Palaeobiogeographic implications of the first report of the eocrinoid genus Ascocystites Barrande (Echinodermata,Blastozoa) in the Upper Ordovician of the Ougarta Range (Western Algeria)

Several specimens of the genus Ascocystites (Blastozoa, Eocrinoidea) are described for the first time in Late Ordovician deposits (Bou M’Haoud Formation) from the Ougarta Range, Algeria. This genus was previously known in Darriwilian–Sandbian deposits of four other areas of the Mediterranean Province (Czech Republic, France, Morocco and Portugal). The Algerian material completes its palaeobiogeographic distribution in the peri-Gondwanan area, restricted in shallow water settings.     

La macrofaune du Pléistocène supérieur d’El Harhoura 2 (Témara, Maroc) : données préliminaires

The El Harhoura 2 cave is situated in the commune of El Harhoura in the Temara region (Morocco). The entrance faces west, towards the ocean, around 200 m away from the current shoreline and about 16 m above sea level. The stratigraphy has currently 11 levels numbered from top to bottom. Level 1 is attributed to the early Neolithic. The archaeological material from level 2 places it within the Upper Palaeolithic (Iberomaurusian). Levels 3 to 11 are assigned to the Middle Palaeolithic (Aterian). Since 1977, a surface of 37 m² has been excavated, yielding particularly abundant remains of a large mammal fauna study in this paper.     

Évolution des environnements littoraux du bassin de Melilla-Nador (Nord-Est Maroc) au Messinien supérieur entre 6,0 et 5,77 Ma

This study of late Messinian deposits (E2 = TCC) from Melilla-Nador carbonate complex (Morocco) allowed us to characterize the environmental evolution between 6.0 and 5.77 Ma, which is a period considered as contemporaneous of the onset and development of the “messinian salinity crisis”. A statistical approach coupled together with classical petrographic study allows us to constrain the sedimentary dynamic of platforms and to propose deposition environment models. The Melilla-Nador platforms E2 deposits show a progressive continuous evolution of littoral environments in two major stages of available space creation and infill. The first stage corresponds to the build up of a reef complex between 6.0 and 5.85 Ma then the second stage, between 5.85 and 5.77 Ma, corresponds to the development of a mixed oolitic shallow ramp during the progressive infill of Melilla-Nador basin. The comparison with the Sorbas basin (Spain) TCC deposits exhibits a very close evolution of late Messinian littoral environments, which could be used as a correlation tool.     

Les grands mammifères du Villafranchien supérieur des travertins du Bassin de Denizli (Sud-Ouest Anatolie,Turquie)

The large mammals from travertine deposits in the Denizli basin include the following species: Archidiskodon meridionalis meridionalis, Equus cf. altidens s. l., E. cf. apolloniensis, Stephanorhinus cf. etruscus, Metacervoceros rhenanus, Cervalces (Libralces) ex gr. minor-gallicus, Palaeotragus sp., Bovinae gen. and sp. indet. This association resembles those from the late Villafranchian of Southern and Eastern Europe, and, to some extent, fromWestern Asia, and could be older than 1.2 Ma.     

Phylloceratoidea du Jurassique moyen et supérieur du Nord-Est de l’Iran (Monts Binalud)

Phylloceratidae are often the most frequent taxa within the Middle and Late Jurassic ammonite faunas of North Iran but so far they have not been noticed much within the existing literature and have been described only sporadically. In this study, the Phylloceratidae of north Iran are treated systematically and described comprehensively. These include 16 taxa, most of them being previously unpublished for Iran.     

Enregistrement des fluctuations du niveau marin dans l’Holocène supérieur du lac Retba (Sénégal) par les foraminifères

A study of sea level fluctuation records based on foraminiferal assemblages was carried out for the first time in a sediment core taken at the Lac Retba edge. A total of 37 foraminifera species were collected along the core, among which most are known in the modern estuaries and lagoons of Senegal. A succession of four assemblages dominated by Ammonia parkinsoniana and A. tepida were identified; they correspond to different stages of evolution of the lake. The first association, at the base of the core, indicates a lagoon slightly opened to the sea and bordered with vegetation, under relatively humid climate. The second has the richest and most diversified microfauna with a high proportion of coastal, marine benthic and planktonic species (about 10%) that indicate a small coastal gulf bordered with mangroves. The third association is oligospecific and typical of a closed and hypersaline lagoon under a dry climate. The last association again contains coastal, marine benthic and scarce planktonic species (3%) indicating a change to a saltier and more open lagoon under dry climate. The disappearance of planktonic foraminifera at the top of the core indicates the closure of the lagoon. The biocenotic indicators are evidence for two marine intrusions that are referred to the Dakarian (3000 years B.P.) and Saint-Louisian (2140-680 years B.P.) stages of the Upper Holocene of the senegalo-mauritanian stratigraphic scale. Evidence of the Lac Retba closure since 680 years B. P. appears in the core by high reduction of foraminifers’ abundance and diversity absence of planktonic species, and salts precipitated at the bottom of the lake.     

