Late Ordovician-Early Silurian fluctuations in sea level from eastern Anticosti Island, Quebec

Over 1,000 m of Upper Ordovician to Lower Silurian mixed carbonate and clastic strata on Anticosti Island are nearly tectonically undisturbed, despite their proximity to the Northern Appalachians fronting Quebec's Gulf of St. Lawrence. Natural cliffs exposed along the coast and rivers in the eastern part of the island make a relatively conformable sequence belonging to the Ashgill and Llandovery Series. Fossil communities interpreted as depth-associated in life are especially repetitious in the Becscie, Gun River, Jupiter, and Chicotte Formations (Llandovery Series), and to a lesser degree in the Upper Vaureal and Ellis Bay Formations (Ashgill Series). Preliminary study of the pattern of changeovers in Eocoelia, Pentamerus and Stricklandia communities suggests that Anticosti seas deepened and shallowed three and a half times during the Early Silurian. High water peaks were reached during B₁-B₂, C₁-C₂ and C₄-C₅ times, with a final deepening trend beginning in late C₅ time. Age determinations of these events are based on the occurrence of graptolites (with some new records from Anticosti) keyed to the standard graptolite zones, and species of the Eocoelia lineage are also useful for correlation. The profile of the Anticosti sea-level curve compares well with other curves reconstructed from the Lower Silurian of New York, Michigan, and Iowa. Widespread synchronism in sea-level changes on the North American platform is thus corroborated.     

Chitinozoan zonation of the Late Ordovician and the Early Silurian of the island of Anticosti, Québec, Canada

A reappraisal of chitinozoan distribution across the Ordovician-Silurian boundary on the Island of Anticosti has led to the recognition of a new zone, the Ancyrochitina ellisbayensis biozone, in the uppermost part of the Ellis Bay Formation. This biozone lies between the well defined Upper Ordovician Spinachitina taugourdeaui biozone and the lowest Silurian (Rhuddanian) Plectochitina nodifera biozone of the Becscie Formation. The occurrence of such diagnostic species as P. nodifera, Belonechitina postrobusta, Conochitina electa and Ancyrochitina ramosaspina in the Lower Silurian of Anticosti points to a close similarity to faunas in Estonia and north Latvia and indicates an age ranging from the Parakidograptus acuminatus to the Coronograptus cyphus in terms of graptolite zones. The chitinozoan biozonation harmonizes with that based on conodonts and, to a lesser extent, with the known graptolite faunal succession. Five new species: Ancyrochitina ellisbayensis sp. nov., Clathrochitina postconcinna sp. nov., Conochitina gunriveris sp. nov., Clathrochitina perexilis sp. nov., Bursachitina basiconcava sp. nov. and three species in open nomenclature are described.     

The trace fossil Sinusichnus from the Upper Ordovician of Anticosti Island,eastern Canada

Branched, sinusoidal burrows assigned to Sinusichnus cf. seilacheri Knaust et al. 2016; occur in Upper Ordovician (Hirnantian) carbonates on Anticosti Island. These are abundant and monospecific at the base of the Oncolite Bed of the uppermost Ellis Bay Formation, marking a regional discontinuity surface during the multiphase Ordovician mass-extinction event. In contrast to Mesozoic and Cenozoic records of Sinusichnus, commonly attributed to the activity of decapod and isopod crustaceans, the burrow systems from Anticosti Island were probably produced by other arthropods of unknown affinity. A combined dwelling, locomotion and feeding behaviour of their trace maker is assumed. This is the oldest record of the ichnogenus Sinusichnus, previously only known since the Triassic, and its first evidence from North America.     

