Weather-Related Indices of Autumn– Winter Dabbling Duck Abundance in Middle North America

ABSTRACT Research on effects of key weather stimuli influencing waterfowl migration during autumn and winter is limited. We investigated relationships between changes in relative abundances of mallard (Anas platyrhynchos) and other dabbling ducks (Anas spp.) and weather variables at midlatitude locations in North America. We used waterfowl survey data from Missouri Conservation Areas and temperature and snow cover data from the Historical Climatology Network to evaluate competing models to explain changes in relative abundance of ducks in Missouri, USA, during autumn-winter, 1995–2005. We found that a cumulative weather severity index model (CumulativeWSI; calculated as mean daily temp - degrees C + no. of consecutive days with mean temp ≤ 0° C + snow depth + no. of consecutive days with snow cover) had the greatest weight of evidence in explaining changes in relative abundance of ducks. We concluded the CumulativeWSI reflected current and cumulative effects of ambient temperatures on energy expenditure by ducks, and snow cover and wetland icing, on food availability for ducks. The CumulativeWSI may be useful in determining potential changes in autumn-winter distributions of North American waterfowl given different climate change projections and associated changes in habitat conservation needs. Future investigations should address interactions between CumulativeWSI and landscape habitat quality, regional waterfowl populations, hunter harvest, and other anthropogenic influences to increase understanding of waterfowl migration during autumn-winter.     

Waterfowl distribution and productivity in the Prairie Pothole Region of Canada: tools for conservation planning

Species conservation requires an understanding of the factors and interactions affecting species distribution and behavior, habitat availability and use, and corresponding vital rates at multiple temporal and spatial scales. Opportunities to investigate these relationships across broad geographic regions are rare. We combined long-term waterfowl population surveys, and studies of habitat use and breeding success, to develop models that identify and incorporate these interactions for upland-nesting waterfowl in the Prairie Pothole Region (PPR) of Canada. Specifically, we used data from the annual Waterfowl Breeding Population and Habitat Survey (1961–2009) at the survey segment level and associated habitat covariates to model and map the long-term average duck density across the Canadian PPR. We analyzed nest location and fate data from approximately 25,000 duck nests found during 3 multi-year nesting studies (1994–2011) to model factors associated with nest survival and habitat selection through the nesting season for the 5 most common upland nesting duck species: mallard (Anas platyrhynchos), gadwall (Mareca strepera), blue-winged teal (Spatula discors), northern shoveler (Spatula clypeata), and northern pintail (Anas acuta). Duck density was highly variable across the Canadian PPR, reflecting positive responses to local wetland area and count, and amounts of cropland and grassland, a regional positive response to latitude, and a negative response to local amounts of tree cover. Nest survival was affected by temporal and spatial variables at multiple scales. Specifically, nest survival demonstrated interactive effects among species, nest initiation date, and nesting cover type and was influenced by relative annual wetness, population density, and surrounding landscape composition at landscape scales, and broad geographic gradients (east-west and north-south). Likewise, species-specific probability of nest habitat selection was influenced by timing of nest initiation, population density, relative annual wetness, herbaceous cover, and tree cover in the surrounding landscape, and location within the Canadian PPR. We combined these models, with estimates of breeding effort (nesting, renesting, and nest attempts) from existing literature, in a stochastic conservation planning model that estimates nest distribution and success given spatiotemporal variation in duck density, habitat availability, and influential covariates. We demonstrate the use of this model by examining various conservation planning scenarios. These models allow estimation of local, landscape, and regional influence of conservation investments and other landscape changes on the productivity of breeding duck populations across the PPR of Canada. These models lay the groundwork for the incorporation of conservation delivery costs for full return-on-investment analyses and scenario analyses of climate, habitat, and land use change in regional and continental population models.     

Wetland Selection by Mallard Broods in Canada's Prairie-Parklands

ABSTRACT Although brood survival has a pronounced effect on population growth in mallards (Anas platyrhynchos), knowledge of brood ecology is more limited than for other vital rates. During 1993–1997 we collected wetland selection data from 210 radiomarked mallard broods on 15 study areas located throughout the Canadian Prairie-Parklands. We used information-theoretic approaches to select the best-approximating model of habitat selection in relation to wetland characteristics. Wetland permanence, cover type, width of flooded emergent vegetation, and interactions between these variables and date, moisture level, and dominant species of emergent vegetation were all important predictors of wetland selection. Mallard broods selected deeper wetlands, especially later in the brood-rearing season. Mallard broods also selected wetlands with large central expanses of open water and wide peripheral zones of flooded emergent cover. These habitat characteristics can most easily be met in landscapes that already contain an abundance and diversity of natural wetland habitats. Where such wetlands are unavailable, restoration or management of deeper wetlands may be necessary to meet the habitat requirements of mallard ducklings.     