Sols et structures d’habitat du Paléolithique supérieur, nouvelles données depuis les Cantabres : la Galerie Inférieure de La Garma (Cantabrie, Espagne)

A brief review is done of the history of the research on Upper Paleolithic floors and structures, with a general outline of the main substantive and theoretical-methodological aspects. On the resulting “state of the art” we present the preliminary results of the study of the occupation evidences from the Lower Gallery of La Garma. In essence, it constitutes an astonishing ensemble of extensive Middle Magdalenian occupation floors, exceptionally rich and quite well preserved, including several structures and associated with rock art paintings and carvings of that period. The potential of the site is superlative, and the research’s project already begins to offer some interesting results of which this article is a fairly small advance.     

Un nouveau Bovini dans la faune du Pléistocène supérieur d’Algérie

During the study of the Quaternary Maghrebian bovines in 1985, the review of the Bovini (aurochs and buffalos) found in Upper Pleistocene sites of Algeria showed that in comparison with most of the African bovine assemblage the attribution of genus Pelorovis to the species antiquus was inappropriate. The biometric and cladistic data revealed a close kinship with genus Syncerus. Later, other authors’conclusions confirmed its relevance. The antique buffalo of the Aterian site of the Phacochoeri has all the characteristics of genus Syncerus, but is different in that it shows a certain number of specific characteristics described in this article.     

Zaozer'e, un nouveau site du Paléolithique supérieur ancien dans le nord-est de l'Europe. Rapport préliminaire

This paper presents preliminary archaeological and chronostratigraphical data obtained during pilot excavations of one of the northernmost European Early Upper Palaeolithic sites, Zaozer'e, situated on the western slope of the Middle Urals.     

Foraminifères benthiques complexes du Bathonien supérieur au Kimméridgien inférieur de la marge atlantique marocaine (« Bassin » d’Agadir) : stratigraphie et paléobiogéographie

The carbonate sediments of the Agadir Basin (Morocco) deposited during the Upper Bathonian-Lower Kimmeridgian p.p. interval are rich in agglutinated complex benthic foraminifera. The detailed inventory of several field-sections allows to identify and to describe these benthonic foraminifera, some of them being recorded for the first time in Morocco. They belong to the main genera Archaeosepta, Andersenolina, Praekurnubia, Pseudocyclammina, Everticyclammina, Alveosepta, Kurnubia, Parurgonina, Neokilianina, Labyrinthina and Rectocyclammina. The stratigraphic and paleobiogeographic distribution of these foraminifera seem to be related to the geodynamic events occurring on the Moroccan Atlantic margin during this time-interval. The stratigraphic distribution of the principal species is generally controlled by the occurrence of ammonites, brachiopods and of some foraminifera which are considered as good stratigraphic markers.     

L’industrie du site néandertalien de la grotte de Vindija (Croatie) : une révision des matières premières lithiques

The new analysis of the Middle Paleolithic industry from the cave Vindija (Croatia), showed a necessity of revision of the previous obtained data especially considering the use and origin of the raw materials (Kurtanjek and Marci, 1990). The results presented in this study pointed out some new aspects of interpretation. First, the significance of quartz in the Middle Paleolithic was underestimated. At the same time it is obvious that the major change in the use of the raw material was at the beginning of the Upper Paleolithic and of the Epigravettian, especially in the use of chert. The second change is registered in the use of chert between Middle and Upper Paleolithic: the chalky nodules of chert are more frequent in the Upper Paleolithic then the river pebbles used in the production of Middle Paleolithic artefacts. Still, the origin of this new raw material is at the moment unknown.     

Évolution du trait de côte à l’Holocène supérieur dans la Sebkha El-Guettiate de Skhira (Golfe de Gabès, Tunisie) à travers sa faune d’ostracodes et de foraminifères