贵州思南奥陶纪末至志留纪初大息场组的笔石

贵州思南奥陶纪末至志留纪初大息场组的笔石研究结果表明,大息场组共产出笔石6属7种:Styracograptus chiai(Mu,1949),Normalograptus mirnyensis(Obut and Sobolevskaya,1967),Normalograptus ajjeri(Legrand,1977),Metaclimacograptus hughesi(Nicholson,1869),Avitograptus avitus(Davies,1929),Atavograptus atavus(Jones,1909)和Korenograptus laciniosus(Churkin and Carter,1970)。自下而上可识别出相当Dicellograptus complexus带、Akidograptus ascensus带和Cystograptus vesiculosus-Coronograptus cyphus带的3个含笔石地层带。虽然大息场组的笔石动物群分异度较低,但全都是上扬子区奥陶-志留系之交的常见属种,除S.chiai(Mu)之外,其它都是全球广布的物种。     

江西崇义早奥陶世茅坪组的笔石

记述早奥陶世茅坪组笔石9属25种,其中2新种.根据一些新材料的发现和笔石群的详细研究,将茅坪组笔石划分为3个带,即上部的Adelograptus-Clonograptus 带,中部的Aletograptus-Trograptus带,下部的Staurograptus-Anisograptus带.     

Fossil microphytoplankton dynamics across the Ordovician–Silurian boundary

A critical review of all available data on acritarch biostratigraphy and diversity dynamics across the Late Ordovician through the early Silurian, permits a better appreciation of the potential of acritarchs for the recognition of the systemic boundary. This analysis also reveals the response of marine microphytoplankton populations to the Late Ordovician palaeoenvironmental crisis (Hirnantian glaciation). Previous zonal schemes are improved, and an updated acritarch biostratigraphic chart is proposed, plotted against the most recent chronostratigraphic subdivisions. Sections from Anticosti Island (Québec, Canada), Algeria, Morocco, and Estonia preserve the best palynological record for the investigated interval. The present analysis shows that no true mass-extinction event occurred in latest Ordovician times in connection with the well known glacial event. “Pre-glacial” Ashgill acritarch suites are dominated by species of Baltisphaeridium, Multiplicisphaeridium, Ordovicidium, Orthosphaeridium, and netromorph acritarchs. An important proportion of these taxa (excluding Ordovicidium and Orthosphaeridium) survive the onset of glacial conditions in Hirnantian (latest Ordovician) times and continues through the early Silurian. The development of morphological polymorphism appears as a response (a survival strategy?) to the establishment of glacial conditions.In glacial-related sediments of Hirnantian age in North Africa, acritarch assemblages display a burst of relative abundance and intra-specific morphological variability (polymorphism) of long-ranging taxa such as Veryhachium, Multiplicisphaeridium, Dactylofusa, Poikilofusa, and Evittia. The extinction of several species characteristic of Upper to uppermost Ordovician strata occurs near the boundary, in “post-glacial” Ashgill (uppermost Hirnantian). This extinction event is counterbalanced by the almost contemporaneous (within the limits of stratigraphic resolution) appearance of several new taxa showing already a clear “Silurian affinity”, e.g., Tylotopalla, Cymbosphaeridium, and Visbysphaera. This origination event is observable in, and correlatable between the North African, the Bohemian and the Anticosti sections, making it global in extent.The completion of the palynological turnover and the establishment of a diverse Silurian acritarch suite occurs well above the base of the Silurian, during Aeronian times. The strong survival capability of the oceanic plankton through periods of palaeoenvironmental crisis in latest Ordovician times (but also throughout the Phanerozoic) could have played an important role in the post-extinction rebounds of metazoan clades, by assuring the continuity of marine trophic resources to consumers and avoiding irreversible disruptions of the trophic chains.     

A relict Ordovician brachiopod fauna from the Parakidograptus acuminatus Biozone (lower Silurian) of the English Lake District

At Yewdale Beck in northern England, a brachiopod fauna dominated by species of Hindella, Kinnella, Mirorthis, Paromalomena and Plectothyrella , occurs interbedded with a graptolite fauna which includes Akidograptus ascensus , Atavograptus ceryx , Persculptograptus parvulus and Normalograptus spp. (including probable examples of N. normalis , N. angustus and N. medius ) within the lower part of the Skelgill Formation. This suggests that taxa of the terminal Ordovician Hirnantia fauna occur within the lower P. acuminatus Biozone, representing the youngest documented occurrence of the Hirnantia brachiopod fauna. Biostratigraphy, brachiopods, graptolites, Ordovician-Silurian boundary.     

