Spatial distribution patterns of calcareous nannofossils across the Callovian–Oxfordian transition in the French Subalpine Basin

Callovian–Oxfordian calcareous nannofossils are documented from four sections distributed along a proximal–distal (slope to central basin) transect in the French Subalpine Basin (south-eastern France). The sections were dated and correlated using integrated ammonite, dinoflagellate and nannofossil biostratigraphies. The nannofossil assemblages are constantly dominated by Watznaueria britannica. However, significant spatial changes in nannofossil assemblage composition are recorded. Nannofossil assemblages recovered in slope and slope-to-basin transition sections are characterized by high abundances, lowest diversities, highest percentages of smallest-sized morphotypes of W. britannica, subordinated W. britannica/manivitiae (a morphotype described in this study), and are indicative of turbulent, unstable and/or eutrophic conditions. Towards the central part of the basin, as the nutrient content and turbulence declined and paleoenvironmental stability increased, the nannofossil assemblages become more diverse and characterized by lower relative abundances of W. britannica but higher contribution of larger morphotypes and higher relative abundances of W. barnesiae/fossacincta, Biscutum dubium and Zeugrhabdotus erectus. These latter small coccoliths, markers of high surface-waters fertility for the Cretaceous, were probably not competitive with respect to smaller-sized W. britannica in unstable and eutrophic surface waters for the Jurassic. An increase in the surface water productivity is recorded both in proximal and distal sections at the Callovian–Oxfordian transition, and is demonstrated both by an increase in nannofossil total abundances and in the relative abundances of taxa adapted to high-trophic levels. Thermal minimum in surface waters, as indicated by an increase in the percentages of the cool-water nannofossil taxa, is associated to increased productivity. Further studies will be necessary to demonstrate whether surface-water productivity changes, already recorded in the eastern Paris Basin, were global.     

Stable isotope analyses of belemnites from the Kachchh Basin,western India: paleoclimatic implications for the Middle to Late Jurassic transition

Stable isotope analyses of 61 diagenetically unaltered belemnite rostra from the Middle to Late Jurassic of the Kachchh Basin of western India suggest stable paleotemperatures across the Callovian-Oxfordian boundary (~14°C). Only at the end of the Middle Oxfordian water temperatures drop for more than 3°C before reaching again higher values during the Kimmeridgian (~12.3°C). The data do not support polar glaciations proposed for the Middle to Late Jurassic transition, which necessarily would have led to a global temperature and sea-level minimum at the boundary. Callovian to Oxfordian rocks in the Kachchh Basin point to a gradual shallowing corresponding to a slight fall in relative sea level. However, the magnitude of this regression is comparatively small, and the sea-level minimum is reached in the late Early Oxfordian and not close to the boundary. Results from the Kachchh Basin therefore, imply almost stable climatic conditions during the Middle to Late Jurassic transition and do not show any evidence for polar glaciations.     

Pliensbachian (Early Jurassic) calcareous nannofossils from the Peniche section (Lusitanian Basin,Portugal): A clue for palaeoenvironmental reconstructions

Quantitative analysis of Early Jurassic calcareous nannofossil assemblages from the Peniche section in Portugal have been performed in order to interpret palaeoenvironmental changes occurring in the Lusitanian Basin during the Late Pliensbachian Davoei, Margaritatus and Spinatum Ammonite Zones. Nannofossil data are compared to already published carbon and oxygen stable isotope data, organic matter content (wt.%Total Organic Carbon, TOC), and biomarker analysis. A significant change in calcareous nannofossil assemblages and species diversity at the transition between the Margaritatus and Spinatum Ammonite Zones matches with the pattern shown by geochemical data. This suggests that a profound change in environmental conditions occurred at that time. In the Davoei and Margaritatus Ammonite Zones, in a context of general sea-level rise, the Lusitanian Basin was characterized by water column stratification that favoured the sedimentation and preservation of organic matter. Biomarkers and oxygen isotope trends suggest that stratification of water masses occurred because of low salinity in surface waters. The shallowest part of the water column, characterized by oligotrophic conditions, was inhabited by the (probable) calcareous dinocyst Schizosphaerella spp., while the deep-dweller Crepidolithus crassus flourished in the lower photic-zone layers. In the Spinatum Ammonite Zone, a regressive trend occurred and a salinity increase is inferred on the basis of oxygen isotope values. Water masses were probably less stratified at that time. The upper photic-zone nannofossil assemblages were still dominated by Schizosphaerella spp. whilst, in the deep photic-zone, Mitrolithus jansae (a Mediterranean taxon) replaced C. crassus (a taxon with NW-European affinity). This pattern may indicate a change in palaeoceanographic conditions related to surface current circulation. The sea-level fall occurring during the Spinatum Ammonite Zone may have resulted in the partial isolation of the Lusitanian Basin from the NW-Europe basins because of the creation of a threshold. Alternatively, tectonic drowning of North African carbonate platforms in the Late Pliensbachian may have promoted better exchanges of nannoplankton between the Lusitanian Basin and the Mediterranean Tethys via the creation of new seaway connections.     

