华南贵州宾夕法尼亚亚纪牙形类Neognathodus动物群:演化及地层对比意义

以华南斜坡相的罗甸纳庆剖面为代表,我国石炭纪宾夕法尼亚亚纪早–中期地层建立起了基于多个属种的牙形类分带,并可与全球其他地区同期地层进行广泛对比。Neognathodus属是这一时期一类常见的浅水相牙形类分子,在北美、东欧等地的浅水相地层中被广泛用作地层划分对比标志。该属分子在我国北方地区浅水相地层中是重要的带化石分子,而在华南斜坡相剖面中占比非常低。结合现有的生物地层框架,本文厘清了华南贵州罗甸附近3条斜坡相剖面中Neognathodus属分子的延限,尝试利用牙形类P1分子齿台的对称特征将其演化过程归纳为五个阶段,并提出阶段4中对称分子(如:N. bothrops, N. colombiensis, N. nataliae等)的再次出现可作为全球莫斯科阶底界的辅助对比标志。     

Conodontos carboníferos de la sección del río Cares (Unidad de Picos de Europa, Zona Cantábrica, NO de España)

The first Carboniferous conodonts studied from the Cares river section correspond to fifteen species and one subspecies from seven genera, some of them reported from the Picos de Europa Unit for the first time. The conodonts belong to several zones between the upper Tournaisian and the upper Serpukhovian-lower Bashkirian. Furthermore, the species Idiognathodus incurvus Dunn is reported for the first time in the Cantabrian Mountains and southern Europe, from upper Bashkirian or lower Moscovian beds.     

Conodont chronostratigraphical resolution and Declinognathodus evolution close to the Mid‐Carboniferous Boundary in the Barcaliente Formation type section,NW Spain

The Barcaliente Formation in its type section provides significant conodont occurrences in the Mid‐Carboniferous Boundary interval (between the Mississippian and Pennsylvanian sub‐periods). A sequence of Declinognathodus morphotype appearances (considered here as species) is recognized from the uppermost Serpukhovian to the lower Bashkirian. These morphotypes belong to Declinognathodus noduliferus (Ellison & Graves) sensu lato, the primary marker for the worldwide correlation of the Mid‐Carboniferous Boundary. Among them, D. inaequalis (Higgins) appears to have been the first species to occur at the Mid‐Carboniferous Boundary bed in the Arrow Canyon GSSP. A morphocline between D. bernesgae (Sanz‐López et al.) and D. inaequalis is studied here to identify the bed correlated with the GSSP. Other diagnostic first occurrences of conodonts (Rachistognathus minutus (Higgins & Bouckaert), D. noduliferus, D. lateralis (Higgins & Bouckaert), Idiognathoides asiaticus Nigmadganov & Nemirovskaya and I. corrugatus (Harris & Hollingsworth)) are evaluated as chrono‐markers throughout their worldwide distribution, particularly for the Eurasian faunas. The available secondary markers provide a better resolution and support chrono‐correlation in sections other than that at the GSSP. The worldwide first and late occurrences of studied taxa are considered within the background of the Mid‐Carboniferous Extinction Event and the conodont overturn, where crises progenitor Declinognathodus gave rise to members of the Family Idiognathodontidae. Firstly, progenitor taxa are good biostratigraphic tools, whereas local extinction of genera considered Mississippian in age and new taxa arising later in the post‐crises radiation suggest an early Bashkirian palaeogeography and climatic conditions confining temporarily the biogeographic distribution of conodonts.     

The Guadalupian–Lopingian boundary (Permian) in a pelagic sequence from Panthalassa recognized by integrated conodont and radiolarian biostratigraphy

Guadalupian–Lopingian sedimentary rocks are widely distributed in accretionary complexes in Japan, but the Guadalupian–Lopingian boundary (G–LB) is not well documented from these pelagic sediments. To identify the G–LB and to better correlate an extinction event that occurred around the Guadalupian–Lopingian boundary, we examined the conodont biostratigraphy from a Permian pelagic chert sequence in the Gujo-hachiman section, Gifu, southwest Japan. Age-diagnostic conodonts, including Clarkina postbitteri postbitteri, were found in this section. The biostratigraphic occurrences of these age-diagnostic conodonts can pinpoint the “G–L transitional zone” in the Gujo-hachiman section by comparison with well-studied sections from south China, including the GSSP section. The transitional zone was recognized by the first occurrence horizons of both Clarkina postbitteri hongshuiensis and C. p. postbitteri. The G–LB has been placed at or above the first occurrence horizon of the radiolarians Albaillella yamakitai or Albaillella cavitata in previous studies from China and Japan. We detected the first occurrence horizon of A. yamakitai below the base of the “G–L transitional zone,” in the Upper Capitanian. The conodont biostratigraphy is consistent with the radiolarian biostratigraphy in this section, which can be correlated to relevant sections in China.     