Discovery of Anticostia uniformis from the Xiazhen Formation at Zhuzhai,South China and its stratigraphic implication

The Xiazhen Formation is an Upper Ordovician lithostratigraphic unit in the Jiangshan-Changshan-Yushan (JCY) area, which contains series of Late Ordovician reef successions. The reef successions of the Xiazhen Formation at Zhuzhai are critical for evaluation of the Late Ordovician marine diversity and palaeoecology. However, their age has long been uncertain and generally is regarded as of upper Katian, based on the occurrences of shelly fossils and correlation with the stratigraphic equivalent Changwu Formation. The newly discovered graptolite species Anticostia uniformis, in the Xiazhen Formation, together with the combined evidence of brachiopods and sedimentology, indicates an age range for the graptolite locality from the Dicellograptus complanatus Biozone to the Diceratograptus mirus Subzone of late Katian, but the graptolites do not rule out the possibility that it is Hirnantian.     

Population structure of graptolite assemblages

Graptolite rhabdosomes display a diverse suite of morphologies. The range of morphotypes present within most moderate- to high-diversity assemblages from the Ordovician and Silurian is similar, despite the different taxonomic composition of the faunas at different times. Survivorship analyses of graptolite faunas from the Ordovician and Silurian demonstrate strong similarities in the mortality rates of unrelated graptolites of similar functional morphology. They also show a strong correlation between decreasing mortality rates among more mature colonies with increasing rhabdosome complexity. This similarity in both functional morphology and life history of graptolites suggests that they lived within a very stable planktic community structure.     

An Unusually Diverse Llandovery (Silurian) Diplograptid Fauna from the Southern Urals of Russia and Its Evolutionary Significance

This vesiculosus Biozone assemblage of chemically-isolated graptolites from the Orenburg District of Russia was obtained from limestones associated with pillow lavas of the Sakmara Formation of Silurian–Early Devonian age. It comprises 30 species, of which 24 are described. Nine are new, one is a new subspecies, and eight are left under open nomenclature. The assemblage is unique in its diversity of minute biserial graptolites referable to the genera ' Glyptograptus ', Normalograptus , Sudburigraptus , and Metaclimacograptus ; and is unusual in that well-preserved graptolite faunas are rare at this stratigraphical level, which is fixed by the occurrence of Cystograptus vesiculosus (Nicholson) s.l. and Paraclimacograptus innotatus (Nicholson). The following new forms are described: ' Glyptograptus ' tamariscus praecursor subsp. nov., G. mazovoensis sp. nov., G. nikitini sp. nov., G. pavlinovi sp. nov., G. telarius sp. nov., G. zimai sp. nov., Normalograptus legrandi sp. nov., N. melchini sp. nov., N. sobolevskayae sp. nov., and Metaclimacograptus bjerreskovae sp. nov. Five evolutionary lineages are defined within the ' Glyptograptus ' tamariscus (Nicholson) plexus, modifying and extending the pioneer work of Packham (1962). The diverse ' Glyptograptus ' faunas are considered to be a replacement for the Normalograptus fauna which peaked in the late Ordovician and early Silurian. Metaclimacograptus , although reasonably diverse in its speciation, exhibits a conservative style of evolution with little fundamental change throughout the Llandovery. It is suggested that cortical bandages have species-specific dimensions and micro-features. Some problematical micro- and ultrastructural features are described, notably fibrillar textures, parallel to the fusellar margins, in several species.     