Habitat distribution of the mallard in relation to vegetation structure,food, and population density

The mallard (Anas platyrhynchos Linnaeus) is a generalist feeder, breeding in a wide range of habitats, yet showing considerable between site differences in density. Variations in density and habitat use may result from inter- and intea-specific competition, habitat structure or food.We studied habitat selection of the mallard in four regions of Finland and Sweden. In each region, ten lakes were chosen ranging from oligotrophic to eutrophic. Habitat distribution of the mallard did not differ between regions despite variation in the density of the species and congenerics. Mallard density did not correlate with vegetation structure, but increased with food abundance and the number and density of congenerics although there were regional differences in mallard response.     

Sackville Waterfowl Park,Sackville, New Brunswick,Canada. Baseline characterization,towards long-term monitoring

The Sackville Waterfowl Park, contains a 19 hectare shallow freshwater wetland created by reflooding a saltwater marsh that was drained three centuries ago. Its primary purpose is to provide wetland habitat and wildlife viewing opportunities to tourists and residents. This newly created, eutrophic wetland supports high densities of waterfowl, 2.1 and 3.3 brood ha^–1 in 1991 and 1992 respectively. It is hoped that long term monitoring of the Park's waterfowl population and wetland habitat will contribute to a better understanding of factors controlling breeding waterfowl populations.     

Integrated modeling predicts shifts in waterbird population dynamics under climate change

Climate change has been identified as one of the most important drivers of wildlife population dynamics. The in‐depth knowledge of the complex relationships between climate and population sizes through density dependent demographic processes is important for understanding and predicting population shifts under climate change, which requires integrated population models (IPMs) that unify the analyses of demography and abundance data. In this study we developed an IPM based on Gaussian approximation to dynamic N‐mixture models for large scale population data. We then analyzed four decades (1972–2013) of mallard Anas platyrhynchos breeding population survey, band‐recovery and climate data covering a large spatial extent from North American prairies through boreal habitat to Alaska. We aimed to test the hypothesis that climate change will cause shifts in population dynamics if climatic effects on demographic parameters that have substantial contribution to population growth vary spatially. More specifically, we examined the spatial variation of climatic effects on density dependent population demography, identified the key demographic parameters that are influential to population growth, and forecasted population responses to climate change. Our results revealed that recruitment, which explained more variance of population growth than survival, was sensitive to the temporal variation of precipitation in the southern portion of the study area but not in the north. Survival, by contrast, was insensitive to climatic variation. We then forecasted a decrease in mallard breeding population density in the south and an increase in the northwestern portion of the study area, indicating potential shifts in population dynamics under future climate change. Our results implied that different strategies need to be considered across regions to conserve waterfowl populations in the face of climate change. Our modelling approach can be adapted for other species and thus has wide application to understanding and predicting population dynamics in the presence of global change.     

Nest Site Selection Influences Cinnamon Teal Nest Survival in Colorado

Nest survival of ducks is partially a function of the spatiotemporal characteristics of the site at which a bird chooses to nest. Nest survival is also a fundamental component of population growth in waterfowl but is relatively unstudied for cinnamon teal (Spatula cyanoptera). We investigated cinnamon teal nest survival in a managed wetland complex in southern Colorado, USA, and assessed nest site selection to determine whether nest site characteristics were adaptive. We monitored 85 nests in 2015–2017 on Monte Vista National Wildlife Refuge, Colorado and did not detect a difference in nest survival across years. Based on nest site selection data from 2017, cinnamon teal selected nest sites characterized by a lower proportion of forbs than available sites. The relationships between habitat characteristics and nest survival were variable. Microhabitat characteristics exhibited only weak effects on nest survival during the laying stage. Nest survival during incubation was negatively related to the proportion of forbs at the nest site and, to a lesser extent, the proportion of grasses. Nest site selection was predictive of future nest survival based on the percent of forbs and grasses around the nest site, suggesting teal select nest locations to benefit reproductive success. These results have the potential to guide local habitat management actions for breeding waterfowl. © 2020 The Wildlife Society.     