Episodic sedimentation on a peri-Tethyan ridge through the Middle–Late Jurassic transition (Villány Mountains, southern Hungary)

The Villány area, as a central part of the Tisza microcontinent/terrane along the European margin of Tethys, was characterized by intense subsidence in the Early and Middle Triassic, followed by a long interruption of subsidence in the Late Triassic to Middle Jurassic. During the Middle–Late Jurassic transition, marine sedimentation started with three distinct sedimentary episodes dated as Late Bathonian, Early Callovian, and Middle–Late Callovian, respectively. The succession is terminated by a thick limestone of Middle Oxfordian age. The sedimentary features, microfacies, and macroinvertebrate associations of these four stratigraphic units are documented and illustrated. The Middle to Late Jurassic sedimentary episodes of the Villány succession record an interplay of local and global factors and paleogeographical changes. At the beginning, local tectonic movements governed the main features of sedimentation, though the role of eustasy was also essential. From the mid-Callovian onwards, global climatic, biotic, and paleoceanographical changes controlled the nature and formation of the local carbonate sediments. The Callovian stromatolites are attributed to the activity of sulphate-reducing bacteria in a deep sublittoral, current-swept environment. Upwelling of eutrophic Tethyan waters is recorded by the prevalence of the Bositra filament microfacies in the Callovian. The long submarine hiatus at around the Callovian–Oxfordian transition mirrors a serious restriction of the carbonate budget, due to sudden cooling and a change in the oceanic current system (opening of a circumglobal Tethyan Passage), and to a higher amount of dissolved CO₂. In the Middle Oxfordian, the carbonate production considerably increased in accordance with the sudden global warming.     

Paleoenvironmental controls on the morphology and abundance of the coccolith Watznaueria britannica (Late Jurassic,southern Germany)

The coccolithophore species Watznaueria britannica is dominant in Middle-Upper Jurassic calcareous nannofossil assemblages and presents morphological variation, including different coccolith size, shape and length of the central area and of the bridge. Six morphotypes can be recognized in the polarizing light microscope. The aim of this work is to better understand the morphological variability of W. britannica and determine if this variability is controlled by paleoecological factors. In order to investigate the potential paleoecological controls on W. britannica morphology and abundance, we carried out a biometric study on a restricted temporal interval: the Late Oxfordian, in the Swabian Alb (southern Germany), characterized by increasing carbonate production linked to climatic changes. The Balingen–Tieringen section, where previous works on sedimentology, nannofossil assemblage composition, and δ¹⁸O and δ¹³C analyses were performed, was selected for this study. The variations in morphology and abundances of W. britannica were studied on 40 samples of the Balingen–Tieringen section, presenting variable lithologies and calcium carbonate contents. For each level, seven biometric parameters (coccolith length, width and ellipticity, central area length, width and ellipticity and central area proportion with respect to the coccolith) were measured or calculated on digitally captured images of the first 100 W. britannica coccoliths observed in the light microscope. The relationships between the different biometric variables were described using bivariate and Principal Component Analyses. Biometric parameters and Principal Component factors extracted from nannofossil assemblages as well as other paleoenvironmental proxies, were investigated using regression, and their stratigraphic trends were compared. Principal component analysis of the six biometric variables (3938 measurements) on W. britannica coccoliths shows a reduced morphological variability compared to a significant size gradient. An allometric trend recognized on the total placolith and on the central area within the W. britannica assemblages suggests that the different morphotypes may represent intra-specific variability rather than different species. The general trend through Late Oxfordian shows an increase in size of W. britannica coccoliths, mainly driven by an increase in the contribution of the large morphotypes. Increasing placolith size is associated with drier and warmer climatic conditions during the latest Oxfordian.     