内蒙古西部额济纳旗晚奥陶世生物地理和奥陶-志留系分界

通过最近对内蒙古额济纳旗东南珠斯楞海尔罕一带(阿拉善地块)奥陶—志留系界线地层和生物群的野外调查和室内研究,首次在上奥陶统上部巴丹吉林组中发现阿什极尔中期(mid Ashgill)腕足动物化石巨大全嘴贝(Holorhymchus giganteus Kiaer),这是该属在我国奥陶系中的首次记录。根据未成年个体的切片所发现的腹壳顶端发育中隔板构造,修订该属定义,并证明研究小个体对识别物种特征、探讨个体发育和系统演化有重要意义。分析了全嘴贝属的群落生态、生物地理和演化意义后,发现它在奥陶纪末大灭绝过程中,因居群规模和分布范围极大地缩减,至今未发现其化石记录;但在志留纪初环境好转后继续存活,可暂视其为复活属。根据在拐子湖组近底部发现志留系最下部Akidograptus ascensus带的重要分子Normalograptus lubricus Chen et Lin,确定本区的奥陶—志留系分界。阿拉善地块因不发育Hirnantia动物群,表明奥陶纪末期它与扬子区、西藏、滇西等富含该动物群的地区属于不同的生物地理区系;在阿什极尔中期可能位于热带海域,与祁连山、中亚、乌拉尔等地有重要联系。文中描记H．giganteus种,评述正常笔石类(normalograptid)的系统分类位置并描记N．lubricus种。     

The effects of the final stages of the Late Ordovician glaciation on marine palynomorphs (chitinozoans, acritarchs, leiospheres) in well Nl-2 (NE Algerian Sahara)

Palynomorph assemblages, especially chitinozoans and acritarchs, from the Upper Ordovician of well Nl-2 (north-east of the Algerian Sahara) are studied in order to precisely date the ultimate effects of the Late Ordovician glaciation and to document the impact of this major climatic stress on the diversity of the palynoplankton. References are made to stable isotope excursions and to global eustatic sea level variations in order to improve the local age determination. The Hassi el Hadjar Formation, i.e. glacio-marine diamictites, is interpreted as a transgressive event resulting from the melting of the northern Gondwana ice cap. It yields poorly preserved and moderately diverse chitinozoans of late Hirnantian age. Acritarchs are more abundant in the lower part of these "microconglomeratic clays", but display a low diversity and are badly preserved throughout the whole formation. Reworked individuals are recorded in both groups. The marine sediments of the M'Kratta Formation of latest Hirnantian age contain better preserved, more abundant and more diverse palynomorph assemblages, especially in the Upper Member. The composition of this palynoplankton indicates a fairly good faunal and phytoplankton recovery after the early Hirnantian climatic stress.The extinction of the Ordovician forms, and the appearance of Silurian type taxa occur only in the uppermost Hirnantian, i.e. following with a slight delay the glacial event. The overlying black shales of Wenlock age (lower part of the Oued Mehaiguène Formation) are indicative of marine anoxic environments. They yield a virtually exclusive, but enormous number of Tasmanacea. The very peculiar composition of this palynoplankton seems to be independent of the Late Ordovician glaciation and is most likely related to the factors that, later, generated and maintained anoxic conditions in this area.A new species of chitinozoan, Spinachitina oulebsiri sp. nov. from the latest Hirnantian M'Kratta Formation, is described and illustrated.     

HIRNANTIAN (LATEST ORDOVICIAN) GRAPTOLITES FROM THE UPPER YANGTZE REGION, CHINA

Abstract:  The Upper Yangtze region yields a Hirnantian (latest Ordovician) graptolite fauna that includes 41 species assigned to 13 genera. This fauna is particularly important for understanding the Late Ordovician mass extinction event because it is the most diverse known from this interval. In addition, it records the survival, well into the Hirnantian, of many taxa of the Dicranograptidae-Diplograptidae-Orthograptidae (DDO) fauna, which was previously regarded as having gone extinct at the beginning of the Hirnantian. Taxa exhibiting six different astogenetic patterns, including taxa with reclined stipes, scandent, biserial, full-periderm and 'archiretiolitid' rhabdosome forms occur in the lower Normalograptus extraordinarius-N. ojsuensis Biozone. In contrast, in the upper N. persculptus Biozone only four genera remain, all but one of which are Normalograptidae: scandent and biserial taxa with Pattern H astogeny. Normalograptids are the dominant form of the succeeding, lower Rhuddanian, faunas. The Yangtze faunas also document the early expansion of normalograptids coeval with the decline of the DDO fauna. Many previously identified species considered endemic to China have been synonymized; 24 of the 41 species recorded here have been recognized elsewhere. No new taxa are described.     