Multi-scale habitat selection affects offspring survival in a precocial species

In theory, habitat preferences should be adaptive. Accordingly, fitness is often assumed to be greater in preferred habitats; however, this assumption is rarely tested and, when it is, the results are often equivocal. Habitat preferences may not directly convey fitness advantages if animals are constrained by tradeoffs with other selective pressures like predation or food availability. We address unresolved questions about the survival consequences of habitat choices made during brood-rearing in a precocial species with exclusive maternal care (mallard Anas platyrhynchos, n = 582 radio-marked females on 27 sites over 8 years). We directly linked duckling survival with habitat selection patterns at two spatial scales using logistic regression and model selection techniques. At the landscape scale (55–80 km²), females that demonstrated stronger selection of areas with more cover type 4 wetlands and greater total cover type 3 wetland area (wetlands with large expanses of open water surrounded by either a narrow or wide peripheral band of vegetation, respectively) had lower duckling survival rates than did females that demonstrated weaker selection of these habitats. At finer scales (0.32–7.16 km²), females selected brood-rearing areas with a greater proportion of wetland habitat with no consequences for duckling survival. However, females that avoided woody perennial habitats composed of trees and shrubs fledged more ducklings. The relationship between habitat selection and survival depended on both spatial scale and habitats considered. Females did not consistently select brood-rearing habitats that conferred the greatest benefits, an unexpected finding, although one that has also been reported in other recent studies of breeding birds.     

Waterbird Response to Wetlands Restored Through the Conservation Reserve Enhancement Program

Abstract: Conservation programs that facilitate restoration of natural areas on private land are one of the best strategies for recovery of valuable wetland acreage in critical ecoregions of the United States. Wetlands enrolled in the Conservation Reserve Enhancement Program (CREP) provide many ecological functions but may be particularly important as habitat for migrant and resident waterbirds; however, use of, and factors associated with use of, CREP wetlands as stopover and breeding sites have not been evaluated. We surveyed a random sample of CREP wetlands in the Illinois River watershed in 2004 and 2005 to quantify use of restored wetlands by spring migrating and breeding waterbirds. Waterbirds used 75% of wetlands during spring migration. Total use-day abundance for the entire spring migration ranged from 0 to 49,633 per wetland and averaged 6,437 ± 1,887 (SE). Semipermanent wetlands supported the greatest total number of use-days and the greatest number of use-days relative to wetland area. Species richness ranged from 0 to 42 (x̄ = 10.0 ± 1.5 [SE]), and 5 of these species were classified as endangered in Illinois. Density of waterfowl breeding pairs ranged from 0.0 pairs/ha to 16.6 pairs/ha (x̄ = 1.9 ± 0.5 [SE] pairs/ha), and 16 species of wetland birds were identified as local breeders. Density of waterfowl broods ranged from 0.0 broods/ha to 3.6 broods/ha and averaged 0.5 ± 0.1 (SE) broods/ha. We also modeled spring stopover use, waterbird species richness, and waterfowl reproduction in relation to spatial, physical, and floristic characteristics of CREP wetlands. The best approximating models to explain variation in all 3 dependent variables included only the covariate accounting for level of hydrologic management (i.e., none, passive, or active). Active management was associated with 858% greater use-days during spring than sites with only passive water management. Sites where hydrology was passively managed also averaged 402% greater species richness than sites where no hydrologic management was possible. Density of waterfowl broods was 120% greater on passively managed sites than on sites without water management but was 29% less on sites with active compared to passive hydrologic management. Densities of waterfowl broods also were greatest when ratios of open water to cover were 70:30. Models that accounted for vegetation quality and landscape variables ranked lower than models based solely on hydrologic management or vegetation cover in all candidate sets. Although placement and clustering of sites may be critical for maintaining populations of some wetland bird species, these factors appeared to be less important for attracting migrant waterbirds in our study area. In the context of restored CREP wetlands, we suggest the greatest gains in waterbird use and reproduction may be accomplished by emphasizing site-specific restoration efforts related to hydrology and floristic structure. (JOURNAL OF WILDLIFE MANAGEMENT 72(3):654–664; 2008)     

Habitat selection by postbreeding female diving ducks: influence of habitat attributes and conspecifics