An evaluation of biostratigraphic markers across multiple geological sections in the search for the GSSP of the base of the Serpukhovian Stage (Mississippian)

This paper aims to evaluate potential biostratigraphic markers for the Viséan–Serpukhovian boundary in sections of Europe and Asia, to help identify the base of the global Serpukhovian Stage, which is a high priority task for Carboniferous biostratigraphy. Sections in the Serpukhovian stratotype area in the Moscow Basin contain a gap at the base of the Tarusian Regional Substage (basal in the classical Serpukhovian), so the traditional boundary defined in these sections cannot be precisely correlated with other successions worldwide. The IUGS Task Group to establish a GSSP close to the traditional Viséan–Serpukhovian boundary focused on the search for a new boundary marker, primarily on the first appearance datum (FAD) of the conodont Lochriea ziegleri in the lineage Lochriea nodosa to Lochriea ziegleri, considering it to be a suitable biostratigraphic event. The FOD (first occurrence datum) of L. ziegleri has been recognized in many successions worldwide, although only in a few sections the supposed evolutionary lineage of L. ziegleri was inferred. There are serious impediments to the FAD of L. ziegleri being universally accepted as the boundary marker. This paper presents a review of the FOD levels of L. ziegleri documented so far from multiple sections along with other correlatable markers (foraminifers and ammonoids) that can serve as additional points of reference in sections where a conodont record is poor or absent. The reviewed sections are Naqing Section (South China), Verkhnyaya Kardailovka and Kugarchi sections (South Urals, Russia), Mariinsky Log and Ladeinaya Mountain Sections (western slope of the Middle Urals, Russia), Novogurovsky Section (Moscow Basin, Russia), Vegas de Sotres Section (Cantabrian Mountains, Spain), Lugasnaghta Section (County Leitrim, Ireland), Wenne River Bank Section (Germany), and Milivojevića Kamenjar Section (Družetić, NW Serbia). We also included a compilation of data from sections of North England and southern Scotland. In this paper, we will mainly focus on newly described sections, while the discussion of most previously described sections was summarized by Nikolaeva et al. (2001, 2002, 2005, 2009b) and other publications, so they are only briefly mentioned in this review. It should be added that there is no such a thing as a perfect GSSP section, as each section has certain disadvantages, either lithological, paleontological, or both, so it is important to hear and discuss all the different opinions to develop the optimum strategy for future research. In addition, we analyze published records from several sites in North England and southern Scotland. We discuss the first appearances of the ammonoid genera Cravenoceras, Edmooroceras, Lyrogoniatites, Dombarites, and Platygoniatites, the foraminifers Neoarchaediscus postrugosus, Hemidiscopsis muradymica, H. hemisphaerica, species of Janischewskina and Monotaxinoides, Eostaffella pseudostruvei group, Eostaffellina decurta, and Endothyranopsis plana. We publish here for the first time the useful accounts of foraminifers and conodonts from the Mariinsky Log Section and Ladeinaya Mountain Section (Middle Urals, Russia), and re-figure several important type specimens from Europe and the Urals.     