Contribution of calcareous nannoplankton to carbonate deposition: a new approach applied to the Lower Jurassic of central Italy

The Upper Pliensbachian–Lower Toarcian Somma section (Central Italy), which displays alternation of limestone–marl lithotypes, was investigated for its calcareous nannofossil and CaCO₃ contents. In order to evaluate the contribution of calcareous phytoplankton to the carbonate fraction of rocks a new method was applied combining a recently developed technique for nannofossil absolute quantification and measurements of nannofossil size. Our study suggests that nannofossils did not produce the bulk of carbonate mud in the studied section, although they contribute to different extents to the pelagic carbonate fraction. The contribution of both coccoliths and (mainly) the Incertae sedis Schizosphaerella to the carbonate fraction was important only in periods of reduced accumulation rates and/or during clay deposition. The highest estimated carbonate production by nannofossils in the studied section is around 83% of the total carbonate content, but in a sample containing only 12.8 wt% CaCO₃. The observed limestone–marl alternations therefore result from cyclical export of carbonate mud from the shallow Latium–Abruzzi carbonate platform to the Umbria–Marche Basin and, only subordinately, from changes in nannoplankton productivity.     

Calcareous nannofossil changes during the late Callovian–early Oxfordian cooling phase

Calcareous nannofossil quantitative and biostratigraphic analyses integrated with carbon and oxygen stable isotopes were carried out on the core ANDRA (Agence Nationale pour la gestion des Déchets Radio-Actifs—FRANCE) HTM 102 across the Callovian/Oxfordian boundary drilled at Cirfontaines-en-Ornois, Départment de Haute-Marne, eastern France. Calcareous nannofossil assemblages at the Callovian–Oxfordian transition are dominated by the genus Watznaueria. An increase in abundance of Biscutum spp. and A-group, which consists of Axopodorhabdus spp. (A. atavus, A. rahla, and A. cylindratus), Podorhabdus grassei, Octopodorhabdus decussatus, Hexapodorhabdus cuvillieri (family Axopodorhabdaceae), and Triscutum spp., correlates with a significant positive excursion in δ¹⁸O suggesting that these groups were probably adapted to cooler surface waters. A positive increase in δ¹³C values is coupled with high abundances of eutrophic taxa such as Zeughrabdotus erectus, Biscutum spp., and small-sized Watznaueria britannica, and a decrease in abundance of the big and oligotrophic taxa Schizosphaerella punctulata and Watznaueria manivitae. Climate cooling across the Callovian/Oxfordian boundary probably triggered a breakdown in stratification of surface waters leading to more intense nutrient recycling and higher primary productivity that favoured the shift in abundance of small-sized eutrophic taxa in the East Paris Basin.     

An eastern Tethyan (Tibetan) record of the Early Jurassic (Toarcian) mass extinction event

A record of the Early Jurassic mass extinction event is reported from eastern Tethyan (Tibetan) locations for the first time. In the Mount Everest region a thick Lower Jurassic carbonate formation, here named the Yungjia Formation, is developed within the predominantly clastic Triassic–Jurassic succession. Within the formation a sharp transition from peloidal packstones/grainstones to thin‐bedded, pyritic micrite‐shales interbeds records a sharp pulse of deepening and development of dysoxic bottom waters. Both the lithiotid bivalves and the lituolid foraminifera are important constituents of the lower Yungjia Formation but they disappear at this flooding surface or a short distance below it. This extinction event is comparable to that seen at the base of the Pliensbachian/Toarcian boundary in western Tethyan platform carbonates but the Tibetan events occurred late in the Toarcian Stage as indicated by nannofossil biostratigraphy and C isotope chemostratigraphy. The Early Jurassic extinction event (and the associated spread of oxygen‐poor waters) was therefore not synchronous throughout the Tethyan region.     