Habitat selection studies of postbreeding waterfowl have rarely focused on within‐wetland attributes such as water depth, escape cover, and food availability. Flightless waterfowl must balance habitat selection between avoiding predation risks and feeding. Reproductively successful female ducks face the greatest challenges because they begin the definitive prebasic molt at or near the end of brood rearing, when their body condition is at a low point. We assessed the relative importance of habitat attributes and group effects in habitat selection by postbreeding female lesser scaup Aythya affinis on a 2332‐ha montane wetland complex during the peak flightless period (August) over seven years. Hypothesis‐based habitat attributes included percent open water, open water:emergent edge density, water depth, percent flooded bare substrate, fetch (distance wind can travel unobstructed), group size, and several interactions representing functional responses to interannual variation in water levels. Surveys of uniquely marked females were conducted within randomly ordered survey blocks. We fitted two‐part generalized linear mixed‐effects models to counts of marked females within survey blocks, which allowed us to relate habitat attributes to relative probability of occurrence and, given the presence of a marked female, abundance of marked individuals. Postbreeding female scaup selected areas with water depths > 40 cm, large open areas, and intermediate edge densities but showed no relation to flooded bare substrate, suggesting their habitat preferences were more influenced by avoiding predation risks and disturbances than in meeting foraging needs. Grouping behavior by postbreeding scaup suggests habitat selection is influenced in part by behavioral components and/or social information, conferring energetic and survival benefits (predation and disturbance risks) but potentially also contributing to competition for food resources. This study demonstrates the importance of incorporating group effects and interannual variability in habitat conditions when investigating habitat selection, particularly for seasons when waterfowl are aggregated.     

Not all sanctuaries are created equal: variation in protected area selection by wintering mallards

Public lands managed for wildlife frequently provide various forms of sanctuary to increase residency times and allow access to energetic and other habitat resources for waterfowl. The influence of sanctuary type and disturbance regime on resource use and fine-scale movements of waterfowl has not been investigated extensively using currently available transmitter technologies. We examined mallard (Anas platyrhynchos) use of various types of waterfowl sanctuary and non-sanctuary areas in the Mississippi Alluvial Valley region of eastern Arkansas, USA, during winters of 2019–2021. We deployed 105 global positioning system transmitters on mallards at 4 closed-access spatial sanctuaries on or adjacent to Dale Bumpers White River National Wildlife Refuge. We used hourly transmitter locations to examine mallard use of public sanctuary areas, public hunt areas, and private lands using integrated step selection analysis. Public sanctuary areas provided varying levels of protected status, public hunt areas allowed for varying levels of hunting intensity by duck hunters, and private lands were open to waterfowl hunting and other forms of private uses but may or may not have been hunted at any specific frequency. Mallards selected spatial sanctuary and avoided public hunt areas, other sanctuary types, and private lands during the day. In contrast, mallards selected for private lands over spatial sanctuary at night. Mallards tended to avoid areas that allowed duck hunting or used them during the night when risk of harvest mortality was removed. After the hunting season closed, mallards began using areas that previously allowed duck hunting during the day, suggesting that risk was the primary factor influencing site use. Moreover, mallards were 1.6 times more likely to use public daily hunt areas and 2.1 times more likely to use private lands potentially open to hunting during the day than spatial sanctuary 2 weeks after the close of duck hunting season in February. Spatial sanctuaries appear more effective in influencing mallard use than temporal sanctuaries or inviolate sanctuaries, which are commonly used by state and federal agencies. Partial daily, daily, or activity-specific (e.g., no hunting past noon, no hunting 3 days/week, no waterfowl hunting) closures to encourage mallard use of temporal sanctuaries do not appear to reduce the perceived harvest-related risk to mallards enough for them to view these areas as accessible or significantly increase their use.     

Effects of Habitat on Mallard Duckling Survival in the Great Lakes Region

Abstract: Habitat provides food and shelter resources for prefledgling waterfowl and thus plays a critical role in their growth, development, and survival. However, few studies have examined whether and how particular elements of habitat affect duckling survival. We investigated relationships of duckling survival rates with distance of overland travel, wetland vegetation composition, water permanency, and surrounding upland vegetation for 116 mallard (Anas platyrhynchos) broods in the Great Lakes region from 2001 to 2003. We found that the probability, on hatch day, that a mallard duckling will survive to 55 days was positively related to the proportion of wetland area that was vegetated and negatively related to the proportion of forest cover within 500 m of duckling locations. We found little support for relationships between duckling survival rates and the proportions of grasslands or seasonal wetlands or to distances traveled overland by broods. Our results suggest that conservation groups and wildlife managers in the Great Lakes region can improve mallard duckling survival rates by managing for, creating, and protecting vegetated wetlands and focusing efforts within lightly-forested areas.     