A 122 cm-long core was taken in the El-Guettiate Sebkha of Skhira (southeastern part of Tunisia) in order to investigate the recent palaeoenvironmental evolution of this region. The quantitative and qualitative analysis of ostracod and benthic foraminifera assemblages coupled with a correspondence analysis allows the reconstruction of palaeoenvironmental changes during the Holocene in this area. Four typical associations of ostracods (open marine, coastal marine, lagoonal and estuarine brackish) and two associations of benthic foraminifera (coastal and lagoonal) were distinguished. The onset of restricted lagoonal environments linked to the building-up of sand spit led to the onset of restricted lagoon and brackish environment at cal. 5408 years BP. The Shannon and equitability index of diversity were used to decipher the structural variations of the populations of ostracods and benthic foraminifera along the sampled core. We note a reduction in the Shannon index from the bottom to the top, which indicates a progressive isolation of the biotope. The open lagoonal episode is characterized by high values of diversity. During restricted lagoonal episodes the Shannon and equitability index are reduced. The correspondence analysis reveals an environmental gradient related to the marine influence. It shows an antagonism between the widely opened estuarian lagoonal species and those of restricted lagoon. The less opened estuarian lagoonal taxa occupy an intermediate position. Based on these evidences, microfauna carried out in the El-Guettiate Sebkha allows us to recognize four phases beginning with a widely opened estuarian lagoon (ca. cal. 7460 years PB), followed by a restricted lagoon (ca. cal. 5408 years BP) and finally a brackish lagoon evolving towards the present-day sebkha environment. The opened estuarine lagoon is characterized by high values of species richness and diversity indices.     

Le bassin de l’homme du Pléistocène supérieur de Liujiang, Sud de la Chine : taille du corps, forme du corps et encéphalisation

The Late Pleistocene hominin fossil assemblage from Liujiang, South China include a fairly well-preserved cranium, a right os coxa, a complete sacrum, and other postcranial elements all belonging to a single individual. This rare discovery offers us a unique and singular opportunity in understanding this Late Pleistocene hominin's body proportion and relative cranial capacity (encephalization quotient [EQ]), and also pelvic morphology. Using the available right innominate and its mirror-imaged left side, we reconstruct Liujiang hominin's pelvis. Our analysis of the pelvis indicates that the Liujiang hominin has a very gracile and modern-like pelvic morphology. Indeed, all of the pelvic dimensions are smaller than those of other Pleistocene hominins. Moreover, the pelvic characteristics typical of Middle and Late Pleistocene hominins including Neanderthals cannot be identified in the Liujiang pelvis. In contrast, both Liujiang's metric and non-metric features indicate affinities to more recent human populations including our modern Chinese collections from Guangxi of south China. Further support of this assessment comes from the EQ value of 5.754 for Liujiang which is closer to Minatogawa 2 and modern Chinese populations than to Middle and Late Pleistocene fossil hominins. Our analysis of body shape shows that Liujiang has body proportion (i.e. body height relative to body breadth) typical of warm-adapted populations. Based on these findings, we reason that the modern physical characteristics of Liujiang may allude to a more recent geological age. Alternatively, its morphological “modernity” could also point to a much higher degree of skeletal variation within Late Pleistocene hominins in East Asia.     

Échinodermes de l'Ordovicien supérieur (Ashgill) de Sardaigne et d'Algérie

Research on the influence of sea level variations on the benthic faunas have been carried out in the Upper Ordovician of Sardinia. Study of the depositional facies and sequence analysis of the upper part (Lower Ashgill) of the Portixeddu Formation led to the identification of the sedimentary environments. Cystoids and crinoids are associated to bryozoans and brachiopods in most levels. The numerical analysis of associations and megaguilds shows that crinoids and cystoids have a higher frequency in the proximal and median facies of the upper offshore. The columnal association characterized by Conspectocrinus celticus and the coronoid Mespilocystites tregarvanicus has been discovered in the upper part of the formation. This material and complementary samples from Upper Ordovician of Sardinia and Kabylia (Algeria) bring additional data on the systematic and show the wide distribution of this fauna outside of the Ibero-armorican domain. The distribution of this echinoderm association supports a palaeogeographical position of the Ibero-armorican domain and Sardinia within the north gondwanan margin during the Lower Palaeozoic.     

L’industrie aurignacienne de la grotte de Šandalja II (Istrie, Croatie) dans le contexte de la région de l’Est de l’Adriatique

This paper presents the results obtained by lithic analysis of Early Upper Palaeolithic levels in Šandalja II Cave, Istria, Croatia. Technological and typological analysis of stone and typological analysis of bone artefacts have been carried out. Production of flakes in Aurignacian levels (G, F, E and E/F) is dominant, but blade and bladelets production is also present. Blades and bladelets were produced by direct soft hammer technique. Aurignacian people of Šandalja II produced their debitage mainly on local grey chert, which is often patinated. Šandalja II is one of the rare—if not the only—site with Aurignacian industry in eastern Adriatic region. The main reason for absence of such industry at other sites can be seen in scarce population or abandonment of some parts of this region.     