Calcareous nannofossils as paleoproductivity indicators in Upper Cretaceous organic-rich sequences in Israel

Abundant and diverse calcareous nannofossil assemblages were found in organic-rich carbonate sequences which accumulated in a Campanian-Maastrichtian upwelling belt along the southeastern Tethys. The sequences studied represent the inner (shallower) and the outer (deeper) parts of the upwelling belt. The paleoenvironmental significance of selected nannofossil taxa and their utility in productivity reconstruction was established by comparing their distribution to foraminifera and dinocyst-based productivity profiles.Based on the calcareous nannofossil assemblages, a high-productivity group and a low-productivity group of species were determined. The distribution of these groups agrees well with the dinocyst- and foraminifera-based productivity curves and, hence, can be used to record paleoproductivity changes.The ratio between the high-productivity and low-productivity nannofossil groups, the Nannofossil Index of Productivity (NIP), is proposed here as a productivity proxy that can be utilized in reconstructing basinal productivity development.A quantitative analysis of the nannofossil assemblages indicates that their abundance and diversity increase towards the open sea, in the outer and less productive part of the upwelling belt. In the inner and more productive part of the upwelling belt, the nannofossils assemblages become less abundant and less diverse relative to those of the open marine environment.Micula decussata and Watznaueria barnesae are common to abundant in most samples. M. decussata becomes more abundant in poorly-preserved samples. On the other hand, the distribution of W. barnesae matches better with intermediate productivity levels.     

Faunal turnover of marine tetrapods during the Jurassic–Cretaceous transition

Marine and terrestrial animals show a mosaic of lineage extinctions and diversifications during the Jurassic–Cretaceous transition. However, despite its potential importance in shaping animal evolution, few palaeontological studies have focussed on this interval and the possible climate and biotic drivers of its faunal turnover. In consequence evolutionary patterns in most groups are poorly understood. We use a new, large morphological dataset to examine patterns of lineage diversity and disparity (variety of form) in the marine tetrapod clade Plesiosauria, and compare these patterns with those of other organisms. Although seven plesiosaurian lineages have been hypothesised as crossing the Jurassic–Cretaceous boundary, our most parsimonious topology suggests the number was only three. The robust recovery of a novel group including most Cretaceous plesiosauroids (Xenopsaria, new clade) is instrumental in this result. Substantial plesiosaurian turnover occurred during the Jurassic–Cretaceous boundary interval, including the loss of substantial pliosaurid, and cryptoclidid diversity and disparity, followed by the radiation of Xenopsaria during the Early Cretaceous. Possible physical drivers of this turnover include climatic fluctuations that influenced oceanic productivity and diversity: Late Jurassic climates were characterised by widespread global monsoonal conditions and increased nutrient flux into the opening Atlantic‐Tethys, resulting in eutrophication and a highly productive, but taxonomically depauperate, plankton. Latest Jurassic and Early Cretaceous climates were more arid, resulting in oligotrophic ocean conditions and high taxonomic diversity of radiolarians, calcareous nannoplankton and possibly ammonoids. However, the observation of discordant extinction patterns in other marine tetrapod groups such as ichthyosaurs and marine crocodylomorphs suggests that clade‐specific factors may have been more important than overarching extrinsic drivers of faunal turnover during the Jurassic–Cretaceous boundary interval.     

Palynostratigraphie du Callovien-Oxfordien (Jurassique) dans les Terres noires, sud-est France

The Terre noires are monotonous sedimentary deposits comprising thick, dark marls and thin carbonaceous layers. They constitute a lithostratigraphical group known in the major part of the SE French Basin. In the East of this basin this group, dated from late Bajocian to mid Oxfordian, is well developed; on the other hand on the West, it is reduced until it disappears locally on the border of the Massif Central. In the East, the succession is precisely dated by ammonites, whereas in the west, ammonites are not consistently present, and do not permit the same precision. Analysis of dinoflagellate cyst assemblages, at the Callovian-Oxfordian boundary, show that it is possible to correlate the eastern to the western succession. Two marker species allow the correlation of four outcrop sections. The first index, Wanaea fimbriata, is well known; its first appearance datum is exactly at the Callovian-Oxfordian limit in the Boreal and Mediterranean realms. It is the index taxon of the palynological parazone Wfi. The second index, Stephanelytron ceto, an endemic taxon in the SE French, has its last appearance datum within the Scarburghense horizon, the second subzone of the oldest zone of the Oxfordian (Mariae zone). The first section studied was sampled in the east part of the basin (Hautes Alpes department). It corresponds to the maxima of marine deposits and is used as the palynostratigraphical reference. The three other sections analyzed, sampled in the west part of the basin (Ardèche department) where marine deposits are more condensed, are correlated and dated due to dinoflagellate cyst markers observed in the reference section. These results indicate that dinoflagellate cysts are a reliable correlation tool. Batiacasphaera rugosa (Courtinat, 1980) nov. comb is proposed.     