Influence of fish assemblage and shallow lake productivity on waterfowl communities in the Boreal Transition Zone of western Canada

1. Shallow lakes in the Boreal Transition Zone (BTZ) in Alberta, Canada are naturally productive systems that provide important breeding and moulting habitat for many waterfowl (Anseriformes). To examine the relative importance of biotic and abiotic factors on waterfowl population densities, species richness and community composition, we surveyed 30 shallow lakes and evaluated the relationships among fish communities, lake characteristics and waterfowl in both breeding and moulting habitat. Shallow lakes were either fishless (n = 15), contained only small‐bodied fishes (n = 10) or contained large‐bodied, mostly predatory, fish in addition to small‐bodied fish (n = 5). 2. Environmental factors, including water colour, submerged aquatic vegetation, lake area and potassium, explained 24.3% of the variation in breeding waterfowl communities. Fish assemblage contributed independently to a small but significant proportion (13.4%) of the variation, while 13.8% of the explained variation was shared between environmental factors and fish assemblage. In total, 51.5% of the variation in breeding waterfowl communities was explained. 3. Overall, 55.5% of the total variation in moulting waterfowl communities was explained. Environment alone [especially total phosphorus, lake area, maximum depth and dissolved organic carbon (DOC)] and variation shared by fish and environment similarly accounted for most of the explained variation in moulting waterfowl communities (21.7% and 25.7% respectively), while fish assemblage was only one‐third as important (8.1%). 4. Both breeding and moulting waterfowl densities increased with lake productivity, even in eutrophic and hypereutrophic lakes. Breeding waterfowl density was also twice as great in fishless lakes than in lakes with fish, after accounting for lake area. 5. Certain waterfowl taxa were linked to fishless lakes, especially in the moulting season. Canvasback and moulting ring‐necked ducks were linked to small‐bodied fish lakes, whereas moulting common goldeneye were indicators of large‐bodied fish lakes. Knowledge of fish presence and species composition can therefore help guide conservation and management of waterfowl habitat in western Canada. Our results suggest that management efforts to maintain the most productive waterfowl habitat in the BTZ should focus on smaller, shallow, fishless lakes, particularly given that larger fish‐bearing systems have greater regulatory protection.     

The effects of predator removal on mallard production and population change in northeastern North Dakota

In 1994, Delta Waterfowl Foundation began trapping mammalian meso-predators in North Dakota during the breeding season in an attempt to increase waterfowl nest success and enhance recruitment into the fall flight and subsequent breeding population. Multiple studies on these sites demonstrated that removing predators results in near doubling of nest success, which previous simulation modeling suggests is the most influential vital rate influencing the population growth rate of mid-continent mallards (Anas platyrhynchos). We present an assessment of the impact of predator removal on mallard production using population models. We conducted this study on 9 township-sized (93.2 km²) sites (4–8 sites annually per vital rate) in northeastern North Dakota from 2006–2008. Trappers removed mammalian meso-predators on 5 sites and the other 4 served as unmanaged reference sites. To estimate recruitment, we used derived estimates and process variance of pair numbers, hen success (nest survival corrected for renesting), initial brood size, pre-fledging survival, and post-fledging survival, along with previously published estimates of breeding propensity and adult female survival rates. Trapped sites had greater hen success (H = 0.69, = 0.03) than reference sites (H = 0.53, = 0.06), but similar indicated breeding pairs, initial brood size, and pre-fledging survival. We estimated that females on trapped sites added 140 more mallards of both sexes to the fall flight than females on reference sites, at an approximate cost of $74.29 per incremental mallard. Additionally, trapping predators provided a marginal increase (0.04) in finite population growth. We found that predator removal targeted at mammalian nest predators did not produce as many incremental mallards as previously thought and may not be a viable strategy for increasing mallard productivity under conditions similar to those observed during this study. We conducted a sensitivity analysis and determined that pre-fledging survival was the most influential factor regulating mallard population growth. Although hen success increased as a result of trapping, duckling survival became a limiting factor. We suggest that waterfowl managers assess multiple vital rates to determine the likelihood that management actions focused on a single parameter, such as nest success, will yield desired population level effects. © 2012 The Wildlife Society.     