The cyanophyte Wetheredella in Ordovician reefs and off-reef sediments

Wetheredella tumulus n.sp. was an important constituent, both binder and frame builder, in reef and off-reef sediments of the Ashgillian Ellis Bay Formation of Anticosti Island in eastern Canada. In individual small, stromatolite-like mounds, it built a skeletal platform for attachment of associated serpulid worm tubes, bryozoans, crinoids and small brachiopods. In larger bioherms, Wetheredella was an encruster, binding waveswept reef tops of halysitid and heliolitid corals. Wetheredella is interpreted as a blue-green alga related to Silurian Rothpletzella from Gotland and the genus Girvanella.     

Cryptospore Assemblages from the Ordovician/silurian Boundary in the Puna Region, North-west Argentina

Palynomorph assemblages have been recovered from deposits believed to straddle the Ordovician/Silurian boundary, from the upper member of the Salar del Rincón Formation, in the Puna region of north-west Argentina. The palynomorph assemblages are dominated by terrestrial cryptospores, but also contain marine elements (acritarchs, prasinophycean algae and chitinozoans). The cryptospore assemblages are similar in composition to those described from coeval deposits worldwide, suggesting that the producers were cosmopolitan and tolerated a wide range of climatic conditions. They are correlated with the Imperfectotriletes spp. Interval Biozone (sub-biozone α) of the Imperfectotriletes spp.- Laevolancis divellomedia Assemblage Biozone, indicating a Hirnantian (latest Ordovician) age. Acritarchs include late Ordovician species such as Eupoikilosusa striata and Villosacapsula cf. setosapellicula , that coexist with the Llandovery species Dactylofusa estillis. Based on lithological and palynological evidence, an early Llandovery, or a late Hirnantian (post-glacial) age is proposed for the basal part of the upper member of the Salar del Rincón Formation.     

东秦岭下志留统的笔石

笔石采自河南淅川下志留统张湾组,其时代与华中区龙马溪组相当。包含Ｄ．ｔｒｉａｎｇｕｌａｔｕｓ,Ｏ．ｃｏｍｍｕｎｉｓ和Ｍ．ｓｅｄｇｗｉｃｋｉｉ３个笔石带,可以与国内外同时代笔石带相对比。共记述正笔石目２科９属１５种,其中含２新种和１新亚种。     

Upper Ordovician microphytoplankton of the Bill's Creek Shale and Stonington Formation,Upper Peninsula of Michigan,U.S.A.: Biostratigraphy and paleogeographic significance

An abundant, diverse, and well-preserved organic-walled microphytoplankton assemblage is described from the Upper Ordovician Bill's Creek Shale and the lower Stonington Formation (Bay de Noc Member) in the Upper Peninsula of Michigan, U.S.A. Based on graptolite and conodont evidence, the Bill's Creek Shale and Stonington Formation are Richmondian (=Ashgill) in age. The assemblage is dominated by acritarchs, which comprise 29 species (including the enigmatic palynomorph Gloeocapsomorpha prisca) assigned to 20 genera. The prasinophyte phycomata are represented by undifferentiated species of Leiosphaeridia and Tasmanites. In addition, chitinozoans are abundant, and scolecodonts and graptolite fragments are common. Paleontologic-palynologic and sedimentologic evidence indicates that the Bill's Creek Shale was deposited in a low-energy, shallow, nearshore marine environment. The overlying Bay de Noc Member of the Stonington Formation also accumulated in a low-energy, normal marine environment, but in a more offshore, somewhat deeper water setting. Both formations experienced minor transgressive and regressive episodes as indicated by fluctuations in the composition of the palynoflora. The combined Bill's Creek/Stonington acritarch assemblage closely resembles those described from the Richmondian-aged Maquoketa Shale (Missouri and Kansas), Sylvan Shale (Oklahoma), and Vauréal Formation (Anticosti Island, Québec, Canada). The overall composition of the acritarch assemblage from these four formations reflects a distinctive, recognizably Laurentian character. Nonetheless, many of the Bill's Creek/Stonington acritarchs have been reported from Upper Ordovician localities elsewhere, providing additional evidence for Late Ordovician cosmopolitanism of the marine microphytoplankton community. Additionally, the restricted stratigraphic range of many of the taxa further enhances their biostratigraphic application, both regionally and globally, and reaffirms the Richmondian (=Ashgill) age of the Bill's Creek Shale and Stonington Formation.     