Conodonts from the Global Stratotype section for the base of the Guzhangian Stage (Cambrian) at Luoyixi,Hunan, South China

The base of the Guzhangian Stage (Cambrian) has been ratified recently at the lowest occurrence of the cosmopolitan agnostoid trilobite Lejopyge laevigata (base of the Lejopyge laevigata Zone) in the Huaqiao Formation in the Luoyixi section, Guzhang County, Hunan Province, China.Sampling of the Luoyixi section in the boundary interval reveals 16 protoconodont and paraconodont taxa. The conodont succession can be correlated with the Gapparodus bisulcatus-Westergaardodina brevidens and the Shandongodus priscus-Hunanognathus tricuspidatus zones of South China and with the Laiwugnathus laiwuensis and the Shandongodus priscus zones of North China. Laiwugnathus laiwuensis occurs immediately below the lowest occurrence of Lejopyge laevigata. The first occurrence of Shandongodus priscus is 14.95 m above the base of the Lejopyge laevigata Zone. A significant change in the conodont associations occurs in the interval corresponding to the uppermost Lejopyge armata trilobite Zone. This interval is characterized by an increase in diversity with the appearance of the paracondont genera Furnishina and Laiwugnathus. Laiwugnathus laiwuensis, which occurs immediately below the lowest occurrence of Lejopyge laevigata, has a short stratigraphic record and allows confident correlation with North China.     

High-resolution biostratigraphy of the Tournaisian-Visean (Carboniferous) boundary interval, Mokrá quarry, Czech Republic

This paper summarizes the results of investigations carried out in the Mokrá quarry since 2006 on the biostratigraphy of the Tournaisian-Visean (T-V) boundary interval. It also integrates previous results obtained by J. Kalvoda and collaborators. The main focus is on the boundary itself, but stratigraphically lower and higher levels have been investigated as well to provide a biostratigraphical context spanning the late Tournaisian to early Visean. This stratigraphical level has been the focus of intense international research in the recent years under the auspices of the Subcommission on Carboniferous Stratigraphy (SCCS) in order to find a new criterion and reference section (Global Stratotype Section and Point, GSSP) for the base of the Visean Stage. The appearance of Eoparastaffella simplex from its ancestor E. “ovalis” and the Pengchong section (Guangxi, southern China) have recently been proposed by the Task Group on the Tournaisian-Visean Boundary and ratified by the SCCS as the new biostratigraphic criterion and GSSP for the base of the Visean, respectively. The sequence exposed in Mokrá is not suitable as a GSSP, notably because it is an active quarry, but it contains most of the foraminifer and conodont guides allowing a high-resolution biostratigraphy of the boundary interval. In addition, it contains abundant trilobites. For these reasons, it constitutes one of the best sections across the T-V boundary in Europe and can serve as a useful additional reference.     

华南贵州罗旬纳水剖面宾夕法尼亚亚系各阶之界线

贵州罗甸纳水剖面是研究宾夕法尼亚亚系牙形刺和各阶界线的良好剖面.经详细研究,从石炭系中间界线至石炭-二叠系界线,一个新的牙形刺序列为Gnathodus bilineatus bollandensis(密西西比亚系顶部),Declinog-nathodus noduliferus, Idiognathoides sulcatus sulcatus, Id. sinuatus, Id. corrugatus-Id. pacificus, Neognatho-dus symmetricus, Idiognathodus primulus-Neognathodus symmetricus, Idiognathodus primulus-Neognathodus bassleri, Idiognathoides sulcatus parva, Streptognathodus expansus, Idiognathoides ouachitensis, Diplognatho-dus coloradoensis, Di. ellesmerensis, Gondolella donbassica-Go, clarki, Idiognathodus podolskensis, Swadelina subexcelsa, Sw. makhlina-Sw, nodocarinata, Idiognathodus sagittalis, Streptognathodus cancellosus, St. graci-lis, St. guizhouensis, Idiognathodus simulator, I. nashuiensis, Streptognathodus firmus, St. tenuialveus, St. wabaunsensis和St. isolatus(二叠系)等带.Declinognathodus noduliferus和Streptognathodus isolatus的首次现分别代表宾夕法尼亚亚系和二叠系之底;Diplognathodus ellesmerensis, Idiognathodus sagittalis和J. simula-tor的首现则可分别代表莫斯科阶(Moscovian),卡西莫夫阶(Kasimovian)和格舍尔阶(Gzhelian)之底界.作者较为详细讨论了巴什基尔阶与莫斯科阶以及卡西莫夫阶与格舍尔阶之界线,并认为以Diplognathodus ellesmerensis的首现作为巴什基尔阶(Bashkirian)与莫斯科阶之界线足目前最合适的标志.Diplognathodus ellesmerensis是一易于识别、分布广泛及层位稳定的牙形刺种,其产出又十分接近巴什基尔阶与莫斯科阶的传统界线,因此是划分此界线的良好标志.作者拟推荐纳水剖面为莫斯科阶底界之候选层型剖面,这一界线详细的工作正在大量采样分析和深入研究中,成果将在以后陆续发表.     