Berriasian–Aptian black shales of the Maiolica formation (Lombardian Basin, Southern Alps, Northern Italy): local to global events

The Lower Cretaceous Maiolica Formation represents the classic pelagic succession of the Apulian margin of the Tethys Ocean, in the Southern Alps. It consists of white calcilutites with minor marly layers and black shales. A multidisciplinary study including lithostratigraphy, nannofossil assemblage composition and organic geochemistry was performed on two composite sections in order to characterise the marl–shale layers that record production and storage of organic matter (OM) and terrigenous input in the Berriasian–Aptian interval. Such layers are associated with turbidite deposition and bottom current activity in the Late Berriasian–earliest Valanginian, Late Valanginian and Early Aptian. The uppermost Hauterivian–Barremian black shales alternate with pelagic calcilutites, forming an interval barren of redeposited layers. The quantity of organic carbon increases from the Berriasian to the Barremian; the background composition of the kerogen is continental-dominated, with varying proportions of marine-derived OM. Preservation of OM did not necessarily occur under anoxic conditions; on the contrary, storage of OM probably resulted from enhanced terrestrial input of nutrients, siliciclastics, woody and herbaceous fragments, inducing mesotrophic conditions and increased primary productivity. Higher fertility of surface waters during the Late Valanginian and Early Aptian is reflected by distinctive decreases in abundance of oligotrophic nannoconids and increase in abundance of mesotrophic Diazomatholithus in the Valanginian. During the Barremian, the rhythmic establishment of eutrophic conditions, documented by the highest amounts of marine OM, determined high-frequency fluctuations in nannoconid abundance. The Early Aptian nannoconid crisis is interpreted as the result of a major climate change and enhanced primary productivity. Variations of calcareous nannofloras, as well as changes of lithology and OM, which we describe in the Lower Cretaceous Maiolica Formation of the Lombardian Basin, can also be recognised in coeval units from other sites of the Apulian side of the Tethys. Similar episodes of siliciclastic and terrestrial OM accumulation and major changes in calcareous nannofossils are also recorded in the Upper Valanginian and Lower Aptian pelagic sediments of the Tethys, Atlantic and, partly, Pacific oceans. Such nannofossil variations are coeval with positive excursions of the δ¹³C curve and suggest perturbations of the bio-geochemical cycles at a global scale. Additional minor changes in siliciclastic and terrestrial OM accumulation as well as in nannofloras are probably the result of local events, strictly related to the physiography and evolution of the Tethyan margin.     

Palynofacies and TOC analysis of marine and non-marine sediments across the Middle–Upper Jurassic boundary in the central-northern Lusitanian Basin (Portugal)

The Middle–Upper Jurassic transition is a geodynamic benchmark in the evolutionary history of several peri-Atlantic basins. Contrary to the vast Tethyan and peri-Tethyan areas, in the Lusitanian Basin (Portugal), this interval corresponds to a major basin-wide disconformity preceded by a complex forced regression that induced sharp facies variations across the depositional systems. The Middle Jurassic units below are fully marine, whereas the Upper Jurassic sediments (Lower?-Middle Oxfordian), correspond to freshwater/brackish lacustrine, grading into punctuated brackish-restricted lagoonal and shallow-marine paleoenvironments. This study presents total organic carbon and palynofacies data of 34 samples (total analyzed thickness about 85 m) collected from three key sections encompassing the Middle–Upper Jurassic transition in the central-northern sector of the Lusitanian Basin. The palynofacies of the analyzed part of Middle Jurassic units (Cabo Mondego Formation) are characteristic of marine environments at their base, evidencing upwards a regressive trend of the depositional systems. Total organic carbon content is generally low, reaching up to 1.94 wt%. The Upper Jurassic Cabaços Formation presents kerogen assemblages mostly of continental origin, although punctuated by minor intervals of marine influence. Total organic carbon content is more variable, reaching up to 30.56 wt%. Intraclasts of re-deposited fragments of microbial mats were found incorporated in the kerogen assemblages, which point to highly dynamic erosional and depositional processes. Diversity of Botryococcus sp. occurrences was confirmed as an indicator of the degree of paleoenvironmental stability. The vertical distribution and comparison of the kerogen assemblages of the different sections indicate major changes of these parameters among relatively close settings and along narrow vertical intervals, attesting to the high sedimentary dynamics observed in the Lusitanian Basin.     

