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1.
    
Abstract:  Several thousand disarticulated remains together with a few complete enrolled specimens of the lower Cambrian eodiscoid trilobite Calodiscus lobatus ( Hall, 1847 ) have been collected at two outcrop areas in Sweden. The material reveals new details of morphology and morphogenesis during ontogenetic development. Size-frequency analyses show that the material from the Fånån rivulet in Jämtland, central Sweden, represents a natural population dominated by juveniles, whereas the material from Gislövshammar in Scania, southern Sweden, has been sorted during postmortem transport. Three stages of protaspid development can be traced and defined as well as all subsequent ontogenetic stages for the cephalon, hypostome and pygidium. The early meraspid pygidium has a pronounced larval notch, which persists, though becoming progressively less distinct in later meraspides. The number of axial rings in the transitory pygidium increases throughout meraspid development until a third and final thoracic segment is liberated. During ontogeny the articulating half-rings are strongly developed, and both meraspides and holaspides were capable of full sphaeroidal enrollment and outstretched postures. The hypostome undergoes some dramatic modifications; in M0 the anterior margin is axe-shaped, by M1 the area of attachment greatly decreases and the hypostome becomes more elongated and pear-shaped, before attaining its adult form, which has an overall resemblance to that of polymerid trilobites. During ontogeny, the hypostome changes from a conterminant attachment to a natant condition, thereby mirroring hypostomal evolution within trilobites generally. The morphology, ontogeny, enrollment, hypostomal development and the presence of calcified protaspides suggest polymerid rather than agnostoid affinities of the eodiscoids.  相似文献   

2.
    
Abstract: Three thousand seven hundred disarticulated remains together with several articulated specimens of the Cambrian Series 2 ptychopariid trilobite Strenuaeva inflata Ahlberg and Bergström, 1978 have been collected from the Torneträsk area, northern Sweden. The material provides significant new data on the morphology, ontogeny, moulting and enrolment of the species. Two distinct morphotypes, possibly an expression of sexual dimorphism, are recognized. The morph with a pair of bulbs in the frontal area, interpreted as brood pouches, is considered to represent females. Statistical treatment of the length/width ratio in cranidia reveals isometric growth during ontogeny for both morphotypes. The transition from the meraspid to holaspid ontogenetic period has been established through recognition of the successive development of the number of thoracic segments in articulated late meraspides. Throughout its life cycle, Sinflata went through 11 meraspid degrees and at least 17 holaspid growth stages. Inferred moult ensembles and exuviae reveal the successive opening of cephalic sutures and the function of the rostral plate during exuviation. As in other ellipsocephalid trilobites in which enrolment is known, the pygidium and two or three thoracic segments of Sinflata are concealed beneath the cephalon (spiral enrolment) during complete enrolment.  相似文献   

3.
    
Abstract: A large number of complete specimens together with numerous disarticulated sclerites of the eodiscinid trilobite Sinodiscus changyangensis Zhang in Zhou et al., 1977 have been collected from the lower Cambrian Shuijingtuo Formation in Changyang, Hubei Province, South China. An ontogenetic series is established based on the immature and mature exoskeletons including the previously unknown protaspides and meraspides, in particular. No further substages can be differentiated in the protaspid specimens herein. Changes that took place during the meraspid period include the addition of postcephalic segments and prominent pygidial larval notches in early meraspid development which became progressively less distinct and disappeared in degree 2. Two holaspid stages are recognized based on the addition of a new pygidial segment, indicating that the start of the holaspid phase preceded the onset of the epimorphic phase and accordingly, its developmental mode is attributed to the protarthrous pattern. The trunk segmentation schedule of S. changyangensis is discussed, which is similar to other primitive eodiscoid trilobites, that is, as the boundary between the thorax and pygidium migrated posteriorly, there is no change in the number of the trunk segments. The processes of liberation of the thoracic segment and segment insertion into the pygidium are separated from one another, and the two different mechanisms, somitogenesis and tagmosis, progress independently during the ontogenetic development of the postcephalic region of these primitive eodiscinids.  相似文献   