Interspecific interactions and co-existence in dabbling ducks: observations and an experiment

We studied the possible role of resource limitation and interspecific competition in assemblages of dabbling ducks on breeding lakes in Finland and Sweden with observational and experimental data. After initial vegetation mapping and yearly censuses of ducks in 1985–1990, we collected observational data in 1991–1994 from 28 lakes with natural populations of mallard Anas platyrhynchos and teal A. crecca. Mallard and teal co-occur over vast areas in the Holarctic and they are the only breeding dabbling ducks on many oligotrophic lakes. Both species are migratory in our study regions, teal arriving later in spring than mallards. Log-linear model analysis of observational presence/absence data revealed a positive, not a negative, association between the species. This association was independent of habitat diversity as well as of lake size. Mallard-teal interaction was also studied in a cross-over introduction experiment in 32 other lakes in two years. Wing-clipped mallards were introduced to breeding lakes before the arrival of teal to induce resource limitation and interspecific competition, hypothesized to reduce lake use by teal. The density of mallard pairs on experimental lakes was 2.9–8.0 times higher than on controls, but there was no negative response by teal to the treatment. This is the first combined observational-experimental demonstration of lack of interspecific competition in waterfowl. Our results indicate that heterospecific attraction may affect species co-existence in dabbling ducks. Received: 28 October 1996 / Accepted: 13 January 1997     

Livestock grazing in intermountain depressional wetlands: effects on breeding waterfowl

Livestock grazing is a prevalent land use in western North American intermountain wetlands, and physical and biotic changes related to grazing-related disturbance can potentially limit wetland habitat value for waterfowl. We evaluated breeding waterfowl use in 34 wetlands in relation to water retention, amount of wetlands on the landscape, and livestock grazing intensity. The study was conducted over 2 years in the southern intermountain region of British Columbia, Canada. For a subset of 17 wetlands, we measured aquatic invertebrate abundance over 1 year. Waterfowl breeding pairs and broods were classified into three functional groups: dabbling ducks, and two types of diving ducks, overwater and cavity nesters. We evaluated candidate models with variables considered singly and in combination using the Akaike Information Criterion. When selected, bare ground (an indicator of grazing intensity) and wetland density were negatively associated with breeding use while wetland fullness and invertebrate density were positively associated. Each factor was a significant predictor in at least one of the models, but unexpectedly, grazing intensity was the most consistent predictor of waterfowl wetland use (e.g., it was present in more ‘best models’ than wetland fullness). Grazing was associated with declines in the number of waterfowl pairs and broods, likely mediated through effects on wetland vegetation and aquatic macroinvertebrates. Models with site- and landscape-scale variables generally performed better than simpler models. Waterfowl breeding use of wetlands can be improved by reduced livestock grazing intensity adjacent to wetlands and by grazing later in the season. Wetland water retention is also an important constraint on waterfowl use of wetlands and may become more limiting with a shifting climate.     

Occupancy Patterns of Breeding American Black Ducks

Occupancy patterns can assist with the determination of habitat limitation during breeding or wintering periods and can help guide population and habitat management efforts. American black ducks (Anas rubripes; black ducks) are thought to be limited by habitat and food availability during the winter, but breeding sites may also limit the size or growth potential of the population. The Canadian Wildlife Service conducts an annual breeding waterfowl survey that we used to explore the hypothesis that black duck carrying capacity is limited by wetlands available for breeding in Québec, Canada. We applied single-visit, multi-species occupancy models to the 1990–2015 population survey data to determine if there was evidence the black duck population was limited by breeding habitat. Using a dynamic (multi-season) occupancy modeling approach, we estimated latent occupancy (occupancy accounting for imperfect detection) of black ducks and then used latent occupancy estimates to derive occupancy, colonization, and extirpation rates. We jointly modeled the occupancy dynamics of black ducks and other duck species in wetlands where both species were present. Throughout the duration of the survey, 44% of wetlands were never observed to be occupied by black ducks. Occupancy models showed wetland size was positively associated with occupancy at the first time step (initial occupancy) and colonization. All 2-species models indicated initial black duck occupancy, persistence (continued occupancy), and colonization were positively associated with the presence of a second species. Colonization rate over the 26-year period ranged from 7% to 27% across all models. Extirpation rates were similar and were constant through time within each model. Low occupancy rates, combined with approximately equal colonization and extirpation rates, suggest there are available wetlands for breeding black ducks in their core breeding area. If breeding habitats are not saturated, this suggests migration or wintering areas may be more limiting to black duck population abundance. © 2019 The Wildlife Society.     