Reconstructing the palaeoenvironment of the Middle Russian Sea during the Middle–Late Jurassic transition using stable isotope ratios of cephalopod shells and variations in faunal assemblages

Oxygen and carbon isotope data of well-preserved belemnite rostra and ammonite shells are presented from the Callovian–Oxfordian boundary (uppermost Lamberti to lowermost Cordatum zones) of the Dubki section near Saratov in the Russian Platform. Palaeotemperatures calculated for nektobenthic belemnites (averages of 5 °C and 8 °C for cylindroteuthids and belemnopseids, respectively) show the presence of cold bottom waters in the central part of the Middle Russian Sea during the studied interval. Palaeotemperatures calculated for ammonites, which are assumed to have lived in near-surface waters, are considerably higher (average 13 °C). The presented data show a vertical thermal gradient in the Middle Russian Sea. The belemnite oxygen isotope record and the relative abundances of ammonite families in the Dubki section do not correlate with each other probably as a result of different depth habitats of ammonites and belemnites. A review of literature isotope data shows the climatic zonation in European seas at the Middle–Late Jurassic transition. Despite the flux of cold polar waters to the Middle Russian Sea and the area of Scotland there is no evidence for glaciation at the Middle–Late Jurassic transition. Changes in water circulation during a sea-level highstand were likely a source of spreads of cold bottom waters and cardioceratid ammonite fauna in this time period.The belemnite isotope record of the Callovian–Oxfordian boundary in the Russian Platform is characterized by significant scatter of δ¹³C values. No temporal carbon isotope trend is observed. The δ¹³C values of Russian belemnite rostra average 2.6‰ VPDB being 1 to 2‰ higher than the values of coeval Lower Oxfordian belemnites from the area of the Submediterranean ammonite province. Higher (than Submediterranean) δ¹³C values of Russian belemnite rostra are likely related to high biologic productivity and/or high organic matter burial in semi-isolated Boreal–Subboreal marine basins.     

Earliest Zanclean age for the Colombacci and uppermost Di Tetto formations of the “latest Messinian” northern Apennines: New palaeoenvironmental data from the Maccarone section (Marche Province, Italy)

The occurrence of planktonic foraminifers in the latest Messinian deposits (uppermost Di Tetto Formation and Colombacci Formation) of the Marche Province (Apennine foredeep, Italy) has stimulated a debate since the 1970s. An earlier palynological study of the entire Maccarone section revealed a pronounced, and a sudden increasing frequency of saccate pollen grains which indicates more distal conditions, and thus a transgression. At first attributed to tectonic activity, this transgression is now interpreted as representing the Zanclean marine transgression after the discovery of Ceratolithusacutus, the calcareous nannofossil marker of the earliest Zanclean in the Mediterranean Sea. Evidence from marine dinoflagellate cysts and planktonic foraminifers supports this result. The Colombacci Formation and uppermost part of the Di Tetto Formation (i.e. the entire p-ev2 stratigraphic unit) belong to the earliest Zanclean. The so-called Lago Mare no longer has a regional chronostratigraphic sense, and should be understood as the invasion of Paratethyan organisms via surface waters owing to a connection at high sea-level between the Aegean Sea and the Eastern Paratethys (Dacic Basin). A new robust environmental reconstruction of the northern Apennine foredeep is proposed, which respectively considers the effects of tectonics and Mediterranean eustasy.     

