formations developed in the Ouarsenis and Tlemcen Mountains, the Oran High Plains, the Nador and Ksour Mountains. The position of the large bivalve limestones (= Lithiotis limestones) is well established in the different lithologic successions. This facies is widespread in Western Algeria where it is interbedded with brachiopod marker beds, indicating short periods of maximum flooding. The large number of collected brachiopods are distributed into four “faunas” (assemblages) ranging from the Late Sinemurian (= Lotharingian) to the Early Pliensbachian (= Carixian). These faunas have been dated by the age of the species that they have in common with the NW european and western tethyan provinces. These chronological data are confirmed by rare ammonites. All these results evidence the age of the large bivalve facies in Western Algeria. They are contained in the Middle to Late Carixian (Demonense and Dilectum Zones). This datation is in conformity with that known from the Eastern High-Atlas (Bou Dahar). Consequently, the large bivalves cannot be considered as “markers for the unique Domerian” as it has been too often asserted. The palaeontological part of our study shows that the multicostate Zeilleriids (several Tauromenia species from the Late Sinemurian to the Early Carixian) are older than the multicostate Terebratulids (Hesperithyris species from the Middle to Late Carixian).     

Paleogene terrigenous (flysch) sequences in Etoloakarnania region (W. Greece). Plankton stratigraphy and paleoenvironmental implications

Based on calcareous nannofossils and planktonic foraminifera1 biostratigraphic data from flysch sequences, we give evidence for the paleoenvironmental evolution of Gavrovo and Ionian foreland basins (External Hellenides, Etoloakarnania region). Our data suggest that the onset of clastic sedimentation in both foreland basins in the study area is chronostratigraphically placed at Late Eocene (from 36.2-34.4 Ma; nannofossil biozones NP19-20, planktonic foraminifera biozones P16-17). During the earliest Oligocene (NP21-22 nannofossil biozones/34.4-32.45 Ma), both basins represent restricted accumulation of sediments, mainly composed of clays and silts. The presence of thick flysch deposits, accumulated during Early Oligocene (33.4-30 Ma, nannofossil zone NP23), indicates an increasing rate of sediment supply. The flysch sequences in the Ionian basin are associated with a distal depositional environment, while in the same time the sedimentation in the external part of Gavrovo basin is related to a more proximal environment that is gradually deepening. On the contrary, the internal part of Gavrovo basin is characterised by deep-water facies, deposited in the Early Oligocene. At the end of Early Oligocene and the onset of Late Oligocene (nannofossil zone NP24/30-27.2 Ma, planktonic foraminifera zone P21), the deposition of coarse grained sediments in both basins indicates a shift to shallower depositional environment. The accumulation of fine-grained sediments during Late Oligocene (27.2-23.2 Ma, NP25 nannofossil biozone) in the Ionian basin marks the youngest flysch sediments in the Etoloakarnania region and specifies the time of the Gavrovo nappe emplacement on the Ionian zone. Moreover the emplacement of Pindos nappe on the Gavrovo zone is estimated between 30-27.2 Ma (NP24 biozone) as supported by the nannofossil analysis of samples in front of Pindos thrust.