4.
黑龙江伊春早寒武世西伯利亚型动物群   总被引:10,自引:1,他引:10  
记述黑龙江伊春地区早寒武世含多门类小壳化石的三叶虫为主的动物群,并依据东北北部及其紧邻俄罗斯境内的寒武纪地层及动物群特征,论证了东北北部地区的沉积-生物-构造古地理属于西伯利亚构造域,华北与西伯利亚两大构造域的对接主带应在二连浩特-西拉木伦-长春-延吉-符拉迪沃期斯克一线。  相似文献   

5.
Shabaella fengzuensis的蜕壳   总被引:3,自引:1,他引:3  
描述了浙江江山早寒武世三叶虫Shabaella fengzuensis的蜕壳方式,指出Shabaella的蜕壳机制与同时代的Redlichia有别,但与Hunanocephalus,Sinolenus等的蜕壳方式可能接近。  相似文献   

6.
中国寒武纪三叶虫生物地理趋议   总被引:1,自引:0,他引:1  
寒武纪三叶虫除北疆和藏南尚无可靠报道外,在中国各地均有广泛分布。三叶虫证据表明,除兴安区的额尔古纳一兴安地体和早寒武世中天山一北山地体显示了与西伯利亚和劳伦陆块在动物群方面的密切相关外,中国寒武纪的所有板块和大部分地体(包括中及晚寒武世的中天山~北山地体)均系当时东泛冈瓦纳的重要组成部分,在生物地理上彼此密切关联。对这些属于东泛冈瓦纳的中国区块的生物地理格局的进一步探讨,主要侧重于它们台地相浅水三叶虫动物群之间的对比,特别是立足于来自动物群纪录比较完整的华南、塔里木和华北板块的证据。综合分析研究表明这些板块和有关的地体在中及晚寒武世应归属同一生物地理区,而在早寒武世则可划分为两个生物地理亚区:一个由华南、塔里木板块和安南或印支地体组成,另一仅以华北板块为代表。  相似文献   

7.
    
There were multiple origins of metamorphosis‐undergoing protaspides in trilobite evolution: within the superfamilies Remopleuridioidea, Trinucleoidea, and within the Order Asaphida. Recent studies have revealed that the protaspides of the Cambrian representatives of the Remopleuridioidea and the Trinucleoidea did not undergo metamorphosis. However, ontogeny of the Cambrian members of the Order Asaphida has remained unknown. This study documents the ontogeny of the Furongian asaphoidean ceratopygid trilobite, Proceratopyge cf. P. lata Whitehouse, from northern Victoria Land, Antarctica. Two stages for the protaspid phase, five developmental stages for the post‐protaspid cranidia, and ten stages for the post‐protaspid pygidia have been identified. Interestingly, the protaspis directly developed into a meraspis without metamorphosis. A new cladistic analysis resulted in a single most parsimonious tree, according to which the presence of the bulbous commutavi protaspis turns out to be a synapomorphy for Asaphidae + Cyclopygoidea, not a synapomorphy for the Order Asaphida as previously suggested. In addition, it is inferred that there was convergent evolution of indirectly‐developing commutavi protaspides during the Furongian and Early Ordovician. Metamorphosis‐entailing planktonic larvae evolved in many different metazoan lineages near the Cambrian–Ordovician transition, due to the escalating ecological pressure of the Great Ordovician Biodiversification Event. Since the bulbous commutavi protaspid morphology is thought to be an adaptation for a planktonic life mode, the convergent evolution of the indirect development in the three trilobite lineages at this period might have been a result of adaptation to the early phase of the Great Ordovician Biodiversification Event.  相似文献   