A proposed new global stratotype for Aeronian Stage of the Silurian System: Hlásná Třebaň section,Czech Republic

The current Global Stratotype Section and Point (GSSP ) for the Aeronian Stage (Llandovery Series, Silurian System), on Trefawr track in the Llandovery area of Wales, is an inadequate marker for precise, global, correlation. The International Subcommission on Silurian Stratigraphy has, therefore, undertaken the selection of a new GSSP for this level. The lowest occurrence of the graptolite Demirastrites triangulatus , 1.38 m above the base of the black‐shale succession of the ?elkovice Formation at the Hlásná T?ebaň section in Central Bohemia, is proposed to mark the base of the Aeronian Stage. The section, which fulfils all formal requirements for stratotype of a chronostratigraphical unit, should be considered as a candidate for the new GSSP . An abundant, well‐preserved, diverse graptolite fauna occurs through the section along with common chitinozoans, which indicate that the Spinachitina maennili Biozone spans the boundary interval. The section comprises the lower–middle Aeronian (D. triangulatus–Lituigraptus convolutus graptolite biozones) along with underlying Rhuddanian (Akidograptus ascensus–Coronograptus cyphus biozones) and Hirnantian strata. Several graptolite genera of primary importance in global correlation (Demirastrites , Petalolithus , Rastrites and Campograptus ) first appear in the lower part of the triangulatus Biozone. The structurally simple section is somewhat condensed, but there is a uniform succession of black shale without any evidence of disconformity in the broad boundary interval. The C_org isotope record exhibits a minor positive excursion just above the base of the triangulatus Biozone, whereas TOC and N isotope and elemental geochemical records provide evidence for uninterrupted sedimentation under stable, anoxic conditions.     

Integrated biostratigraphy and carbon isotopes from the Cambrian Tangwangzhai section,North China

The Tangwangzhai section, western Shandong Province, North China, the type section for the Cambrian Kushan and Chaomitian formations, yielded a diverse and relatively well-preserved conodont fauna, in which we recognize the Westergaardodina orygma, Westergaardodina matsushitai, Muellerodus? erectus, and Westergaardodina aff. fossa–Prooneotodus rotundatus zones of the North China conodont zonation. The Tangwangzhai conodont succession can be correlated not only with the polymerid trilobites occurring in the section but also with the conodont zones established for South China. The first occurrence of Furnishina longibasis and Furnishina quadrata in the upper part of the Westergaardodina matsushitai Zone allows the recognition of the base of the Paibian Stage and Furongian Series in the upper part of the Kushan Formation. The base of the Jiangshanian Stage, in the uppermost Muellerodus? erectus Zone, can be recognized by the presence of Westergaardodina cf. calix close to the base of the Chaomitian Formation. Chemostratigraphic analyses of the Tangwangzhai section show the onset of a positive carbon isotope excursion, referred to the SPICE event, in the upper part of the Kushan Formation at a level corresponding to the first occurrence of F. longibasis and F. quadrata. The base of the Jiangshanian Stage in the section is close to the demise of the SPICE positive excursion.     

Lower Bashkirian conodonts from the Iraty Formation in the Alduides-Quinto Real Massif (Pyrenees,Spain)

The stratigraphy of the Carboniferous Iraty Formation is described in several sections from the southwestern Alduides-Quinto Real Massif (western Pyrenees), and biostratigraphy is updated based on conodont occurrences. The Ochaverri Member yields upper Serpukhovian conodonts, whereas the lower part of the overlying Asturreta Member is assigned to the lower Bashkirian, according to the occurrences of Declinognathodus noduliferus bernesgae and D. n. noduliferus, respectively. The upper beds of the formation (Baserdi Member) are correlated with the Kinderscoutian to lowermost Marsdenian English substages (lower Bashkirian), and new conodont occurrences are the oldest species of Neognathodus and Idiognathodus known in Spain. We discuss the status of Idiognathoides macer, several species related to I. sinuatus, I. sulcatus and Lochriea glaber. Conodont diversity is related to stratigraphic distribution of species and conodont biofacies. Finally, the lowermost Marsdenian to upper Bashkirian age of the siliciclastic rocks overlying the Iraty Formation is also discussed and compared with known data from localities in the western Pyrenees.     