Placement of the Wenlockian/Ludlovian boundary in western New York State

Despite many years of detailed examination focused on the Silurian strata of New York State, the position of the Wenlockian/Ludlovian boundary has, to date, not been satisfactorily cstablished. Reccnt discovery of early Ludlovian graptoloid graptolites, however, provides the first conclusive biostratigraphic evidence for placement of this chronostratigraphic boundary within the New York sections. The carly Ludlovian genera Saetograptus and Spinograptus were recovered from a thin, but laterally persistent bed of argillaceous dolostone located near the base of the Goat Island Formation. This occurrencc indicates that the Wenlockian/Ludlovian boundary is at or near the contact between the Goat Island Formation and the underlying Gasport Formation, considerably lower than had previously been suspected. □ Silurian, Wenlockian, Ludlovian, graptolites, stratigraphy. biostratigraphy.     

Revision of the Zone 13 graptolite biostratigraphy in the Marathon, Texas, standard succession and its bearing on Upper Ordovician graptolite biogeography

One of the long-standing problems in North American graptolite biostratigraphy is the distinct differences in assemblages of post-Climacograptus bicornis age between the classical graptolite sequences in the New York - Quebec and Marathon, west Texas, regions. These have been attributed either to faunal provincialism or to the presence of a major hiatus between the Woods Hollow and Maravillas formations in Texas. New collections from the key Marathon Picnic Grounds section contain diagnostic Late Ordovician graptolites that confirm the existence of a major stratigraphic gap below the Maravillas Formation. The lower Maravillas Formation (Zone 13) has a Late Ordovician, low-diversity Pacific Province graptolite fauna that includes the biostratigraphically diagnostic species Climacograptus nevadensis, C. tubuliferus, Orthograptus fastigatus and Dicellograptus ornatus. Zone 13 graptolite assemblages from the Marathon region correlate with the C. tubuliferus to D. ornatus zones in the Trail Creek, Idaho, succession, the Ea4-Bo2 interval in Victoria, the O. fastigatus Zone in the Canadian Arctic Islands, the O. quadrimucronatus to D. ornatus zones of the Canadian Cordillera, and the D. complanatus to D. anceps zones in Scotland. The hiatus between the Woods Hollow and Maravillas formations spans an interval corresponding to at least the Eal Ea3 interval in Australia, the C. americanus to upper A. manitoulinensis zones in the New York - Quebec succession, and the D. clingani and P. linearis zones in Scotland. These results agree with the magnitude of the hiatus previously indicated by conodont biostratigraphy. Late Ordovician graptolite distribution patterns in North America can be explained by an extension of Cooper, Fortey & Lindholm's (1991; Lethaia 24) Lower Ordovician graptolite biofacies model into the Upper Ordovician, which incorporates both lateral water-mass specificity and depth stratification. Using this model, we recognize in Laurentia two separate biofacies among tropical-zone Late Ordovician Pacific Province graptolite faunas, a cosmopolitan Oceanic biofacies, and a cratonic Laurentian biofacies. The lower Maravillas Formation graptolite fauna is clearly part of the Oceanic biofacies, whereas the coeval Appalachian faunas represent the Laurentian biofacies. □Graptolites, Ordovician, biostratigraphy, Texas, biofacies, biogeography.