8.
新材料表明定远虫(Tingyuania)和双岛虫(Dinesus)这两个三叶虫属足有效的,前者并不是后者的晚出异名.双岛虫类的地质延限被下延至传统早寒武世晚期的清虚洞组.补允描述首次发现的定远虫的尾部,并描述双岛虫的一新种.  相似文献   

9.
林天瑞 《古生物学报》2002,41(3):428-433
描述采用自南澳大利亚Officer盆地东部Manya-6号钻井的下寒武统Ouldburra组的Abadiella三叶虫的一新种-Abadiella officerensis,同时指出Parabadiella是Abadiella的同义名。  相似文献   

10.
塔里木盆地塔中台地奥陶系海百合茎化石丰富。本文通过研究塔中35井上奥陶统良里塔格组和桑塔木组岩芯中的海百合与其共生生物的关系, 讨论了海百合化石的古生态。在塔中35井上奥陶统取芯中识别出7种海百合茎类型: 圆圆茎、星星茎、星圆茎、椭圆圆茎、星椭圆茎、圆角星星茎和不规则星茎, 根据茎板形态划分为五射茎板组(Pentameri)、椭圆茎板组(Elliptici)、圆茎板组(Cyclici)、特殊茎板组(Varii)四种组合类型; 统计了海百合茎化石各种切面的大小和百分含量; 结合岩性分析了每段地层的沉积环境。研究发现, 在局限泻湖环境下, 蓝细菌–四分珊瑚为主的生物组合中不见海百合化石碎片; 仅见腹足类和介壳类化石的细砂屑滩中, 偶见细小的海百合茎化石; 在开阔海环境中, 海百合广泛与苔藓虫、管孔藻、粗枝藻等共生。在海百合–苔藓虫组合中, 海百合茎板的类型最丰富多样, 五射茎板组、椭圆茎板组、圆茎板组、特殊茎板组等四种茎板组合类型都存在; 在管孔藻–海百合组合中, 海百合茎板主要为圆茎板类型, 粗大、密集, 海百合茎碎片含量最高; 在海百合–粗枝藻组合中, 海百合茎板的数量减少、大小变小, 但茎板的类型比较丰富, 四种茎板组合类型都有。由此认为, 塔中台地上奥陶统沉积期, 海百合生长的首选环境为开阔台地, 并受水动力能量和水体清洁度控制: 水动力能量高, 水体清洁, 海百合繁盛; 水动力能量低, 水体动荡浑浊, 海百合可以生存; 水动力能量低、较为动荡、微生物席发育的环境, 不利于海百合生存。  相似文献   

11.
    
The Global Standard Stratotype-section and Point (GSSP) of the Furongian Series (uppermost series of the Cambrian System) and the Paibian Stage (lowermost stage of the Furongian Series), has been recently defined and ratified by the International Union of Geological Sciences (IUGS). The boundary stratotype is 369 metres above the base of the Huaqiao Formation in the Paibi section, northwestern Hunan Province, China. This point coincides with the first appearance of the cosmopolitan agnostoid trilobite Glyptagnostus reticulatus, and occurs near the base of a large positive carbon isotopic excursion (SPICE excursion).  相似文献   

12.
    
Abstract: Cambrian trilobites mainly lived on the sea floor, and up till now few, if any, unequivocally planktonic trilobites have been reported from earlier than the Ordovician. The late Cambrian (Furongian) to late Ordovician olenids are a distinctive group of benthic (sea‐floor dwelling) or nekto‐benthic trilobites. Here we show, however, that one recently described, miniaturized and very spiny olenid species, Ctenopyge ceciliae must have been planktonic (passively drifting or feebly swimming in the upper waters of the sea). This interpretation is based not only upon body form but also on the analysis of its visual system and may be one of the earliest records of the planktonic realm being invaded by trilobites.  相似文献   

13.
    