Uppermost Devonian–Lower Carboniferous conodonts from the Indert Formation in the Shine Jinst area,South Mongolia

In southwestern Mongolia, conodonts from the Indert Formation at its type locality in Shine Jinst were investigated to improve regional and global correlation of the uppermost Devonian–lowermost Carboniferous. The abundance and diversity of the conodont fauna is quite low in general, whereas representatives of Siphonodella, Protognathodus, Pseudopolygnathus, Polygnathus and Bispathodus have been collected from the Indert-Uul section. Five conodont biointervals, from the Bi. costatus-Pr. kockeli Interregnum (ckI) to the Siphonodella sandbergi Zone, could be recognized from the top Heermorit Member and the lower part of the Shombon Member, while the middle and upper parts of the Shombon Member cannot be zoned based on the conodonts. Absence of index fossils Si. sulcata, Pr. kuehni and Pr. kockeli hampered precise recognition of the Devonian–Carboniferous boundary (DCB) at Indert-Uul, but this transition is well delineated by typical uppermost Famennian and Tournaisian conodont assemblages. The DCB is tentatively placed at the base of the Shombon Member, consistent with the lithological boundary and the resuming of carbonate production during the DCB transitional interval.     

The Carboniferous carbonates of the Dombar Hills (western Kazakhstan) and the problem of the Viséan–Serpukhovian boundary

The Upper Viséan–Serpukhovian ammonoid-rich carbonates in the Dombar Hills (Aktobe Region, western Kazakhstan) provide an excellent opportunity to calibrate the ammonoid and conodont zonations around the base of the Serpukhovian Stage, and are important for interregional correlation. A section in the Dombar Hills spanning the Viséan–Serpukhovian boundary is measured and its fossil content is analyzed. Two ammonoid genozones (Hypergoniatites-Ferganoceras and Uralopronorites-Cravenoceras) and two conodont zones (Lochriea nodosa and Lochriea ziegleri) are recognized. The section displays a contact between the Hypergoniatites-Ferganoceras and Uralopronorites-Cravenoceras ammonoid genozones in a continuous succession and an evolutionarily early appearance of the genus Cravenoceras. The base of the Serpukhovian Stage is drawn at the level of the first appearance datum (FAD) of the conodont Lochriea ziegleri, which, as in the Verkhnyaya Kardailovka section (potential GSSP candidate, South Urals, Russia) enters within the Hypergoniatites-Ferganoceras ammonoid Genozone.     

Multidisciplinary study of the Mechetlino Quarry section (Southern Urals,Russia) — The GSSP candidate for the base of the Kungurian Stage (Lower Permian)

The Mechetlino Quarry section is a potential candidate for the Global Section Stratotype and Point (GSSP) for the base of the Global Kungurian Stage on the International Stratigraphic Chart. In 2017, the Mechetlino Quarry section became the part of the First Geopark in Russia, therefore excavation work was carried out to clear it, a road was laid, and infrastructure was developed. Recent evidence to justify the base Kungurian boundary in the Mechetlino section was obtained. Additional conodonts confirmed the data of V.V. Chernykh on the Artinskian–Kungurian transition. Besides Neostreptognathodus pnevi Kozur and Movshovitsch, the second marker (N. lectulus Chernykh) of the base-Kungurian was established. The Artinskian–Kungurian transition of the Mechetlino Quarry section is characterized by the richest associations of small foraminifers, ammonoids, and other groups. The occurrence of mass Clausiuraloceras above the Kungurian boundary, which changed the representative Artinskian association, was identified. Small foraminifers of the Artinskian–Kungurian transition are numerous and include sufficient species to provide a correlation with Western Tethys, Svalbard and Australia. Data on isotopic ratios of δ¹³C and δ¹⁸O, δ¹³C_org values and the magnetic susceptibility were obtained for the first time.     