Signification paléoécologique des assemblages de diatomées du Messinien dans le bassin de Melilla-Nador (Rif Nord-Oriental, Maroc)

In the Melilla-Nador basin, the diatomites represent one of the main sedimentary components of the Messinian together with reefal carbonate platforms. Two vertical sections were realised in the diatomitic levels from distal platform. Diatom assemblages are for the first time described in this basin. A total of 125 diatom species were determined: 73 centrics and 52 pennates. Significant diatom assemblages defined from the statistical analysis indicate ecological variations concerning diatomitic levels or parts of diatomitic levels. Using the data sets we propose paleoenvironmental models showing the complexity of the ecological interactions. This study point the permanence of opened marine environment, the impulse of cold waters from Atlantic origin, a general decrease in bathymetry in the Messinian marls-diatomitic series, the influence of the carbonate platform progradation and the occurrence of upwelling systems.     

La Grotte des Félins, site paléolithique du Pléistocène supérieur à Dar Bouazza (Maroc)

In the continental sediments filling a cave that developed in Middle Pleistocene deposits near Casablanca, Morocco, a very rich mammal fauna from the last climatic cycle was discovered. It was associated with a Middle Palaeolithic poor and atypical lithic assemblage manufactured on blocks and pebbles. The lower fossiliferous layer contained abundant carbonized remains of felids. Unfortunately, the site was destroyed by quarrying before adequate excavations could be carried out.     

Le Paléolithique moyen et supérieur de la Hongrie du nord : nouvelles investigations dans la région d’Eger

The region of Eger town at the foot of the Bükk Mountains in Northeastern Hungary has particular importance for the understanding the Middle and Upper Palaeolithic periods of the Carpathian Basin. One of the key-problems is the association of carinated tools and foliate points in the lithic assemblages of the sites. New investigations at the Egerszalók-K?vágó site give fundamental data for the animated discussion on the relation between Aurignacian and Szeletian.     

Nouvelles données sur le genre Imerites Rouchadzé, 1933 (Ammonitida,Ancyloceratina) ; réponses et précisions apportées au travail de Bert,Delanoy et Canut, 2009

This work is mainly an answer to the work of Bert, Delanoy and Canut, on the genus Imerites Rouchadzé, and we add also some points to the knowledge of this genus. An answer is given for the implication of one of the authors (J.V.) of this present work, about the origin of the genus Imerites Rouchadzé, and some significant bibliographic references, not named in the 2009 work, are specified and placed in their context. The characteristics of the ornamental stage “Pseudoshasticrioceras” mainly defined by an oval and compressed section of the whorls and by regular ventrolateral clavi, well marked, which frame the venter in a typical way (Bert et al., p. 181), don’t allow to affirm the presence of this stage during the ontogeny of all species of the genus Imerites Rouchadzé, contrary to what Bert, Delanoy and Canut (p. 30, 31) wrote. The type-species of the genus Imerites Rouchadzé is made clear by the application of the rules of the I.C.Z.N. code. The article 23, with the paragraph 23.1 of the I.C.Z.N. code, imply the senior synonymy of Imerites cristatus (Orbigny) on Imerites giraudi (Kilian), even if the Kilian's species is still the type species of the genus Imerites Rouchadzé. The taxinomical validity of the species Imerites cristatus (Orbigny) is confirmed. Its morphological variability is clarified by the distinction of four referential morphotypes, Cristatus, Giraudi, Favrei and Raricostatum. Consequently, the assumption of a shape-dimensional dimorphism into the populations of the genus Imerites Rouchadzé is rejected, awaiting irrefutable proof. A new species, Imerites stephaniae sp. nov., is described. It was collected in the Gerhardtia sartousiana Zone, in the uppermost part of the Gerhardtia provincialis Subzone, and it is a probable ancestor of the cogeneric species of the Imerites giraudi Zone.     

Succession des peuplements d’échinides du Crétacé inférieur dans le Haut Atlas atlantique (Maroc) ; révision systématique et intérêt stratigraphique

The rich echinoid fauna from the Lower Cretaceous of the Agadir, Haha and Essaouira basins has been revised taxonomically, based on new and abundant collections. Forty-seven species have been identified, of which 20 are reported for the first time from this region and a further three represent new species left in open nomenclature. Their stratigraphic ranges have been determined accurately by reference to the regional ammonite zonation. However, it is also possible to use a local biostratigraphical scheme based on the echinoid succession, where ammonites are rare. The Toxasteridae, which are very abundant and underwent rapid evolution, are the most useful biostratigraphically. The genus Toxaster appears in the Lower Berriasian and continues until the Gargasian passing through the following succession: T. africanus, T. granosus, T. kiliani, T. lorioli, T. obtusus, T. exilis, T. retusus, T. maurus, T. peroni, T. collegnoi. The presence of Eoholaster, Macraster, Douvillaster, Palhemiaster, Heteraster and Pygopyrina is signaled for the first time in the Atlantic High Atlas region.