Abstract:  The ontogeny of Pseudopetigurus deprati Turvey et al ., 2006 from the Ordovician Dawan Formation (Arenigian), Anhui Province, South China, is described. The presence of an anterior cranidial border in Pseudopetigurus is recognized for the first time. On account of the short (tr.), spindle-shaped anterior border, and the distinct, steeply inclined, wall-like pygidial margin, Pseudopetigurus is assigned to the family Dimeropygidae, the first genus of this family endemic to Gondwana. It has previously been regarded as a member of family Isocolidae. Cladistic analysis of species assigned to Dimeropygidae supports the monophyly of a clade including Dimeropyge , Dimeropygiella , Ischyrotoma , Pseudohystricurus and Pseudopetigurus , which may represent a subfamily Dimeropyginae. The distribution of Dimeropyginae shows a predominantly palaeotropical distribution, while the Gondwana Pseudopetigurus must have diverged from the dimeropygine common ancestor in pre-Arenigian time.  相似文献   

14.
    
Polymeroid trilobite fauna from two stratigraphic sections (Godbondar and Kuhbanan) of the Kuhbanan Formation in northern Kerman (central Iran) were studied and subjected to biostratigraphic analysis. Eleven genera and species are recognised from the latest Early Cambrian and Middle Cambrian Peri-Gondwanian successions of the study sections. The recognised fauna includes Afghanocare lategenatum, Blountia blountia, Iranoleesia sp., Iranoleesia pisiformis, Kermanella kuhbananensis, Kermanella lata lata, Kermanella lata minuta, Kermanella sp., Redlichia chinensis, Redlichia noetlingi and Redlichia sp. Based on trilobite distribution, three trilobite biozones were recognised in the study sections, namely Redlichia noetlingi biozone, Kermanella kuhbananensis biozone and Iranoleesia pisiformis biozone. The age of the study sections is late Early Cambrian to late Middle Cambrian based on the recognised trilobite biozones. The recognised late Early Cambrian trilobite assemblages (especially Redlichia and Kermanella) from northern Kerman are similar to those found from some other parts of Gondwana or Peri-Gondwana terrains (north India, Pakistan, northwestern Kashmir, Tajikistan, South Australia, South China and Afghanistan) and show affinities with fauna found in some other parts of Iran (Alborz, northern Iran; Tabas, eastern Iran; southeast Karman).  相似文献   

15.
The morphogenesis of the early members of the genus Peronopsis (Trilobita, Agnostina) is studied. The pygidium of Agnostina exhibits growth-related changes more clearly than the cephalon. In Peronopsis inarmata Hutchinson, 1962 and P. brighamensis Resser, 1938 the axial furrow develops around the anteroglabella in meraspid degree 0. The most significant changes occur in the shape of the pygidium axis and the postaxial field; posterolateral spines do not change throughout growth. Specific characters continue to develop until the advanced holaspid stage. Within the genus Peronopsis, three distinct growth patterns of the pygidium are recognized. Different relative growth rates of different parts of the pygidium were responsible for the evolutionary transitions between the growth patterns. The growth pattern when the pygidium lacks a postaxial furrow was the most primitive and gave rise to a number of species with a long axis reaching the border, a growth pattern that in turn gave rise to the species with the postaxial furrow (a character typical of Agnostina). The evolution of these growth patterns is inferred from ontogenetic observations and is supported by the stratigraphic succession of Peronopsis and their immediate descendants in different regional faunas.  相似文献   

16.
    