Re‐evaluation of the conodont Iapetognathus and implications for the base of the Ordovician System GSSP

Terfelt, F, Bagnoli, G. & Stouge, S. 2011: Re‐evaluation of the conodont Iapetognathus and implications for the base of the Ordovician System GSSP. Lethaia, Vol. 45, pp. 227–237. In 2000, the International Union of Geological Sciences (IUGS) ratified the decision from the International Working Group on the Cambrian–Ordovician Boundary (COBWG) to place the Global boundary Stratotype Section and Point (GSSP) for the base of the Ordovician System in the Green Point section, Newfoundland, Canada, at a point coinciding with the first appearance of the conodont Iapetognathus fluctivagus. However, a restudy of the conodont successions from Green Point shows that this species is not present at the boundary interval, and as a consequence the section does not fulfil the biostratigraphical requirements of a GSSP. The GSSP horizon as now defined is based on a level part‐way through the range of I. preaengensis– a species with lower first appearance datum (FAD). The true FAD of I. fluctivagus is above the FAD of planktonic graptolites and well above the FAD of I. preaengensis. As a consequence of these problems, a restudy of the GSSP section and the other sections in the Cow Head Group is necessary. A redefinition of the GSSP horizon is suggested. The following four alternative horizons have potential as new horizons for the GSSP level: the FAD of Cordylodus intermedius; the FAD of Cordylodus andresi; the FAD of Eoconodontus notchpeakensis; and the FAD of the agnostoid Lotagnostus americanus. □Boundary, Cambrian, conodont, Global boundary Stratotype Section Point, Iapetognathus, Ordovician.     

Conodont biostratigraphy of the Upper Devonian-Lower Carboniferous Shahmirzad section, central Alborz, Iran

The conodont fauna from the Devonian-Carboniferous Shahmirzad section, located in the Central Alborz Mountains (North Iran), have been studied mainly for biostratigraphic purposes. Some levels were barren of conodonts, whereas others yielded a not very abundant, but quite differentiated fauna. No conodonts have been found from the mainly terrigenous and shaly Geirud Formation, whereas representative of genera Bispathodus, Clydagnathus, Gnathodus, Hindeodus, Mehlina, Polygnathus, Protognathodus, Pseudopolygnathus and Siphonodella have been collected from the mainly calcareous overlaying Mobarak Formation. The fauna allowed to discriminate five biointervals, from the sulcata Zone to a “Lower typicus - anchoralis-latus interval” in the central part of the section, while the lower and upper parts cannot be zoned on the basis of conodonts. This paper is the first report on lowermost Carboniferous conodonts from the Mobarak Formation in central Alborz.     

Biostratigraphical analysis of the Moscovian–Kasimovian transition at the neostratotype of Kasimovian Stage (Afanasievo section, Moscow Basin, Russia)

The Afanasievo Quarry, approximately 90 km southeast of Moscow and about 5 km southwest of Voskresensk, starts with Late Moscovian limestone of the Peski Formation, which is overlain by shallow-water carbonates of the Krevyakinian Horizon (Substage) (Suvorovo and Voskresensk formations) and the lower part of the Khamovnikian Horizon (Substage) (Ratmirovo and Neverovo formations). These rocks were formed under the strong influence of glacio-eustatic sea-level fluctuations and are separated by the palaeosol horizons and minor stratigraphic gaps. The Moscovian–Kasimovian transition interval contains fusulinids, brachiopods, bryozoans, corals, and conodonts. It was recently proposed [Villa, E., Task Group, 2005. Report of the Task Group to establish GSSPs at the Moscovian–Kasimovian and Kasimovian-Gzhelian boundaries. Newsletter on Carboniferous Stratigraphy 23, 9–10] that the appearance of the conodont Idiognathodus sagittalis Kozitskaya is one of the best markers for definition of the base of the Kasimovian on the global chronostratigraphic scale. The first appearance of this species is 2 m above the base of Neverovo Formation, and is close to the first appearance of the fusulinid Montiparus in the section. The possible ancestor of I. sagittalis occurs in the lower Suvorovo Formation, but is more abundant and more advanced in the middle to upper parts of the Voskresensk Formation. This lineage has potential for defining a GSSP at the first appearance of I. sagittalis.     