Lutetian calcareous nannofossil events in the Agost section (Spain): implications toward a revision of the Middle Eocene biomagnetostratigraphy

This paper presents a detailed calcareous nannofossil biostratigraphy of the entire Lutetian of the Agost section (Betic Cordillera, SE Spain). This investigation integrates and improves on previous study performed through the Ypresian/Lutetian boundary by the authors on this succession. The new revision of the integrated bio‐magnetobiochronology of the Early/Middle Eocene interval revealed highly diversified calcareous nannofossil assemblages, characterizing more than 8 Myr of climatic variability. The studied interval spans from Zone CP11 to Subzone CP14a and from the upper part of Zone NP13 to the base of Zone NP16 of calcareous nannofossil standard zonations. The revision of the calcareous nannofossil content enabled the identification of numerous secondary events which greatly improved the stratigraphic resolution of this time interval. An important re‐organization of the nannoflora was observed during the Y/L transition, when Reticulofenestra and Dictyococcites (Noelaerhabdaceae) became the most important genera in terms of abundance and dispersal, dominating the Middle Eocene nannofossil assemblages and replacing Toweius and Discoaster taxa characteristic of the lower Eocene. Pentaliths and Blackites experience a great expansion and diversification, whereas Discoaster and Chiasmolithus which are well diversified but never abundant during the Lutetian show a slow turnover. A reassessment of the major bio‐events observed in the Noelaerhabdaceae family as well as revision and correlation of these events with the classical Italian sections (Contessa and Bottaccione) are presented. The new results show that biostratigraphic problems related to the Middle Eocene chronology are not limited to the correlation between calcareous nannofossils and planktonic foraminiferans at the Y/L transition but extend to calcareous nannofossil events commonly used for correlating the Bartonian.     

Spatial distribution of Late Pliensbachian (Early Jurassic) calcareous nannofossils within the Lusitanian basin (Portugal)

Quantitative analyses of Pliensbachian calcareous nannofossils have been carried out on a proximal-distal transect in the Lusitanian Basin (Portugal). The studied sections Vale Venteiro near Tomar and Peniche represent proximal and distal environments with respect to the emerged land of the Iberian Meseta (to the East). The upper portion of the Vale das Fontes Fm (Davoei and Margaritatus ammonite Zones) is studied in both sections and correlated by means of ammonite and nannofossil integrated biostratigraphy. A careful analysis of the preservation state of nannofossils is performed. Preservation state is moderate to good in the two settings; changes in nannofossil assemblages are therefore considered as primary. Samples were analysed for nannofossil absolute and relative abundances, species diversity and wt%CaCO₃. The analysis of calcareous nannofossils (absolute abundance, percentage, average abundance) shows changes in the community structure in space (i.e., from proximal to distal), and vertically within the water column. This pattern suggests a partitioning of habitats within the photic zone, and with respect to emerged lands. Proximal environments within the Lusitanian Basin were probably more eutrophic, because of the proximity of emerged lands to the East (Iberian Meseta). This was the source area for nutrients delivered into the basin by river transport. Distal environments were likely characterized by a greater water depth and by a deeper light penetration leading to a relatively more expanded photic zone. The assemblages recorded in the proximal setting exhibit high mean relative abundance of placolith-bearing coccolithophorids (Lotharingius spp., Biscutum spp. and Similiscutum spp.) and of Schizosphaerella spp., while distal assemblages are dominated by Crepidolithus crassus and Schizospharella spp. Crepidolithus crassus is interpreted as a deep-dweller coccolithophorid, inhabiting preferentially distal and deeper settings in the Lusitanian Basin. Placolith-bearing coccolithophorids were more abundant in proximal settings with respect to emerged lands, under relatively elevated trophic conditions. The probable calcareous dinocyst Schizospharella spp. proliferated in surface waters of both proximal and distal environments.     