Abstract:  Ontogenies are described for the first time for three species of Early Cambrian oryctocephalid trilobites: the oryctocephaline Changaspis elongata Lee, in Chien and the oryctocarines Duyunaspis duyunensis Chang and Chien, in Zhou et al . and Balangia balangensis Chien from the Balang Formation in eastern Ghuizhou Province, south-west China. The complete protaspid to holaspid ontogeny for Duyunaspis duyunensis and incomplete meraspid to holaspid ontogenies for Changaspis elongata and Balangia balangensis are described. The relative degree of ontogenetic variation within the lineage Duyunaspis duyunensis – Balangia balangensis is analysed and indicates that Balangia may have evolved from Duyunaspis by paedomorphosis. Decrease in thoracic segment number and concomitant increase in pygidial segment number in the later genus mirrors the pattern seen in the evolution of another oryctocephalid genus, Arthricocephalus . In both cases, selection was probably targeting an increase in cephalic convexity. This resulted in the evolution of a larger pygidium due to increased elevation of the axis above the substrate. The ontogenetic development of Changaspis provides evidence supporting the view that the Oryctocarinae may be considered as paedomorphic descendants of the Oryctocephalinae.  相似文献   

17.
    
Several hundred specimens of a tiny olenid trilobite, Ctenopyge ceciliae sp. nov., have been found in stinkstone nodules in the upper Cambrian Peltura scarabaeoides Zone in southern Sweden. This exceptionally spinose form is known only from disarticulated specimens, but is quite well preserved, and all growth stages are represented. The early ontogenetic stages are exceptionally small, the protaspis being only half the size of that of the associated Peltura species. There may have been no more than three thoracic segments. Thus the whole ontogeny was compressed, and this together with the very small size of the adult indicates a true miniaturisation. Whereas the likely control of the origin of the tiny C. ceciliae was basically progenesis, the extreme spinosity had a different origin; allometric growth or possibly peramorphosis. C. ceciliae is small enough for the spines to have appreciably retarded sinking through frictional effects, and this small trilobite is interpreted as a free-swimming or floating form.  相似文献   

18.
Juvenile development of the Middle Cambrian agnostid trilobite Pentagnostus proanabarensis Fedoseev, 1999 is studied. A sequence of eight juvenile stages is defined based on changes in morphology and measurements. Three of them belong to meraspid degree 0, another three refer to meraspid degree 1, and two represent early holaspid stages. Neither the frequency distribution of the length of cephalons and pygidia nor the length-width scatter diagrams give clear clusters of molts; the number of juvenile molts can be estimated through the analysis of the posteroaxis length distribution. Thus, the measurements and morphology of some juvenile pygidia with vestiges of the pleural and postaxial furrows allow reconstruction of the number of segments during individual growth. At the initial stage of development, the pygidial axis of P. proanabarensis probably had four segments, the pygidial axis increased up to no less than six segments in adults.  相似文献   

19.
This study is focused on the growth pattern of agnostids, a group of organisms that were very abundant in the Cambrian. The developmental patterns of six agnostid species are compared. It is shown that, like other trilobites, agnostids have two developmental stages, i.e., meraspid degrees 0 and 1 and the holaspid stage. Meraspid degrees are named according to the number of released thoracic segments. Several molts took place during every meraspid stage resulting in successive instars with a unique set of features. Meraspid degree 0 includes two or three instars, the first and last of which have a distinctive morphology and have been encountered in most of the species studied. During meraspid degree 1, the animal molted from two to four times. The number of molt stages appears to be species-specific. The first and last instars are easily distinguished from other developmental forms. Early holaspids of the majority of agnostid species studied also have certain features in common.  相似文献   

20.
Duyunaspis Zhang and Qian in Zhou et al.,1977是黔东、湘西北杷榔组掘头虫类三叶虫的重要分子。本文描述剑河八郎\"清虚洞组\"的Duyunaspis,这些新材料的头鞍中部膨大且向前略有收缩,前颊类型面线,成虫具10个胸节;而杷榔组的Duyunaspis duyunensis Zhang and Qian in Zhou et al.,1977头鞍桶状,背沟较平直,后颊类型面线,成虫10个胸节,二者明显不同。本文将\"清虚洞组\"的Duyunaspis定为一个新种:剑河都匀盾壳虫Duyunaspis jianheensis sp.nov.。研究显示,D.jianheensis与杷榔组的D.duyunensis具有形态相似性,后者可能为其祖先种。  相似文献   

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