Early evolution of Declinognathodus close to the Mid‐Carboniferous Boundary interval in the Barcaliente type section (Spain)

Upper Serpukhovian to lower Bashkirian conodonts studied from the lower and middle part of the Barcaliente Formation type section (NW Spain) are not abundant, but the sedimentary record seems unusually well represented. The first occurrence of Declinognathodus bernesgae occurs more than 140 m below the first occurrence of D. inaequalis, the taxon of the D. noduliferus species group appearing in the bed of the Global Standard Stratotype‐section and Point (GSSP) for the Mid‐Carboniferous Boundary (Arrow Canyon, Nevada). P1 elements transitional between D. bernesgae and D. inaequalis are described and considered relevant to the global correlation of the Mid‐Carboniferous Boundary. In addition, D. praenoduliferus, D. cf. noduliferus, D. lateralis, Idiognathoides asiaticus, I. corrugatus and Neognathodus sp. are described and illustrated from the Barcaliente Formation. A phylogeny is proposed for the early species of these genera on the basis of the morphological changes of the P1 element, where the rostral parapet and area, the surface roughness and the length and depth of the adcarinal troughs are diagnostic characters at the specific level, while caudal ornamentation is a secondary taxonomic character.     

U–Pb zircon age,sedimentary facies,and sequence stratigraphy of the Devonian–Carboniferous boundary,Daposhang Section,Guizhou, China

The Devonian–Carboniferous (D/C) boundary marks a major extinction event and an important evolutionary turnover in Paleozoic biotas, but understanding D/C boundary events is hampered by a lack of accurate and consistent ages. As a result, although a global stratotype section and point (GSSP) for the D/C boundary was defined in the section La Serre trench E’ (Montagne Noire, France) and accepted and ratified by the IUGS, problems arose concerning both the lithology of the La Serre section and the identification of S. sulcata. Due to these problems, in the very recent years, the ICS decided to reconsider the GSSP. The Daposhang section in Muhua, Guizhou, China, is one of the most important reference sections for the D/C boundary: it is a conformable succession across the D/C boundary that contains conodont faunas characteristic of restricted subtidal facies. A biostratigraphically well constrained tuff near the D/C boundary in the Daposhang section provides an excellent opportunity to date this important event in Paleozoic history. Sensitive high-mass resolution ion microprobe (SHRIMP) U–Pb analyses were conducted on single zircons extracted from the tuff (between beds 0 and 1, which may be temporally equivalent to the Hangenberg black shale) as well as an analysis of sedimentary facies and sequence stratigraphy across the D/C boundary in the Daposhang section. Bed 0 is below the D/C boundary, in the Upper praesulcata conodont zone, and bed 1 is directly above the D/C boundary, in the sulcata conodont zone. Zircon analyses yielded 20 concordant dates that form a cluster with a ²⁰⁶Pb/²³⁸U concordia age of 359.6 ± 1.9 Ma. An abundance of bioclastic packstone and wackestone and a lack of obvious high-energy sedimentary structures or evidence of reworking in layers spanning the D/C boundary suggest conformable deposition in a quiet, restricted subtidal paleoenvironment. Fifty-nine fifth-order cycles combine to depict a shallowing-upward succession (beds 029-01; highstand systems tract of a third-order sequence representing the terminal Devonian). The highstand systems tract is overlain by a deepening-upward succession (beds 0-29; maximum flooding surface and transgressive systems tract of a third-order sequence representing predominantly the basal Carboniferous) that begins at the D/C boundary. Fischer plot analysis of high-frequency cycles at the D/C boundary confirms this well developed sea-level pattern, which, together with the tuff and black shale depositional events, has potential for global correlation. On the basis of this new single-zircon SHRIMP age and previously published work on the biostratigraphy and sequence stratigraphy of the D/C transition interval, the deposition of tuff took place at approximately 359.6 Ma, and the age of the D/C boundary at Daposhang, Guizhou, China, is estimated at 359.58 Ma.     

Evolution of the fusulinid Depratina in the Bashkirian–Moscovian interval

The genera Staffellaeformes and Depratina are widespread in Lower and Middle Pennsylvanian rocks of Eurasia and are important for Pennsylvanian biostratigraphy. The lineage Staffellaeformes–Depratina represents one of several major trends in the fusulinid evolution recorded in the Bashkirian–Moscovian. The genus Depratina first appeared in the late Bashkirian but continued and thrived in the Moscovian. In the Southern Urals, Depratina prisca first appeared close to the base of the Moscovian, and can be used as a marker for this boundary.