The record of mid Cretaceous oceanic anoxic events from the Ionian zone of southern Albania

Along the Sopoti section of Mali Gjere mountain are found two distinct mid-Cretaceous marly-shaly-siliceous intervals that occur near the top of the Vigla Limestone Formation (Aptian-Albian) of the Ionian zone in southern Albania. The lower interval is relatively rich in carbonate content (36% CaCO₃ on average) and radiolaria (10% on average). It contains black shale levels rich in organic matter (up to 8.5% total organic carbon (TOC)) of marine origin, which did not experience any strong thermal maturation as suggested by their very low T_max values. The age of the lower interval is latest Barremian to Early Aptian based on its calcareous nannofossil and radiolarian assemblages. It is therefore regarded as the equivalent of the Fourcade Level of Greece, reflecting the Oceanic Anoxic Event 1a (OAE1a). The upper interval is richer in both carbonate content (60% CaCO₃ on average) and radiolaria (20% on average) but it is practically devoid of any preserved organic matter. Its age straddles the Aptian-Albian boundary based on integrated biochronologic data of dinoflagellates, calcareous nannofossils and radiolaria. It is tentatively considered as the sedimentary expression of OAE1b (sensu Leckie et al., 2002) in the Ionian zone of Albania. The presence of large Assipetra nannoliths in both shaly-siliceous intervals and the relative abundance of radiolaria suggest that their accumulation took place during periods of higher productivity in the Ionian zone of Albania.     

Microbial carbonates and corals on the marginal French Jura platform (Late Oxfordian,Molinges section)

Molinges was located on an Upper Jurassic ramp system of low-energy regime that developed at the southern margin of the French Jura platform. The sedimentary succession is characterized by the transition from a mixed siliciclastic-carbonate to a carbonate depositional setting that occurred during a long-term shallowing-upward trend. The disappearance of siliciclastics is explained by a climatic change, from humid and cold to drier and warmer conditions, previously identified in Late Oxfordian adjacent basins. The base of the section shows marl-limestone alternations of outer ramp. In its middle part, the section displays oncolitic marls, coral-microbialite beds and oncolitic limestones that deposited in a mid ramp position. Finally, the upper section part is made of oolitic limestones of inner ramp. In outer- to mid-ramp settings submitted to terrigenous inputs, the stacking pattern of deposits and facies evolution allow the identification of elementary, small-, medium-, and large-scale sequences. Small amplitudes of sea-level variations probably controlled rapid shifts of facies belts and reef window occurrences. In small-scale sequences, the coral beds developed during periods of sea-level rise. The decreasing rate of sea-level rise is marked by the downramp shift of the oncolitic limestone belt that led to the demise of coral-microbialite beds. These bioconstructions are mainly represented by thin biostromes in which corals never reach great sizes. The coral assemblages mainly include the genera Enallhelia, Dimorpharaea, Thamnasteria, and some solitary forms (Montlivaltia and Epistreptophyllum). They suggest relatively low-mesotrophic conditions in marine waters during the edification of the primary framework. Relatively cold water temperatures and periods of more elevated nutrient contents are probably responsible of the reduced coral development and the formation of a large amount of microbialites.     

Changes in calcareous nannofossil assemblages during the Mid-Pleistocene Revolution

The distribution of calcareous nannofossils are quantitatively analysed in high-resolution deep-sea core samples from North Atlantic DSDP Site 607 and Eastern Mediterranean ODP Site 967, between marine oxygen isotope stage (MIS) 35 and 15, in order to observe the response of calcareous nannofossils to the Mid-Pleistocene Revolution (MPR). Nannofossil abundance patterns show major variations through MIS 25–22 and distinct higher amplitude changes from MIS 21 upward, sometime clearly correlatable to glacial–interglacial cycles. High amplitude, short-term, fluctuations of small placoliths and F. profunda characterize the interval through MIS 24 to 22/21, suggesting strong instability in nutricline dynamics and surface water productivity. Principal Component Analysis and Shannon–Weaver indices (diversity and dominance) highlight significant shifts, mainly related to variations in the characteristics of surface waters. In both sites a minimum in diversity is recorded at MIS 22/21, corresponding to a maximum in the dominance of Reticulofenestra spp. at Site 607 and of Pseudoemiliania lacunosa at Site 967. Above MIS 21 the increase in abundance of various taxa (C. leptoporus, Umbilicosphaera spp., Syracosphaera spp., Rhabdosphaera spp., Oolithotus spp.) and a general increase in diversity are related to a distinct trend toward more stable and oligotrophic surface waters in contrast to the higher productivity and less stratified surface waters of the Early Pleistocene.