New species records and changes in abundance of waterfowl in northwest Greenland

Breeding populations of Nearctic and Palearctic waterfowl have undergone significant changes in abundance and distribution over the past 50 years. The Avanersuaq District in northwest Greenland is home to an assemblage of waterfowl from both geographic areas; however, minimal historic or current information is available on species abundance. In 2008 and 2009, we conducted field surveys in Greenland from 76.00° to 77.35°N for breeding and non-breeding waterfowl and have collected anecdotal field notes of avian observations over a 20-year period (1993–2012). During these periods, we documented the first observation of a Ross’s goose (Chen rossii) and the first confirmed breeding by lesser snow geese (Chen caerulescens caerulescens) in Greenland. Northern pintails (Anas acuta) were observed for the first time in northwest Greenland, and a previously unknown breeding location for brent geese (Branta bernicla hrota) was also identified. Local populations of greater snow (C. c.) and Canada geese (B. canadensis) have increased in size. The Booth Sound and Drown Bay wetland areas and many islands throughout the Avanersuaq District were identified as critical habitat for both breeding and non-breeding waterfowl. Further increases in waterfowl abundance, including more frequent rare and new visitors, are likely in the study area as breeding populations further south continue to increase and an ameliorating climate allows for a longer breeding season. These results will prove useful as a baseline for comparisons with future surveys.     

Dynamics of habitat distribution in breeding mallards: assessing the applicability of current habitat selection models

The applicability of ideal free, ideal despotic and ideal preemptive habitat selection models to explain dynamics of habitat distribution of breeding mallards ( Anas platyrhynchos ) was explored. Data from 35 lakes studied between 1985 and 2000 were used to examine overall habitat distribution of breeding pairs, breeding success in different habitats, within-year order of habitat occupation, and density dependence of habitat distribution and breeding success. Two habitat types, rich and poor, were defined based on the structure and luxuriance of shore vegetation; each lake belonged to one or the other of the habitat types. Breeding pairs used the rich habitat more than expected, breeding density also being higher there than in the poor habitat. Both average brood density and breeding success were higher in the rich habitat than in the poor. Breeding success was not density dependent, neither when analysed separately for the habitat types, nor in the study area in general. Within season, arriving mallard pairs did not occupy rich lakes earlier than poor lakes. An isodar analysis based on between-year variation of the breeding density in rich and poor habitats revealed that habitat distribution of breeding pairs was not density dependent. By contrast, density in the rich habitat increased and proportional use of the poor habitat decreased with increasing overall population density, i.e. the rich habitat got increasingly crowded. None of the habitat selection models considered was applicable to explain the dynamics of habitat distribution of breeding mallards.     

Tracking palustrine water seasonal and annual variability in agricultural wetland landscapes using Landsat from 1997 to 2005

Wetlands densely populate the ecoregion transecting the center of the Prairie Pothole Region (PPR) known as the Missouri Coteau and epicenter to the most productive waterfowl‐breeding habitat in North America. These palustrine, depressional basin waters vacillate with regional drought and deluge, so surface water fluctuations over time modulate wetland productivity, habitat, and water quality functions. Models predict formidable effects of climate change on glacial basin surface waters, yet large‐scale, long‐term observation data are lacking to compare against predicted changes. Current, optical‐based water detection models do not delineate marsh vegetation from shallow, turbid, high‐chlorophyll waters common to the region. We developed a palustrine wetland spectral model for tracking open surface waters using Landsat imagery, which we evaluated for a 2500 km² landscape that estimates seasonal and annual open water variability for thousands of individual wetlands in the Missouri Coteau ecoregion. Detection accuracy of 96% was achieved for water bodies greater than a half‐pixel in size. We identified shifts in the distribution of water permanence classes within and between years for waters emerging in spring, mid‐summer, and late summer from 1997 to 2005 and identified a maximum of 19 047 basins with open water (12% of the landscape) populating 2500 km². For the 2005 growing season, we observed only 8757 basins with open water (6% of the landscape) for the same area. Declines were greatest for water bodies detected only in spring, suggesting a loss of those wetlands functioning to recharge groundwater stores early in the season and a high sensitivity to observed reductions in snowfall. If landscape factors driving open water coverage and wetland density are similar for the entire Missouri Coteau, we estimate the number of basins containing at least a pixel of water for this region declined from 577 600 to 266 000 between 1997 and 2005.